PDB entry 2fym

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HEADER    LYASE                                   08-FEB-06   2FYM              
TITLE     CRYSTAL STRUCTURE OF E. COLI ENOLASE COMPLEXED WITH THE               
TITLE    2 MINIMAL BINDING SEGMENT OF RNASE E.                                  
COMPND    MOL_ID: 1;                                                            
COMPND   2 MOLECULE: ENOLASE;                                                   
COMPND   3 CHAIN: A, C, D, F;                                                   
COMPND   4 SYNONYM: 2-PHOSPHOGLYCERATE DEHYDRATASE, 2-PHOSPHO-D-                
COMPND   5 GLYCERATE HYDRO-LYASE;                                               
COMPND   6 EC: 4.2.1.11;                                                        
COMPND   7 ENGINEERED: YES;                                                     
COMPND   8 MOL_ID: 2;                                                           
COMPND   9 MOLECULE: RIBONUCLEASE E;                                            
COMPND  10 CHAIN: B, E;                                                         
COMPND  11 FRAGMENT: RESIDUES 833-850;                                          
COMPND  12 SYNONYM: RNASE E;                                                    
COMPND  13 EC: 3.1.4.-;                                                         
COMPND  14 ENGINEERED: YES                                                      
SOURCE    MOL_ID: 1;                                                            
SOURCE   2 ORGANISM_SCIENTIFIC: ESCHERICHIA COLI;                               
SOURCE   3 ORGANISM_COMMON: BACTERIA;                                           
SOURCE   4 GENE: ENO;                                                           
SOURCE   5 EXPRESSION_SYSTEM: ESCHERICHIA COLI;                                 
SOURCE   6 EXPRESSION_SYSTEM_COMMON: BACTERIA;                                  
SOURCE   7 EXPRESSION_SYSTEM_STRAIN: BL21(DE3);                                 
SOURCE   8 EXPRESSION_SYSTEM_VECTOR_TYPE: PLASMID;                              
SOURCE   9 EXPRESSION_SYSTEM_PLASMID: PET11A;                                   
SOURCE  10 MOL_ID: 2;                                                           
SOURCE  11 SYNTHETIC: YES;                                                      
SOURCE  12 OTHER_DETAILS: THE E. COLI RNASE E PEPTIDE WAS SYNTHESIZED.          
KEYWDS    RNA DEGRADOSOME, ENOLASE, RNASE E                                     
EXPDTA    X-RAY DIFFRACTION                                                     
AUTHOR    V.CHANDRAN,B.F.LUISI                                                  
REVDAT   2   11-APR-06 2FYM    1       JRNL                                     
REVDAT   1   28-FEB-06 2FYM    0                                                
JRNL        AUTH   V.CHANDRAN,B.F.LUISI                                         
JRNL        TITL   RECOGNITION OF ENOLASE IN THE ESCHERICHIA COLI RNA           
JRNL        TITL 2 DEGRADOSOME                                                  
JRNL        REF    J.MOL.BIOL.                   V. 358     8 2006              
JRNL        REFN   ASTM JMOBAK  UK ISSN 0022-2836                               
REMARK   1                                                                      
REMARK   2                                                                      
REMARK   2 RESOLUTION. 1.60 ANGSTROMS.                                          
REMARK   3                                                                      
REMARK   3 REFINEMENT.                                                          
REMARK   3   PROGRAM     : REFMAC 5.2.0005                                      
REMARK   3   AUTHORS     : MURSHUDOV,VAGIN,DODSON                               
REMARK   3                                                                      
REMARK   3    REFINEMENT TARGET : MAXIMUM LIKELIHOOD                            
REMARK   3                                                                      
REMARK   3  DATA USED IN REFINEMENT.                                            
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.60                           
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 19.99                          
REMARK   3   DATA CUTOFF            (SIGMA(F)) : NULL                           
REMARK   3   COMPLETENESS FOR RANGE        (%) : 90.9                           
REMARK   3   NUMBER OF REFLECTIONS             : 205585                         
REMARK   3                                                                      
REMARK   3  FIT TO DATA USED IN REFINEMENT.                                     
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT                      
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM                          
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.167                           
REMARK   3   R VALUE            (WORKING SET) : 0.165                           
REMARK   3   FREE R VALUE                     : 0.201                           
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.000                           
REMARK   3   FREE R VALUE TEST SET COUNT      : 10847                           
REMARK   3                                                                      
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.                                  
REMARK   3   TOTAL NUMBER OF BINS USED           : 20                           
REMARK   3   BIN RESOLUTION RANGE HIGH           : 1.60                         
REMARK   3   BIN RESOLUTION RANGE LOW            : 1.64                         
REMARK   3   REFLECTION IN BIN     (WORKING SET) : 15113                        
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 91.22                        
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2010                       
REMARK   3   BIN FREE R VALUE SET COUNT          : 820                          
REMARK   3   BIN FREE R VALUE                    : 0.2640                       
REMARK   3                                                                      
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.                    
REMARK   3   ALL ATOMS                : 14807                                   
REMARK   3                                                                      
REMARK   3  B VALUES.                                                           
REMARK   3   FROM WILSON PLOT           (A**2) : NULL                           
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 20.67                          
REMARK   3   OVERALL ANISOTROPIC B VALUE.                                       
REMARK   3    B11 (A**2) : -0.02000                                             
REMARK   3    B22 (A**2) : -0.02000                                             
REMARK   3    B33 (A**2) : 0.03000                                              
REMARK   3    B12 (A**2) : 0.00000                                              
REMARK   3    B13 (A**2) : -0.01000                                             
REMARK   3    B23 (A**2) : 0.00000                                              
REMARK   3                                                                      
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.                                 
