PDB entry 2a5y

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HEADER    APOPTOSIS                               01-JUL-05   2A5Y              
TITLE     STRUCTURE OF A CED-4/CED-9 COMPLEX                                    
COMPND    MOL_ID: 1;                                                            
COMPND   2 MOLECULE: APOPTOSIS REGULATOR CED-9;                                 
COMPND   3 CHAIN: A;                                                            
COMPND   4 FRAGMENT: RESIDUES 48-251;                                           
COMPND   5 SYNONYM: CELL DEATH PROTEIN 9;                                       
COMPND   6 ENGINEERED: YES;                                                     
COMPND   7 MUTATION: YES;                                                       
COMPND   8 MOL_ID: 2;                                                           
COMPND   9 MOLECULE: CED-4;                                                     
COMPND  10 CHAIN: B, C;                                                         
COMPND  11 ENGINEERED: YES                                                      
SOURCE    MOL_ID: 1;                                                            
SOURCE   2 ORGANISM_SCIENTIFIC: CAENORHABDITIS ELEGANS;                         
SOURCE   3 EXPRESSION_SYSTEM: ESCHERICHIA COLI;                                 
SOURCE   4 EXPRESSION_SYSTEM_COMMON: BACTERIA;                                  
SOURCE   5 EXPRESSION_SYSTEM_STRAIN: BL21(DE3);                                 
SOURCE   6 EXPRESSION_SYSTEM_VECTOR_TYPE: PLASMID;                              
SOURCE   7 EXPRESSION_SYSTEM_VECTOR: PBB75, PET-21B;                            
SOURCE   8 MOL_ID: 2;                                                           
SOURCE   9 ORGANISM_SCIENTIFIC: CAENORHABDITIS ELEGANS;                         
SOURCE  10 EXPRESSION_SYSTEM: ESCHERICHIA COLI;                                 
SOURCE  11 EXPRESSION_SYSTEM_COMMON: BACTERIA                                   
KEYWDS    CED-4, CED-9, CED-3 ACTIVATION, APOPTOSIS                             
EXPDTA    X-RAY DIFFRACTION                                                     
AUTHOR    N.YAN,Q.LIU,Q.HAO,L.GU,Y.SHI                                          
REVDAT   2   21-MAR-06 2A5Y    1       JRNL                                     
REVDAT   1   11-OCT-05 2A5Y    0                                                
JRNL        AUTH   N.YAN,J.CHAI,E.S.LEE,L.GU,Q.LIU,J.HE,J.W.WU,                 
JRNL        AUTH 2 D.KOKEL,H.LI,Q.HAO,D.XUE,Y.SHI                               
JRNL        TITL   STRUCTURE OF THE CED-4-CED-9 COMPLEX PROVIDES                
JRNL        TITL 2 INSIGHTS INTO PROGRAMMED CELL DEATH IN                       
JRNL        TITL 3 CAENORHABDITIS ELEGANS.                                      
JRNL        REF    NATURE                        V. 437   831 2005              
JRNL        REFN   ASTM NATUAS  UK ISSN 0028-0836                               
REMARK   1                                                                      
REMARK   2                                                                      
REMARK   2 RESOLUTION. 2.60 ANGSTROMS.                                          
REMARK   3                                                                      
REMARK   3 REFINEMENT.                                                          
REMARK   3   PROGRAM     : CNS                                                  
REMARK   3   AUTHORS     : BRUNGER,ADAMS,CLORE,DELANO,GROS,GROSSE-              
REMARK   3               : KUNSTLEVE,JIANG,KUSZEWSKI,NILGES, PANNU,             
REMARK   3               : READ,RICE,SIMONSON,WARREN                            
REMARK   3                                                                      
REMARK   3  REFINEMENT TARGET : ENGH & HUBER                                    
REMARK   3                                                                      
REMARK   3  DATA USED IN REFINEMENT.                                            
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.60                           
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 40.00                          
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.000                          
REMARK   3   DATA CUTOFF HIGH         (ABS(F)) : NULL                           
REMARK   3   DATA CUTOFF LOW          (ABS(F)) : NULL                           
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : NULL                           
REMARK   3   NUMBER OF REFLECTIONS             : 52166                          
REMARK   3                                                                      
REMARK   3  FIT TO DATA USED IN REFINEMENT.                                     
REMARK   3   CROSS-VALIDATION METHOD          : NULL                            
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM                          
REMARK   3   R VALUE            (WORKING SET) : 0.249                           
REMARK   3   FREE R VALUE                     : 0.277                           
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : NULL                            
REMARK   3   FREE R VALUE TEST SET COUNT      : 2433                            
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : NULL                            
REMARK   3                                                                      
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.                                  