REMARK   3   ESU BASED ON R VALUE                            (A): 0.125         
REMARK   3   ESU BASED ON FREE R VALUE                       (A): 0.091         
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): 0.052         
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): 3.234         
REMARK   3                                                                      
REMARK   3 CORRELATION COEFFICIENTS.                                            
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.968                         
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.954                         
REMARK   3                                                                      
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES        COUNT    RMS    WEIGHT      
REMARK   3   BOND LENGTHS REFINED ATOMS        (A): 13050 ; 0.009 ; 0.022       
REMARK   3   BOND LENGTHS OTHERS               (A):  NULL ;  NULL ;  NULL       
REMARK   3   BOND ANGLES REFINED ATOMS   (DEGREES): 17604 ; 1.157 ; 1.969       
REMARK   3   BOND ANGLES OTHERS          (DEGREES):  NULL ;  NULL ;  NULL       
REMARK   3   TORSION ANGLES, PERIOD 1    (DEGREES):  1744 ; 5.413 ; 5.000       
REMARK   3   TORSION ANGLES, PERIOD 2    (DEGREES):   517 ;40.267 ;25.745       
REMARK   3   TORSION ANGLES, PERIOD 3    (DEGREES):  2277 ;12.064 ;15.000       
REMARK   3   TORSION ANGLES, PERIOD 4    (DEGREES):    46 ;18.674 ;15.000       
REMARK   3   CHIRAL-CENTER RESTRAINTS       (A**3):  2013 ; 0.083 ; 0.200       
REMARK   3   GENERAL PLANES REFINED ATOMS      (A):  9690 ; 0.004 ; 0.020       
REMARK   3   GENERAL PLANES OTHERS             (A):  NULL ;  NULL ;  NULL       
REMARK   3   NON-BONDED CONTACTS REFINED ATOMS (A):  7024 ; 0.194 ; 0.200       
REMARK   3   NON-BONDED CONTACTS OTHERS        (A):  NULL ;  NULL ;  NULL       
REMARK   3   NON-BONDED TORSION REFINED ATOMS  (A):  9218 ; 0.302 ; 0.200       
REMARK   3   NON-BONDED TORSION OTHERS         (A):  NULL ;  NULL ;  NULL       
REMARK   3   H-BOND (X...Y) REFINED ATOMS      (A):  1764 ; 0.137 ; 0.200       
REMARK   3   H-BOND (X...Y) OTHERS             (A):  NULL ;  NULL ;  NULL       
REMARK   3   POTENTIAL METAL-ION REFINED ATOMS (A):  NULL ;  NULL ;  NULL       
REMARK   3   POTENTIAL METAL-ION OTHERS        (A):  NULL ;  NULL ;  NULL       
REMARK   3   SYMMETRY VDW REFINED ATOMS        (A):    69 ; 0.171 ; 0.200       
REMARK   3   SYMMETRY VDW OTHERS               (A):  NULL ;  NULL ;  NULL       
REMARK   3   SYMMETRY H-BOND REFINED ATOMS     (A):    78 ; 0.147 ; 0.200       
REMARK   3   SYMMETRY H-BOND OTHERS            (A):  NULL ;  NULL ;  NULL       
REMARK   3   SYMMETRY METAL-ION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL       
REMARK   3   SYMMETRY METAL-ION OTHERS         (A):  NULL ;  NULL ;  NULL       
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.     COUNT   RMS    WEIGHT      
REMARK   3   MAIN-CHAIN BOND REFINED ATOMS  (A**2):  8843 ; 0.844 ; 1.500       
REMARK   3   MAIN-CHAIN BOND OTHER ATOMS    (A**2):  NULL ;  NULL ;  NULL       
REMARK   3   MAIN-CHAIN ANGLE REFINED ATOMS (A**2): 13607 ; 1.237 ; 2.000       
REMARK   3   SIDE-CHAIN BOND REFINED ATOMS  (A**2):  4702 ; 2.697 ; 3.000       
REMARK   3   SIDE-CHAIN ANGLE REFINED ATOMS (A**2):  3994 ; 3.078 ; 4.500       
REMARK   3                                                                      
REMARK   3 ANISOTROPIC THERMAL FACTOR RESTRAINTS.    COUNT   RMS   WEIGHT       
REMARK   3   RIGID-BOND RESTRAINTS          (A**2): 13545 ; 2.495 ; 3.000       
REMARK   3   SPHERICITY; FREE ATOMS         (A**2):  1934 ; 3.556 ; 3.000       
REMARK   3   SPHERICITY; BONDED ATOMS       (A**2): 12873 ; 2.843 ; 3.000       
REMARK   3                                                                      
REMARK   3  NCS RESTRAINTS STATISTICS                                           
REMARK   3   NUMBER OF DIFFERENT NCS GROUPS : 0                                 
REMARK   3                                                                      
REMARK   3  TLS DETAILS                                                         
REMARK   3   NUMBER OF TLS GROUPS  : 0                                          
REMARK   3                                                                      
REMARK   3  BULK SOLVENT MODELLING.                                             
REMARK   3   METHOD USED : MASK                                                 
REMARK   3   PARAMETERS FOR MASK CALCULATION                                    
REMARK   3   VDW PROBE RADIUS   : 1.20                                          
REMARK   3   ION PROBE RADIUS   : 0.80                                          
REMARK   3   SHRINKAGE RADIUS   : 0.80                                          
REMARK   3                                                                      
REMARK   3  OTHER REFINEMENT REMARKS: HYDROGENS HAVE BEEN ADDED IN THE          
REMARK   3  RIDING POSITIONS                                                    
REMARK   4                                                                      
REMARK   4 2FYM COMPLIES WITH FORMAT V. 3.0, 1-DEC-2006                         
REMARK   4                                                                      
REMARK   4 THIS IS THE REMEDIATED VERSION OF THIS PDB ENTRY.                    
REMARK   4 REMEDIATED DATA FILE REVISION 3.100 (2007-03-17)                     
REMARK 100                                                                      
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB.                               
REMARK 100 THE RCSB ID CODE IS RCSB036466.                                      
REMARK 200                                                                      
REMARK 200 EXPERIMENTAL DETAILS                                                 
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION                  
REMARK 200  DATE OF DATA COLLECTION        : 08-MAY-2004; 24-FEB-2004; 09-      
REMARK 200                                   MAR-2004; 08-MAY-2004              
REMARK 200  TEMPERATURE           (KELVIN) : 100; 100; 100; 100                 
REMARK 200  PH                             : 7.0; 8.2; 6.0; 7.5                 
REMARK 200  NUMBER OF CRYSTALS USED        : 1                                  
REMARK 200                                                                      
REMARK 200  SYNCHROTRON              (Y/N) : Y; Y; N; Y                         
REMARK 200  RADIATION SOURCE               : ESRF; ESRF; ROTATING ANODE;        
REMARK 200                                   ESRF                               
REMARK 200  BEAMLINE                       : ID14-4; ID29; ID14-4               
REMARK 200  X-RAY GENERATOR MODEL          : NULL; RIGAKU                       
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M; M; M; M                         
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.0; 0.9782; 1.54; 1.0             
REMARK 200  MONOCHROMATOR                  : GE(220) CRYSTAL; SI(111)           
REMARK 200                                   CRYSTAL; GE(220) CRYSTAL           
REMARK 200  OPTICS                         : NULL                               
REMARK 200                                                                      
REMARK 200  DETECTOR TYPE                  : Q315R ADSC DETECTOR; ADSC          
REMARK 200                                   Q210 2D; IMAGE PLATE; Q315R        
REMARK 200                                   ADSC DETECTOR                      
REMARK 200  DETECTOR MANUFACTURER          : ADSC QUANTUM 4; ADSC QUANTUM       
REMARK 200                                   4; RIGAKU RAXIS IV; ADSC           
REMARK 200                                   QUANTUM 4                          
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : DENZO                              
REMARK 200  DATA SCALING SOFTWARE          : HKL                                
REMARK 200                                                                      
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 216432                             
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.600                              
REMARK 200  RESOLUTION RANGE LOW       (A) : 20.000                             
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 2.000                              
REMARK 200                                                                      
REMARK 200 OVERALL.                                                             
REMARK 200  COMPLETENESS FOR RANGE     (%) : 91.1                               
REMARK 200  DATA REDUNDANCY                : 10.300                             
REMARK 200  R MERGE                    (I) : 0.03400                            
REMARK 200  R SYM                      (I) : NULL                               
REMARK 200   FOR THE DATA SET  : 13.7000                            
REMARK 200                                                                      
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.                                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.60                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.63                     
REMARK 200  COMPLETENESS FOR SHELL     (%) : 94.3                               
REMARK 200  DATA REDUNDANCY IN SHELL       : 16.51                              
REMARK 200  R MERGE FOR SHELL          (I) : 0.25300                            
REMARK 200  R SYM FOR SHELL            (I) : NULL                               
REMARK 200   FOR SHELL         : 2.450                              
REMARK 200                                                                      
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH; SINGLE WAVELENGTH;          
REMARK 200                       SINGLE WAVELENGTH; SINGLE WAVELENGTH           
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT        
REMARK 200 SOFTWARE USED: MOLREP                                                
REMARK 200 STARTING MODEL: ENOLASE STRUCTURE 1E9I                               
REMARK 200                                                                      
REMARK 200 REMARK: NULL                                                         
REMARK 280                                                                      
REMARK 280 CRYSTAL                                                              
REMARK 280 SOLVENT CONTENT, VS   (%): 50.11                                     
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.47                     
REMARK 280                                                                      
REMARK 280 CRYSTALLIZATION CONDITIONS: 2.4M SODIUM MALONATE, PH 7.0, VAPOR      
REMARK 280  DIFFUSION, HANGING DROP, TEMPERATURE 298.0K. 40% MPEG 550,          
REMARK 280  0.1M HEPES, PH 8.2, VAPOR DIFFUSION, HANGING DROP, TEMPERATURE      
REMARK 280  298.0K. 8% PEG 4K, 0.2 M IMIDAZOLE MALEATE, PH 6.0, VAPOR           
REMARK 280  DIFFUSION, HANGING DROP, TEMPERATURE 298.0K. 27% PEG 600, 0.1M      
REMARK 280  HEPES, PH 7.5, VAPOR DIFFUSION, HANGING DROP, TEMPERATURE           
REMARK 280  298.0K                                                              
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY                                            
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 1 21 1                         
REMARK 290                                                                      
REMARK 290      SYMOP   SYMMETRY                                                
REMARK 290     NNNMMM   OPERATOR                                                
REMARK 290       1555   X,Y,Z                                                   
REMARK 290       2555   -X,1/2+Y,-Z                                             
REMARK 290                                                                      
REMARK 290     WHERE NNN -> OPERATOR NUMBER                                     
REMARK 290           MMM -> TRANSLATION VECTOR                                  
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS                            
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM             
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY                
REMARK 290 RELATED MOLECULES.                                                   
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   2  0.000000  1.000000  0.000000       62.10050            
REMARK 290   SMTRY3   2  0.000000  0.000000 -1.000000        0.00000            
REMARK 290                                                                      
REMARK 290 REMARK: NULL                                                         
REMARK 300                                                                      
REMARK 300 BIOMOLECULE: 1, 2                                                    
REMARK 300 THIS ENTRY CONTAINS THE CRYSTALLOGRAPHIC ASYMMETRIC UNIT             
REMARK 300 WHICH CONSISTS OF 6 CHAIN(S). SEE REMARK 350 FOR                     
REMARK 300 INFORMATION ON GENERATING THE BIOLOGICAL MOLECULE(S).                
REMARK 350                                                                      
REMARK 350 GENERATING THE BIOMOLECULE                                           
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN           
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE                
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS          
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND                          
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.                               
REMARK 350                                                                      
REMARK 350 BIOMOLECULE: 1                                                       
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B, C                               
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 350 BIOMOLECULE: 2                                                       
REMARK 350 APPLY THE FOLLOWING TO CHAINS: D, E, F                               
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 465                                                                      
REMARK 465 MISSING RESIDUES                                                     
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE                       
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)                
REMARK 465                                                                      
REMARK 465   M RES C SSSEQI                                                     
REMARK 465     ILE B    16                                                      
REMARK 465     VAL B    17                                                      
REMARK 465     ARG B    18                                                      
REMARK 465     GLY C   263                                                      
REMARK 465     ASN C   264                                                      
REMARK 465     LYS C   265                                                      
REMARK 465     ALA C   431                                                      
REMARK 465     GLY D   261                                                      
REMARK 465     GLU D   262                                                      
REMARK 465     GLY D   263                                                      
REMARK 465     ALA D   431                                                      
REMARK 465     ILE E    16                                                      
REMARK 465     VAL E    17                                                      
REMARK 465     ARG E    18                                                      
REMARK 470                                                                      
REMARK 470 MISSING ATOM                                                         
REMARK 470 THE FOLLOWING RESIDUES HAVE MISSING ATOMS(M=MODEL NUMBER;            
REMARK 470 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;          
REMARK 470 I=INSERTION CODE):                                                   
REMARK 470   M RES CSSEQI  ATOMS                                                
REMARK 470     LYS A   2    CG    CD    CE    NZ                                
REMARK 470     LYS A   5    CG    CD    CE    NZ                                
REMARK 470     GLU A 235    CG    CD    OE1   OE2                               
REMARK 470     LYS A 256    CG    CD    CE    NZ                                
REMARK 470     ASN A 264    CG    OD1   ND2                                     
REMARK 470     LYS A 265    CG    CD    CE    NZ                                
REMARK 470     GLN A 282    CG    CD    OE1   NE2                               
REMARK 470     LYS A 310    CG    CD    CE    NZ                                
REMARK 470     LYS A 324    CG    CD    CE    NZ                                
REMARK 470     GLU A 331    CG    CD    OE1   OE2                               
REMARK 470     LYS C 305    CG    CD    CE    NZ                                
REMARK 470     GLU C 417    CG    CD    OE1   OE2                               
REMARK 470     GLU D 100    CG    CD    OE1   OE2                               
REMARK 470     ASP D 160    CG    OD1   OD2                                     
REMARK 470     ASN D 162    CG    OD1   ND2                                     
REMARK 470     LYS D 197    CG    CD    CE    NZ                                
REMARK 470     LYS D 253    CG    CD    CE    NZ                                
REMARK 470     ASP D 254    CG    OD1   OD2                                     
REMARK 470     ASN D 264    CG    OD1   ND2                                     
REMARK 470     GLU D 278    CG    CD    OE1   OE2                               
REMARK 470     LYS D 305    CG    CD    CE    NZ                                
REMARK 470     LYS D 324    CG    CD    CE    NZ                                
REMARK 470     GLU D 417    CG    CD    OE1   OE2                               
REMARK 470     LYS F   2    CG    CD    CE    NZ                                
REMARK 470     GLN F  76    CG    CD    OE1   NE2                               
REMARK 470     LYS F 265    CG    CD    CE    NZ                                
REMARK 470     LYS F 310    CG    CD    CE    NZ                                
REMARK 470     LYS F 324    CG    CD    CE    NZ                                
REMARK 470     LYS F 359    CG    CD    CE    NZ                                
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT                     
REMARK 500                                                                      
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.                            