REMARK   3   TOTAL NUMBER OF BINS USED           : NULL                         
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : NULL                         
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : NULL                         
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : NULL                         
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : NULL                         
REMARK   3   BIN R VALUE           (WORKING SET) : NULL                         
REMARK   3   BIN FREE R VALUE                    : NULL                         
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : NULL                         
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : NULL                         
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : NULL                         
REMARK   3                                                                      
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.                    
REMARK   3   PROTEIN ATOMS            : 8406                                    
REMARK   3   NUCLEIC ACID ATOMS       : 0                                       
REMARK   3   HETEROGEN ATOMS          : 64                                      
REMARK   3   SOLVENT ATOMS            : 260                                     
REMARK   3                                                                      
REMARK   3  B VALUES.                                                           
REMARK   3   FROM WILSON PLOT           (A**2) : NULL                           
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : NULL                           
REMARK   3   OVERALL ANISOTROPIC B VALUE.                                       
REMARK   3    B11 (A**2) : NULL                                                 
REMARK   3    B22 (A**2) : NULL                                                 
REMARK   3    B33 (A**2) : NULL                                                 
REMARK   3    B12 (A**2) : NULL                                                 
REMARK   3    B13 (A**2) : NULL                                                 
REMARK   3    B23 (A**2) : NULL                                                 
REMARK   3                                                                      
REMARK   3  ESTIMATED COORDINATE ERROR.                                         
REMARK   3   ESD FROM LUZZATI PLOT        (A) : NULL                            
REMARK   3   ESD FROM SIGMAA              (A) : NULL                            
REMARK   3   LOW RESOLUTION CUTOFF        (A) : NULL                            
REMARK   3                                                                      
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.                         
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : NULL                            
REMARK   3   ESD FROM C-V SIGMAA          (A) : NULL                            
REMARK   3                                                                      
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.                                   
REMARK   3   BOND LENGTHS                 (A) : 0.008                           
REMARK   3   BOND ANGLES            (DEGREES) : 1.49                            
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : NULL                            
REMARK   3   IMPROPER ANGLES        (DEGREES) : NULL                            
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL MODEL : NULL                                      
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA                
REMARK   3   MAIN-CHAIN BOND              (A**2) : NULL  ; NULL                 
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : NULL  ; NULL                 
REMARK   3   SIDE-CHAIN BOND              (A**2) : NULL  ; NULL                 
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : NULL  ; NULL                 
REMARK   3                                                                      
REMARK   3  BULK SOLVENT MODELING.                                              
REMARK   3   METHOD USED : NULL                                                 
REMARK   3   KSOL        : NULL                                                 
REMARK   3   BSOL        : NULL                                                 
REMARK   3                                                                      
REMARK   3  NCS MODEL : NULL                                                    
REMARK   3                                                                      
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT          
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL                 
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL                 
REMARK   3                                                                      
REMARK   3  PARAMETER FILE  1  : NULL                                           
REMARK   3  TOPOLOGY FILE  1   : NULL                                           
REMARK   3                                                                      
REMARK   3  OTHER REFINEMENT REMARKS: NULL                                      
REMARK   4                                                                      
REMARK   4 2A5Y COMPLIES WITH FORMAT V. 3.0, 1-DEC-2006                         
REMARK   4                                                                      
REMARK   4 THIS IS THE REMEDIATED VERSION OF THIS PDB ENTRY.                    
REMARK   4 REMEDIATED DATA FILE REVISION 3.100 (2007-03-17)                     
REMARK 100                                                                      
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB .                              
REMARK 100 THE RCSB ID CODE IS RCSB033528.                                      
REMARK 200                                                                      
REMARK 200 EXPERIMENTAL DETAILS                                                 
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION                  
REMARK 200  DATE OF DATA COLLECTION        : 01-JUL-2004; 30-SEP-2004           
REMARK 200  TEMPERATURE           (KELVIN) : 100.0                              
REMARK 200  PH                             : 7.50                               
REMARK 200  NUMBER OF CRYSTALS USED        : 1                                  
REMARK 200                                                                      
REMARK 200  SYNCHROTRON              (Y/N) : Y; Y                               
REMARK 200  RADIATION SOURCE               : NSLS; NSLS                         
REMARK 200  BEAMLINE                       : X25; X25                           
REMARK 200  X-RAY GENERATOR MODEL          : NULL                               
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M; M                               
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.9645; 0.9791, 0.9794, 0.9600     
REMARK 200  MONOCHROMATOR                  : NULL                               
REMARK 200  OPTICS                         : NULL                               
REMARK 200                                                                      
REMARK 200  DETECTOR TYPE                  : CCD; CCD                           
REMARK 200  DETECTOR MANUFACTURER          : ADSC QUANTUM 4; ADSC QUANTUM 4     
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : HKL-2000                           
REMARK 200  DATA SCALING SOFTWARE          : SCALEPACK                          
REMARK 200                                                                      
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 54599                              
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.600                              
REMARK 200  RESOLUTION RANGE LOW       (A) : 50.000                             
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 0.000                              
REMARK 200                                                                      
REMARK 200 OVERALL.                                                             
REMARK 200  COMPLETENESS FOR RANGE     (%) : 99.1                               
REMARK 200  DATA REDUNDANCY                : NULL                               
REMARK 200  R MERGE                    (I) : NULL                               
REMARK 200  R SYM                      (I) : NULL                               
REMARK 200   FOR THE DATA SET  : NULL                               
REMARK 200                                                                      
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.                                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.60                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.70                     
REMARK 200  COMPLETENESS FOR SHELL     (%) : 100.0                              
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL                               
REMARK 200  R MERGE FOR SHELL          (I) : NULL                               
REMARK 200  R SYM FOR SHELL            (I) : NULL                               
REMARK 200   FOR SHELL         : NULL                               
REMARK 200                                                                      
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH; MAD                         
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MAD                          
REMARK 200 SOFTWARE USED: SOLVE                                                 
REMARK 200 STARTING MODEL: NULL                                                 
REMARK 200                                                                      
REMARK 200 REMARK: NULL                                                         
REMARK 280                                                                      
REMARK 280 CRYSTAL                                                              
REMARK 280 SOLVENT CONTENT, VS   (%): 50.00                                     
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.40                     
REMARK 280                                                                      
REMARK 280 CRYSTALLIZATION CONDITIONS: PEG3350, HEPES, AMMONIUM SULFATE,        
REMARK 280  PH 7.5, VAPOR DIFFUSION, HANGING DROP, TEMPERATURE 294K             
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY                                            
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 41 21 2                        
REMARK 290                                                                      
REMARK 290      SYMOP   SYMMETRY                                                
REMARK 290     NNNMMM   OPERATOR                                                
REMARK 290       1555   X,Y,Z                                                   
REMARK 290       2555   -X,-Y,1/2+Z                                             
REMARK 290       3555   1/2-Y,1/2+X,1/4+Z                                       
REMARK 290       4555   1/2+Y,1/2-X,3/4+Z                                       
REMARK 290       5555   1/2-X,1/2+Y,1/4-Z                                       
REMARK 290       6555   1/2+X,1/2-Y,3/4-Z                                       
REMARK 290       7555   Y,X,-Z                                                  
REMARK 290       8555   -Y,-X,1/2-Z                                             
REMARK 290                                                                      
REMARK 290     WHERE NNN -> OPERATOR NUMBER                                     
REMARK 290           MMM -> TRANSLATION VECTOR                                  
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS                            
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM             
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY                
REMARK 290 RELATED MOLECULES.                                                   
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000      104.96400            
REMARK 290   SMTRY1   3  0.000000 -1.000000  0.000000       64.44900            
REMARK 290   SMTRY2   3  1.000000  0.000000  0.000000       64.44900            
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000       52.48200            
REMARK 290   SMTRY1   4  0.000000  1.000000  0.000000       64.44900            
REMARK 290   SMTRY2   4 -1.000000  0.000000  0.000000       64.44900            
REMARK 290   SMTRY3   4  0.000000  0.000000  1.000000      157.44600            
REMARK 290   SMTRY1   5 -1.000000  0.000000  0.000000       64.44900            
REMARK 290   SMTRY2   5  0.000000  1.000000  0.000000       64.44900            
REMARK 290   SMTRY3   5  0.000000  0.000000 -1.000000       52.48200            
REMARK 290   SMTRY1   6  1.000000  0.000000  0.000000       64.44900            
REMARK 290   SMTRY2   6  0.000000 -1.000000  0.000000       64.44900            
REMARK 290   SMTRY3   6  0.000000  0.000000 -1.000000      157.44600            
REMARK 290   SMTRY1   7  0.000000  1.000000  0.000000        0.00000            
REMARK 290   SMTRY2   7  1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY3   7  0.000000  0.000000 -1.000000        0.00000            
REMARK 290   SMTRY1   8  0.000000 -1.000000  0.000000        0.00000            
REMARK 290   SMTRY2   8 -1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY3   8  0.000000  0.000000 -1.000000      104.96400            
REMARK 290                                                                      
REMARK 290 REMARK: NULL                                                         
REMARK 300                                                                      
REMARK 300 BIOMOLECULE: 1                                                       
REMARK 300 THIS ENTRY CONTAINS THE CRYSTALLOGRAPHIC ASYMMETRIC UNIT             
REMARK 300 WHICH CONSISTS OF 3 CHAIN(S). SEE REMARK 350 FOR                     
REMARK 300 INFORMATION ON GENERATING THE BIOLOGICAL MOLECULE(S).                