REMARK 500                                                                      
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI                             
REMARK 500   OG   SER F   249     O    HOH    1932              1.93            
REMARK 500   O    HOH     410     O    HOH    1940              1.94            
REMARK 500   O    HOH     485     O    HOH    1934              1.98            
REMARK 500   O    HOH    1236     O    HOH    1933              2.11            
REMARK 500   O    HOH     324     O    HOH    1938              2.16            
REMARK 500   O    HOH     941     O    HOH    1937              2.17            
REMARK 500   O    HOH     303     O    HOH    1926              2.18            
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND LENGTHS                                      
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,2(A3,1X,A1,I4,A1,1X,A4,3X),F6.3)                  
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   RES CSSEQI ATM2   DEVIATION                     
REMARK 500    GLU C 375   CB    GLU C 375   CG     0.060                        
REMARK 500    GLU D 375   CB    GLU D 375   CG     0.076                        
REMARK 500    PRO F  18   CB    PRO F  18   CG    -0.062                        
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES                                       
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)              
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3                                     
REMARK 500    GLU A 375   CA  -  CB  -  CG  ANGL. DEV. =  7.6 DEGREES           
REMARK 500    LYS C 392   N   -  CA  -  C   ANGL. DEV. = -7.2 DEGREES           
REMARK 500    LEU F 236   CA  -  CB  -  CG  ANGL. DEV. =  7.2 DEGREES           
REMARK 500    LEU F 236   CB  -  CG  -  CD2 ANGL. DEV. =  7.4 DEGREES           
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: TORSION ANGLES                                             
REMARK 500                                                                      
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:            
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;               
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                             
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)                    
REMARK 500                                                                      
REMARK 500  M RES CSSEQI        PSI       PHI                                   
REMARK 500    ARG A 398      123.79     78.01                                   
REMARK 500    ARG C 398      123.72     75.67                                   
REMARK 500    ARG D 398      123.00     75.87                                   
REMARK 500    ARG F 398      126.88     75.30                                   
REMARK 525                                                                      
REMARK 525 SOLVENT                                                              
REMARK 525 THE FOLLOWING SOLVENT MOLECULES LIE FARTHER THAN EXPECTED            
REMARK 525 FROM THE PROTEIN OR NUCLEIC ACID MOLECULE AND MAY BE                 
REMARK 525 ASSOCIATED WITH A SYMMETRY RELATED MOLECULE (M=MODEL                 
REMARK 525 NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE          
REMARK 525 NUMBER; I=INSERTION CODE):                                           
REMARK 525                                                                      
REMARK 525  M RES CSSEQI                                                        
REMARK 525    HOH   748        DISTANCE =  5.72 ANGSTROMS                       
REMARK 525    HOH   807        DISTANCE =  5.26 ANGSTROMS                       
REMARK 900                                                                      
REMARK 900 RELATED ENTRIES                                                      
REMARK 900 RELATED ID: 1E9I   RELATED DB: PDB                                   
REMARK 900 THE SAME PROTEIN COMPLEXED WITH MINIMAL BINDING PEPTIDE              
REMARK 900 FROM RNASE E.                                                        
DBREF  2FYM A    1   431  UNP    P0A6P9   ENO_ECOLI        1    431             
DBREF  2FYM C    1   431  UNP    P0A6P9   ENO_ECOLI        1    431             
DBREF  2FYM D    1   431  UNP    P0A6P9   ENO_ECOLI        1    431             
DBREF  2FYM F    1   431  UNP    P0A6P9   ENO_ECOLI        1    431             
DBREF  2FYM B    1    18  UNP    P21513   RNE_ECOLI      833    850             
DBREF  2FYM E    1    18  UNP    P21513   RNE_ECOLI      833    850             
SEQRES   1 A  431  SER LYS ILE VAL LYS ILE ILE GLY ARG GLU ILE ILE ASP          
SEQRES   2 A  431  SER ARG GLY ASN PRO THR VAL GLU ALA GLU VAL HIS LEU          
SEQRES   3 A  431  GLU GLY GLY PHE VAL GLY MET ALA ALA ALA PRO SER GLY          
SEQRES   4 A  431  ALA SER THR GLY SER ARG GLU ALA LEU GLU LEU ARG ASP          
SEQRES   5 A  431  GLY ASP LYS SER ARG PHE LEU GLY LYS GLY VAL THR LYS          
SEQRES   6 A  431  ALA VAL ALA ALA VAL ASN GLY PRO ILE ALA GLN ALA LEU          
SEQRES   7 A  431  ILE GLY LYS ASP ALA LYS ASP GLN ALA GLY ILE ASP LYS          
SEQRES   8 A  431  ILE MET ILE ASP LEU ASP GLY THR GLU ASN LYS SER LYS          
SEQRES   9 A  431  PHE GLY ALA ASN ALA ILE LEU ALA VAL SER LEU ALA ASN          
SEQRES  10 A  431  ALA LYS ALA ALA ALA ALA ALA LYS GLY MET PRO LEU TYR          
SEQRES  11 A  431  GLU HIS ILE ALA GLU LEU ASN GLY THR PRO GLY LYS TYR          
SEQRES  12 A  431  SER MET PRO VAL PRO MET MET ASN ILE ILE ASN GLY GLY          
SEQRES  13 A  431  GLU HIS ALA ASP ASN ASN VAL ASP ILE GLN GLU PHE MET          
SEQRES  14 A  431  ILE GLN PRO VAL GLY ALA LYS THR VAL LYS GLU ALA ILE          
SEQRES  15 A  431  ARG MET GLY SER GLU VAL PHE HIS HIS LEU ALA LYS VAL          