REMARK 350                                                                      
REMARK 350 GENERATING THE BIOMOLECULE                                           
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN           
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE                
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS          
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND                          
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.                               
REMARK 350                                                                      
REMARK 350 BIOMOLECULE: 1                                                       
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B, C                               
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 465                                                                      
REMARK 465 MISSING RESIDUES                                                     
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE                       
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)                
REMARK 465                                                                      
REMARK 465   M RES C SSSEQI                                                     
REMARK 465     LEU A    48                                                      
REMARK 465     PRO A    49                                                      
REMARK 465     SER A    50                                                      
REMARK 465     PRO A    51                                                      
REMARK 465     SER A    52                                                      
REMARK 465     ARG A    53                                                      
REMARK 465     GLN A    54                                                      
REMARK 465     ALA A    55                                                      
REMARK 465     SER A    56                                                      
REMARK 465     THR A    57                                                      
REMARK 465     ARG A    58                                                      
REMARK 465     ARG A    59                                                      
REMARK 465     MET A    60                                                      
REMARK 465     SER A    61                                                      
REMARK 465     ILE A    62                                                      
REMARK 465     GLY A    63                                                      
REMARK 465     GLU A    64                                                      
REMARK 465     SER A    65                                                      
REMARK 465     ILE A    66                                                      
REMARK 465     THR A   161                                                      
REMARK 465     ASP A   162                                                      
REMARK 465     LYS A   242                                                      
REMARK 465     VAL A   243                                                      
REMARK 465     GLY A   244                                                      
REMARK 465     ARG A   245                                                      
REMARK 465     ARG A   246                                                      
REMARK 465     LYS A   247                                                      
REMARK 465     GLN A   248                                                      
REMARK 465     ASN A   249                                                      
REMARK 465     ARG A   250                                                      
REMARK 465     ARG A   251                                                      
REMARK 465     VAL B   311                                                      
REMARK 465     GLY B   312                                                      
REMARK 465     ASP B   417                                                      
REMARK 465     ILE B   418                                                      
REMARK 465     CYS B   419                                                      
REMARK 465     SER B   420                                                      
REMARK 465     ASN B   421                                                      
REMARK 465     GLU B   422                                                      
REMARK 465     GLU B   423                                                      
REMARK 465     GLY B   488                                                      
REMARK 465     ASN B   489                                                      
REMARK 465     ASN B   490                                                      
REMARK 465     ASN B   491                                                      
REMARK 465     VAL B   492                                                      
REMARK 465     SER B   493                                                      
REMARK 465     VAL B   494                                                      
REMARK 465     PRO B   495                                                      
REMARK 465     GLU B   496                                                      
REMARK 465     ARG B   497                                                      
REMARK 465     HIS B   498                                                      
REMARK 465     ILE B   499                                                      
REMARK 465     PRO B   500                                                      
REMARK 465     SER B   501                                                      
REMARK 465     HIS B   502                                                      
REMARK 465     PHE B   503                                                      
REMARK 465     GLN B   504                                                      
REMARK 465     LYS B   505                                                      
REMARK 465     PHE B   506                                                      
REMARK 465     ARG B   507                                                      
REMARK 465     ARG B   508                                                      
REMARK 465     SER B   509                                                      
REMARK 465     SER B   510                                                      
REMARK 465     ALA B   511                                                      
REMARK 465     SER B   512                                                      
REMARK 465     GLU B   513                                                      
REMARK 465     MET B   514                                                      
REMARK 465     TYR B   515                                                      
REMARK 465     PRO B   516                                                      
REMARK 465     LYS B   517                                                      
REMARK 465     THR B   518                                                      
REMARK 465     THR B   519                                                      
REMARK 465     GLU B   520                                                      
REMARK 465     LYS B   544                                                      
REMARK 465     ASN B   545                                                      
REMARK 465     PHE B   546                                                      
REMARK 465     ALA B   547                                                      
REMARK 465     CYS B   548                                                      
REMARK 465     CYS B   549                                                      
REMARK 465     MET C     1                                                      
REMARK 465     LEU C     2                                                      
REMARK 465     CYS C     3                                                      
REMARK 465     GLU C     4                                                      
REMARK 465     ILE C     5                                                      
REMARK 465     GLU C     6                                                      