SEQRES  16 A  431  LEU LYS ALA LYS GLY MET ASN THR ALA VAL GLY ASP GLU          
SEQRES  17 A  431  GLY GLY TYR ALA PRO ASN LEU GLY SER ASN ALA GLU ALA          
SEQRES  18 A  431  LEU ALA VAL ILE ALA GLU ALA VAL LYS ALA ALA GLY TYR          
SEQRES  19 A  431  GLU LEU GLY LYS ASP ILE THR LEU ALA MET ASP CYS ALA          
SEQRES  20 A  431  ALA SER GLU PHE TYR LYS ASP GLY LYS TYR VAL LEU ALA          
SEQRES  21 A  431  GLY GLU GLY ASN LYS ALA PHE THR SER GLU GLU PHE THR          
SEQRES  22 A  431  HIS PHE LEU GLU GLU LEU THR LYS GLN TYR PRO ILE VAL          
SEQRES  23 A  431  SER ILE GLU ASP GLY LEU ASP GLU SER ASP TRP ASP GLY          
SEQRES  24 A  431  PHE ALA TYR GLN THR LYS VAL LEU GLY ASP LYS ILE GLN          
SEQRES  25 A  431  LEU VAL GLY ASP ASP LEU PHE VAL THR ASN THR LYS ILE          
SEQRES  26 A  431  LEU LYS GLU GLY ILE GLU LYS GLY ILE ALA ASN SER ILE          
SEQRES  27 A  431  LEU ILE LYS PHE ASN GLN ILE GLY SER LEU THR GLU THR          
SEQRES  28 A  431  LEU ALA ALA ILE LYS MET ALA LYS ASP ALA GLY TYR THR          
SEQRES  29 A  431  ALA VAL ILE SER HIS ARG SER GLY GLU THR GLU ASP ALA          
SEQRES  30 A  431  THR ILE ALA ASP LEU ALA VAL GLY THR ALA ALA GLY GLN          
SEQRES  31 A  431  ILE LYS THR GLY SER MET SER ARG SER ASP ARG VAL ALA          
SEQRES  32 A  431  LYS TYR ASN GLN LEU ILE ARG ILE GLU GLU ALA LEU GLY          
SEQRES  33 A  431  GLU LYS ALA PRO TYR ASN GLY ARG LYS GLU ILE LYS GLY          
SEQRES  34 A  431  GLN ALA                                                      
SEQRES   1 B   18  ALA SER PRO GLU LEU ALA SER GLY LYS VAL TRP ILE ARG          
SEQRES   2 B   18  TYR PRO ILE VAL ARG                                          
SEQRES   1 C  431  SER LYS ILE VAL LYS ILE ILE GLY ARG GLU ILE ILE ASP          
SEQRES   2 C  431  SER ARG GLY ASN PRO THR VAL GLU ALA GLU VAL HIS LEU          
SEQRES   3 C  431  GLU GLY GLY PHE VAL GLY MET ALA ALA ALA PRO SER GLY          
SEQRES   4 C  431  ALA SER THR GLY SER ARG GLU ALA LEU GLU LEU ARG ASP          
SEQRES   5 C  431  GLY ASP LYS SER ARG PHE LEU GLY LYS GLY VAL THR LYS          
SEQRES   6 C  431  ALA VAL ALA ALA VAL ASN GLY PRO ILE ALA GLN ALA LEU          
SEQRES   7 C  431  ILE GLY LYS ASP ALA LYS ASP GLN ALA GLY ILE ASP LYS          
SEQRES   8 C  431  ILE MET ILE ASP LEU ASP GLY THR GLU ASN LYS SER LYS          
SEQRES   9 C  431  PHE GLY ALA ASN ALA ILE LEU ALA VAL SER LEU ALA ASN          
SEQRES  10 C  431  ALA LYS ALA ALA ALA ALA ALA LYS GLY MET PRO LEU TYR          
SEQRES  11 C  431  GLU HIS ILE ALA GLU LEU ASN GLY THR PRO GLY LYS TYR          
SEQRES  12 C  431  SER MET PRO VAL PRO MET MET ASN ILE ILE ASN GLY GLY          
SEQRES  13 C  431  GLU HIS ALA ASP ASN ASN VAL ASP ILE GLN GLU PHE MET          
SEQRES  14 C  431  ILE GLN PRO VAL GLY ALA LYS THR VAL LYS GLU ALA ILE          
SEQRES  15 C  431  ARG MET GLY SER GLU VAL PHE HIS HIS LEU ALA LYS VAL          
SEQRES  16 C  431  LEU LYS ALA LYS GLY MET ASN THR ALA VAL GLY ASP GLU          
SEQRES  17 C  431  GLY GLY TYR ALA PRO ASN LEU GLY SER ASN ALA GLU ALA          
SEQRES  18 C  431  LEU ALA VAL ILE ALA GLU ALA VAL LYS ALA ALA GLY TYR          
SEQRES  19 C  431  GLU LEU GLY LYS ASP ILE THR LEU ALA MET ASP CYS ALA          
SEQRES  20 C  431  ALA SER GLU PHE TYR LYS ASP GLY LYS TYR VAL LEU ALA          
SEQRES  21 C  431  GLY GLU GLY ASN LYS ALA PHE THR SER GLU GLU PHE THR          
SEQRES  22 C  431  HIS PHE LEU GLU GLU LEU THR LYS GLN TYR PRO ILE VAL          
SEQRES  23 C  431  SER ILE GLU ASP GLY LEU ASP GLU SER ASP TRP ASP GLY          
SEQRES  24 C  431  PHE ALA TYR GLN THR LYS VAL LEU GLY ASP LYS ILE GLN          
SEQRES  25 C  431  LEU VAL GLY ASP ASP LEU PHE VAL THR ASN THR LYS ILE          
SEQRES  26 C  431  LEU LYS GLU GLY ILE GLU LYS GLY ILE ALA ASN SER ILE          
SEQRES  27 C  431  LEU ILE LYS PHE ASN GLN ILE GLY SER LEU THR GLU THR          
SEQRES  28 C  431  LEU ALA ALA ILE LYS MET ALA LYS ASP ALA GLY TYR THR          
SEQRES  29 C  431  ALA VAL ILE SER HIS ARG SER GLY GLU THR GLU ASP ALA          
SEQRES  30 C  431  THR ILE ALA ASP LEU ALA VAL GLY THR ALA ALA GLY GLN          
SEQRES  31 C  431  ILE LYS THR GLY SER MET SER ARG SER ASP ARG VAL ALA          
SEQRES  32 C  431  LYS TYR ASN GLN LEU ILE ARG ILE GLU GLU ALA LEU GLY          
SEQRES  33 C  431  GLU LYS ALA PRO TYR ASN GLY ARG LYS GLU ILE LYS GLY          
SEQRES  34 C  431  GLN ALA                                                      
SEQRES   1 D  431  SER LYS ILE VAL LYS ILE ILE GLY ARG GLU ILE ILE ASP          
SEQRES   2 D  431  SER ARG GLY ASN PRO THR VAL GLU ALA GLU VAL HIS LEU          
SEQRES   3 D  431  GLU GLY GLY PHE VAL GLY MET ALA ALA ALA PRO SER GLY          
SEQRES   4 D  431  ALA SER THR GLY SER ARG GLU ALA LEU GLU LEU ARG ASP          
SEQRES   5 D  431  GLY ASP LYS SER ARG PHE LEU GLY LYS GLY VAL THR LYS          
SEQRES   6 D  431  ALA VAL ALA ALA VAL ASN GLY PRO ILE ALA GLN ALA LEU          
SEQRES   7 D  431  ILE GLY LYS ASP ALA LYS ASP GLN ALA GLY ILE ASP LYS          
SEQRES   8 D  431  ILE MET ILE ASP LEU ASP GLY THR GLU ASN LYS SER LYS          
SEQRES   9 D  431  PHE GLY ALA ASN ALA ILE LEU ALA VAL SER LEU ALA ASN          
SEQRES  10 D  431  ALA LYS ALA ALA ALA ALA ALA LYS GLY MET PRO LEU TYR          
SEQRES  11 D  431  GLU HIS ILE ALA GLU LEU ASN GLY THR PRO GLY LYS TYR          
SEQRES  12 D  431  SER MET PRO VAL PRO MET MET ASN ILE ILE ASN GLY GLY          
SEQRES  13 D  431  GLU HIS ALA ASP ASN ASN VAL ASP ILE GLN GLU PHE MET          
SEQRES  14 D  431  ILE GLN PRO VAL GLY ALA LYS THR VAL LYS GLU ALA ILE          
SEQRES  15 D  431  ARG MET GLY SER GLU VAL PHE HIS HIS LEU ALA LYS VAL          
SEQRES  16 D  431  LEU LYS ALA LYS GLY MET ASN THR ALA VAL GLY ASP GLU          
SEQRES  17 D  431  GLY GLY TYR ALA PRO ASN LEU GLY SER ASN ALA GLU ALA          
SEQRES  18 D  431  LEU ALA VAL ILE ALA GLU ALA VAL LYS ALA ALA GLY TYR          
SEQRES  19 D  431  GLU LEU GLY LYS ASP ILE THR LEU ALA MET ASP CYS ALA          
SEQRES  20 D  431  ALA SER GLU PHE TYR LYS ASP GLY LYS TYR VAL LEU ALA          
SEQRES  21 D  431  GLY GLU GLY ASN LYS ALA PHE THR SER GLU GLU PHE THR          
SEQRES  22 D  431  HIS PHE LEU GLU GLU LEU THR LYS GLN TYR PRO ILE VAL          
SEQRES  23 D  431  SER ILE GLU ASP GLY LEU ASP GLU SER ASP TRP ASP GLY          
SEQRES  24 D  431  PHE ALA TYR GLN THR LYS VAL LEU GLY ASP LYS ILE GLN          
SEQRES  25 D  431  LEU VAL GLY ASP ASP LEU PHE VAL THR ASN THR LYS ILE          
SEQRES  26 D  431  LEU LYS GLU GLY ILE GLU LYS GLY ILE ALA ASN SER ILE          
SEQRES  27 D  431  LEU ILE LYS PHE ASN GLN ILE GLY SER LEU THR GLU THR          
SEQRES  28 D  431  LEU ALA ALA ILE LYS MET ALA LYS ASP ALA GLY TYR THR          
SEQRES  29 D  431  ALA VAL ILE SER HIS ARG SER GLY GLU THR GLU ASP ALA          
SEQRES  30 D  431  THR ILE ALA ASP LEU ALA VAL GLY THR ALA ALA GLY GLN          
SEQRES  31 D  431  ILE LYS THR GLY SER MET SER ARG SER ASP ARG VAL ALA          
SEQRES  32 D  431  LYS TYR ASN GLN LEU ILE ARG ILE GLU GLU ALA LEU GLY          
SEQRES  33 D  431  GLU LYS ALA PRO TYR ASN GLY ARG LYS GLU ILE LYS GLY          
SEQRES  34 D  431  GLN ALA                                                      
SEQRES   1 E   18  ALA SER PRO GLU LEU ALA SER GLY LYS VAL TRP ILE ARG          
SEQRES   2 E   18  TYR PRO ILE VAL ARG                                          
SEQRES   1 F  431  SER LYS ILE VAL LYS ILE ILE GLY ARG GLU ILE ILE ASP          
SEQRES   2 F  431  SER ARG GLY ASN PRO THR VAL GLU ALA GLU VAL HIS LEU          
SEQRES   3 F  431  GLU GLY GLY PHE VAL GLY MET ALA ALA ALA PRO SER GLY          
SEQRES   4 F  431  ALA SER THR GLY SER ARG GLU ALA LEU GLU LEU ARG ASP          
SEQRES   5 F  431  GLY ASP LYS SER ARG PHE LEU GLY LYS GLY VAL THR LYS          
SEQRES   6 F  431  ALA VAL ALA ALA VAL ASN GLY PRO ILE ALA GLN ALA LEU          
SEQRES   7 F  431  ILE GLY LYS ASP ALA LYS ASP GLN ALA GLY ILE ASP LYS          
SEQRES   8 F  431  ILE MET ILE ASP LEU ASP GLY THR GLU ASN LYS SER LYS          
SEQRES   9 F  431  PHE GLY ALA ASN ALA ILE LEU ALA VAL SER LEU ALA ASN          
SEQRES  10 F  431  ALA LYS ALA ALA ALA ALA ALA LYS GLY MET PRO LEU TYR          
SEQRES  11 F  431  GLU HIS ILE ALA GLU LEU ASN GLY THR PRO GLY LYS TYR          
SEQRES  12 F  431  SER MET PRO VAL PRO MET MET ASN ILE ILE ASN GLY GLY          
SEQRES  13 F  431  GLU HIS ALA ASP ASN ASN VAL ASP ILE GLN GLU PHE MET          
SEQRES  14 F  431  ILE GLN PRO VAL GLY ALA LYS THR VAL LYS GLU ALA ILE          
SEQRES  15 F  431  ARG MET GLY SER GLU VAL PHE HIS HIS LEU ALA LYS VAL          
SEQRES  16 F  431  LEU LYS ALA LYS GLY MET ASN THR ALA VAL GLY ASP GLU          
SEQRES  17 F  431  GLY GLY TYR ALA PRO ASN LEU GLY SER ASN ALA GLU ALA          
SEQRES  18 F  431  LEU ALA VAL ILE ALA GLU ALA VAL LYS ALA ALA GLY TYR          
SEQRES  19 F  431  GLU LEU GLY LYS ASP ILE THR LEU ALA MET ASP CYS ALA          
SEQRES  20 F  431  ALA SER GLU PHE TYR LYS ASP GLY LYS TYR VAL LEU ALA          
SEQRES  21 F  431  GLY GLU GLY ASN LYS ALA PHE THR SER GLU GLU PHE THR          
SEQRES  22 F  431  HIS PHE LEU GLU GLU LEU THR LYS GLN TYR PRO ILE VAL          
SEQRES  23 F  431  SER ILE GLU ASP GLY LEU ASP GLU SER ASP TRP ASP GLY          
SEQRES  24 F  431  PHE ALA TYR GLN THR LYS VAL LEU GLY ASP LYS ILE GLN          
SEQRES  25 F  431  LEU VAL GLY ASP ASP LEU PHE VAL THR ASN THR LYS ILE          
SEQRES  26 F  431  LEU LYS GLU GLY ILE GLU LYS GLY ILE ALA ASN SER ILE          
SEQRES  27 F  431  LEU ILE LYS PHE ASN GLN ILE GLY SER LEU THR GLU THR          
SEQRES  28 F  431  LEU ALA ALA ILE LYS MET ALA LYS ASP ALA GLY TYR THR          
SEQRES  29 F  431  ALA VAL ILE SER HIS ARG SER GLY GLU THR GLU ASP ALA          
SEQRES  30 F  431  THR ILE ALA ASP LEU ALA VAL GLY THR ALA ALA GLY GLN          
SEQRES  31 F  431  ILE LYS THR GLY SER MET SER ARG SER ASP ARG VAL ALA          
SEQRES  32 F  431  LYS TYR ASN GLN LEU ILE ARG ILE GLU GLU ALA LEU GLY          
SEQRES  33 F  431  GLU LYS ALA PRO TYR ASN GLY ARG LYS GLU ILE LYS GLY          
SEQRES  34 F  431  GLN ALA                                                      
HET     MG  A1431       1                                                       
HET     MG  C1431       1                                                       
HET     MG  D1431       1                                                       
HET     MG  F1431       1                                                       
HETNAM      MG MAGNESIUM ION                                                    
FORMUL   7   MG    4(MG 2+)                                                     
FORMUL  11  HOH   *1930(H2 O)                                                   
HELIX    1   1 ARG A   57  LYS A   61  5                                   5    
HELIX    2   2 VAL A   63  GLY A   72  1                                  10    
HELIX    3   3 GLY A   72  ILE A   79  1                                   8    
HELIX    4   4 ASP A   85  GLY A   98  1                                  14    
HELIX    5   5 GLY A  106  LYS A  125  1                                  20    
HELIX    6   6 PRO A  128  ASN A  137  1                                  10    
HELIX    7   7 GLY A  156  ALA A  159  5                                   4    
HELIX    8   8 THR A  177  LYS A  199  1                                  23    
HELIX    9   9 SER A  217  ALA A  232  1                                  16    
HELIX   10  10 ALA A  247  GLU A  250  5                                   4    
HELIX   11  11 ALA A  260  GLY A  263  5                                   4    
HELIX   12  12 THR A  268  TYR A  283  1                                  16    
HELIX   13  13 ASP A  296  GLY A  308  1                                  13    
HELIX   14  14 ASN A  322  LYS A  332  1                                  11    
HELIX   15  15 LYS A  341  ILE A  345  5                                   5    
HELIX   16  16 SER A  347  ALA A  361  1                                  15    
HELIX   17  17 ALA A  377  THR A  386  1                                  10    
HELIX   18  18 ARG A  398  GLY A  416  1                                  19    
HELIX   19  19 GLU A  417  ALA A  419  5                                   3    
HELIX   20  20 ASN A  422  ILE A  427  5                                   6    
HELIX   21  21 SER B    2  SER B    7  1                                   6    
HELIX   22  22 ARG C   57  LYS C   61  5                                   5    
HELIX   23  23 VAL C   63  GLY C   72  1                                  10    
HELIX   24  24 GLY C   72  ILE C   79  1                                   8    
HELIX   25  25 ASP C   85  GLY C   98  1                                  14    
HELIX   26  26 GLY C  106  LYS C  125  1                                  20    
HELIX   27  27 PRO C  128  GLY C  138  1                                  11    
HELIX   28  28 GLY C  156  ALA C  159  5                                   4    
HELIX   29  29 THR C  177  LYS C  199  1                                  23    
HELIX   30  30 SER C  217  ALA C  232  1                                  16    
HELIX   31  31 ALA C  247  GLU C  250  5                                   4    
HELIX   32  32 THR C  268  TYR C  283  1                                  16    
HELIX   33  33 ASP C  296  GLY C  308  1                                  13    
HELIX   34  34 ASN C  322  GLY C  333  1                                  12    
HELIX   35  35 LYS C  341  GLY C  346  1                                   6    
HELIX   36  36 SER C  347  ALA C  361  1                                  15    
HELIX   37  37 ALA C  377  THR C  386  1                                  10    
HELIX   38  38 ARG C  398  GLY C  416  1                                  19    
HELIX   39  39 GLU C  417  ALA C  419  5                                   3    
HELIX   40  40 ASN C  422  ILE C  427  5                                   6    
HELIX   41  41 ARG D   57  LYS D   61  5                                   5    
HELIX   42  42 VAL D   63  GLY D   72  1                                  10    
HELIX   43  43 GLY D   72  ILE D   79  1                                   8    
HELIX   44  44 ASP D   85  GLY D   98  1                                  14    
HELIX   45  45 GLY D  106  LYS D  125  1                                  20    
HELIX   46  46 PRO D  128  ASN D  137  1                                  10    
HELIX   47  47 GLY D  156  ALA D  159  5                                   4    
HELIX   48  48 THR D  177  LYS D  199  1                                  23    
HELIX   49  49 SER D  217  ALA D  232  1                                  16    
HELIX   50  50 ALA D  247  GLU D  250  5                                   4    
HELIX   51  51 THR D  268  TYR D  283  1                                  16    
HELIX   52  52 ASP D  296  GLY D  308  1                                  13    
HELIX   53  53 ASN D  322  LYS D  332  1                                  11    
HELIX   54  54 LYS D  341  GLY D  346  1                                   6    
HELIX   55  55 SER D  347  GLY D  362  1                                  16    
HELIX   56  56 ALA D  377  THR D  386  1                                  10    
HELIX   57  57 ARG D  398  GLY D  416  1                                  19    
HELIX   58  58 GLU D  417  ALA D  419  5                                   3    
HELIX   59  59 ASN D  422  ILE D  427  5                                   6    
HELIX   60  60 SER E    2  SER E    7  1                                   6    
HELIX   61  61 ARG F   57  LYS F   61  5                                   5    
HELIX   62  62 VAL F   63  GLY F   72  1                                  10    
HELIX   63  63 GLY F   72  ILE F   79  1                                   8    
HELIX   64  64 ASP F   85  GLY F   98  1                                  14    
HELIX   65  65 GLY F  106  LYS F  125  1                                  20    
HELIX   66  66 PRO F  128  ASN F  137  1                                  10    
HELIX   67  67 GLY F  156  ALA F  159  5                                   4    
HELIX   68  68 THR F  177  LYS F  199  1                                  23    
HELIX   69  69 SER F  217  ALA F  232  1                                  16    
HELIX   70  70 ALA F  247  GLU F  250  5                                   4    
HELIX   71  71 THR F  268  TYR F  283  1                                  16    
HELIX   72  72 ASP F  296  GLY F  308  1                                  13    
HELIX   73  73 ASN F  322  LYS F  332  1                                  11    
HELIX   74  74 LYS F  341  GLY F  346  1                                   6    
HELIX   75  75 SER F  347  ALA F  361  1                                  15    
HELIX   76  76 ALA F  377  THR F  386  1                                  10    
HELIX   77  77 ARG F  398  GLY F  416  1                                  19    
HELIX   78  78 GLU F  417  ALA F  419  5                                   3    
HELIX   79  79 ASN F  422  ILE F  427  5                                   6    
SHEET    1   A 3 ILE A   3  ILE A  12  0                                        
SHEET    2   A 3 PRO A  18  LEU A  26 -1  O  HIS A  25   N  LYS A   5           
SHEET    3   A 3 VAL A  31  ALA A  35 -1  O  ALA A  34   N  ALA A  22           
SHEET    1   B 9 VAL A 147  PRO A 148  0                                        
SHEET    2   B 9 GLY A 389  LYS A 392  1  O  GLY A 389   N  VAL A 147           
SHEET    3   B 9 THR A 364  SER A 368  1  N  ILE A 367   O  LYS A 392           
SHEET    4   B 9 SER A 337  ILE A 340  1  N  ILE A 340   O  VAL A 366           
SHEET    5   B 9 GLN A 312  GLY A 315  1  N  GLY A 315   O  LEU A 339           
SHEET    6   B 9 ILE A 285  GLU A 289  1  N  ILE A 288   O  GLN A 312           
SHEET    7   B 9 THR A 241  ASP A 245  1  N  MET A 244   O  GLU A 289           
SHEET    8   B 9 GLU A 167  GLN A 171 -1  N  MET A 169   O  ALA A 243           
SHEET    9   B 9 MET A 150  ASN A 154 -1  N  MET A 150   O  ILE A 170           
SHEET    1   C 3 TYR A 252  LYS A 253  0                                        
SHEET    2   C 3 LYS A 256  VAL A 258 -1  O  LYS A 256   N  LYS A 253           
SHEET    3   C 3 ALA A 266  PHE A 267 -1  O  PHE A 267   N  TYR A 257           
SHEET    1   D 3 ILE C   3  ILE C  12  0                                        
SHEET    2   D 3 PRO C  18  LEU C  26 -1  O  HIS C  25   N  LYS C   5           
SHEET    3   D 3 VAL C  31  ALA C  35 -1  O  ALA C  34   N  ALA C  22           
SHEET    1   E 9 VAL C 147  PRO C 148  0                                        
SHEET    2   E 9 GLY C 389  LYS C 392  1  O  GLY C 389   N  VAL C 147           
SHEET    3   E 9 THR C 364  SER C 368  1  N  ILE C 367   O  LYS C 392           
SHEET    4   E 9 SER C 337  ILE C 340  1  N  ILE C 340   O  VAL C 366           
SHEET    5   E 9 GLN C 312  GLY C 315  1  N  GLY C 315   O  LEU C 339           
SHEET    6   E 9 ILE C 285  GLU C 289  1  N  ILE C 288   O  GLN C 312           
SHEET    7   E 9 THR C 241  ASP C 245  1  N  MET C 244   O  GLU C 289           
SHEET    8   E 9 GLU C 167  GLN C 171 -1  N  MET C 169   O  ALA C 243           
SHEET    9   E 9 MET C 150  ASN C 154 -1  N  MET C 150   O  ILE C 170           
SHEET    1   F 2 TYR C 252  LYS C 253  0                                        
SHEET    2   F 2 LYS C 256  TYR C 257 -1  O  LYS C 256   N  LYS C 253           
SHEET    1   G 3 ILE D   3  ILE D  12  0                                        
SHEET    2   G 3 PRO D  18  LEU D  26 -1  O  HIS D  25   N  LYS D   5           
SHEET    3   G 3 VAL D  31  ALA D  35 -1  O  ALA D  34   N  ALA D  22           
SHEET    1   H 9 VAL D 147  PRO D 148  0                                        
SHEET    2   H 9 GLY D 389  LYS D 392  1  O  GLY D 389   N  VAL D 147           
SHEET    3   H 9 THR D 364  SER D 368  1  N  ILE D 367   O  LYS D 392           
SHEET    4   H 9 SER D 337  ILE D 340  1  N  ILE D 340   O  SER D 368           
SHEET    5   H 9 GLN D 312  GLY D 315  1  N  GLY D 315   O  LEU D 339           
SHEET    6   H 9 ILE D 285  GLU D 289  1  N  ILE D 288   O  GLN D 312           
SHEET    7   H 9 THR D 241  ASP D 245  1  N  LEU D 242   O  VAL D 286           
SHEET    8   H 9 GLU D 167  GLN D 171 -1  N  MET D 169   O  ALA D 243           
SHEET    9   H 9 MET D 150  ASN D 154 -1  N  MET D 150   O  ILE D 170           
SHEET    1   I 3 TYR D 252  LYS D 253  0                                        
SHEET    2   I 3 LYS D 256  LEU D 259 -1  O  LYS D 256   N  LYS D 253           
SHEET    3   I 3 LYS D 265  PHE D 267 -1  O  PHE D 267   N  TYR D 257           
SHEET    1   J 3 ILE F   3  ILE F  12  0                                        
SHEET    2   J 3 PRO F  18  LEU F  26 -1  O  HIS F  25   N  VAL F   4           
SHEET    3   J 3 VAL F  31  ALA F  35 -1  O  ALA F  34   N  ALA F  22           
SHEET    1   K 9 VAL F 147  PRO F 148  0                                        
SHEET    2   K 9 GLY F 389  LYS F 392  1  O  GLY F 389   N  VAL F 147           
SHEET    3   K 9 THR F 364  SER F 368  1  N  ILE F 367   O  LYS F 392           
SHEET    4   K 9 SER F 337  ILE F 340  1  N  ILE F 340   O  VAL F 366           
SHEET    5   K 9 GLN F 312  GLY F 315  1  N  GLY F 315   O  SER F 337           
SHEET    6   K 9 ILE F 285  GLU F 289  1  N  ILE F 288   O  GLN F 312           
SHEET    7   K 9 THR F 241  ASP F 245  1  N  LEU F 242   O  VAL F 286           
SHEET    8   K 9 GLU F 167  GLN F 171 -1  N  MET F 169   O  ALA F 243           
SHEET    9   K 9 MET F 150  ASN F 154 -1  N  MET F 150   O  ILE F 170           
SHEET    1   L 3 TYR F 252  LYS F 253  0                                        
SHEET    2   L 3 LYS F 256  LEU F 259 -1  O  LYS F 256   N  LYS F 253           
SHEET    3   L 3 LYS F 265  PHE F 267 -1  O  PHE F 267   N  TYR F 257           
LINK        MG    MG A1431                 OD2 ASP A 245                        
LINK        MG    MG A1431                 OE2 GLU A 289                        
LINK        MG    MG A1431                 OD2 ASP A 316                        
LINK        MG    MG C1431                 OD2 ASP C 245                        
LINK        MG    MG C1431                 OE2 GLU C 289                        
LINK        MG    MG C1431                 OD2 ASP C 316                        
LINK        MG    MG D1431                 OD2 ASP D 245                        
LINK        MG    MG D1431                 OE2 GLU D 289                        
LINK        MG    MG D1431                 OD2 ASP D 316                        
LINK        MG    MG F1431                 OD2 ASP F 245                        
LINK        MG    MG F1431                 OE2 GLU F 289                        
LINK        MG    MG F1431                 OD2 ASP F 316                        
CRYST1   77.054  124.201   96.076  90.00  90.58  90.00 P 1 21 1      8          
ORIGX1      1.000000  0.000000  0.000000        0.00000                         
ORIGX2      0.000000  1.000000  0.000000        0.00000                         
ORIGX3      0.000000  0.000000  1.000000        0.00000                         
SCALE1      0.012978  0.000000  0.000131        0.00000                         
SCALE2      0.000000  0.008051  0.000000        0.00000                         
SCALE3      0.000000  0.000000  0.010409        0.00000