REMARK 465     CYS C     7                                                      
REMARK 465     ARG C     8                                                      
REMARK 465     ALA C     9                                                      
REMARK 465     LEU C    10                                                      
REMARK 465     SER C    11                                                      
REMARK 465     THR C    12                                                      
REMARK 465     ALA C    13                                                      
REMARK 465     HIS C    14                                                      
REMARK 465     THR C    15                                                      
REMARK 465     ARG C    16                                                      
REMARK 465     LEU C    17                                                      
REMARK 465     ILE C    18                                                      
REMARK 465     HIS C    19                                                      
REMARK 465     ASP C    20                                                      
REMARK 465     PHE C    21                                                      
REMARK 465     GLU C    22                                                      
REMARK 465     PRO C    23                                                      
REMARK 465     ARG C    24                                                      
REMARK 465     ASP C    25                                                      
REMARK 465     ALA C    26                                                      
REMARK 465     LEU C    27                                                      
REMARK 465     THR C    28                                                      
REMARK 465     TYR C    29                                                      
REMARK 465     LEU C    30                                                      
REMARK 465     GLU C    31                                                      
REMARK 465     GLY C    32                                                      
REMARK 465     LYS C    33                                                      
REMARK 465     ASN C    34                                                      
REMARK 465     ILE C    35                                                      
REMARK 465     PHE C    36                                                      
REMARK 465     THR C    37                                                      
REMARK 465     GLU C    38                                                      
REMARK 465     ASP C    39                                                      
REMARK 465     HIS C    40                                                      
REMARK 465     SER C    41                                                      
REMARK 465     GLU C    42                                                      
REMARK 465     LEU C    43                                                      
REMARK 465     ILE C    44                                                      
REMARK 465     SER C    45                                                      
REMARK 465     LYS C    46                                                      
REMARK 465     MET C    47                                                      
REMARK 465     SER C    48                                                      
REMARK 465     THR C    49                                                      
REMARK 465     ARG C    50                                                      
REMARK 465     LEU C    51                                                      
REMARK 465     GLU C    52                                                      
REMARK 465     ARG C    53                                                      
REMARK 465     ILE C    54                                                      
REMARK 465     ALA C    55                                                      
REMARK 465     ASN C    56                                                      
REMARK 465     PHE C    57                                                      
REMARK 465     LEU C    58                                                      
REMARK 465     ARG C    59                                                      
REMARK 465     ILE C    60                                                      
REMARK 465     TYR C    61                                                      
REMARK 465     ARG C    62                                                      
REMARK 465     ARG C    63                                                      
REMARK 465     GLN C    64                                                      
REMARK 465     ALA C    65                                                      
REMARK 465     SER C    66                                                      
REMARK 465     GLU C    67                                                      
REMARK 465     LEU C    68                                                      
REMARK 465     GLY C    69                                                      
REMARK 465     PRO C    70                                                      
REMARK 465     LEU C    71                                                      
REMARK 465     ILE C    72                                                      
REMARK 465     ASP C    73                                                      
REMARK 465     PHE C    74                                                      
REMARK 465     PHE C    75                                                      
REMARK 465     ASN C    76                                                      
REMARK 465     TYR C    77                                                      
REMARK 465     ASN C    78                                                      
REMARK 465     ASN C    79                                                      
REMARK 465     GLN C    80                                                      
REMARK 465     SER C    81                                                      
REMARK 465     HIS C    82                                                      
REMARK 465     LEU C    83                                                      
REMARK 465     ALA C    84                                                      
REMARK 465     ASP C    85                                                      
REMARK 465     PHE C    86                                                      
REMARK 465     LEU C    87                                                      
REMARK 465     GLU C    88                                                      
REMARK 465     ASP C    89                                                      
REMARK 465     TYR C    90                                                      
REMARK 465     ILE C    91                                                      
REMARK 465     ASP C    92                                                      
REMARK 465     PHE C    93                                                      
REMARK 465     ALA C    94                                                      
REMARK 465     ILE C    95                                                      
REMARK 465     ASN C    96                                                      
REMARK 465     GLU C    97                                                      
REMARK 465     PRO C    98                                                      
REMARK 465     ASP C    99                                                      
REMARK 465     LEU C   100                                                      
REMARK 465     LEU C   101                                                      
REMARK 465     ARG C   102                                                      
REMARK 465     PRO C   103                                                      
REMARK 465     VAL C   104                                                      
REMARK 465     VAL C   105                                                      
REMARK 465     ILE C   106                                                      
REMARK 465     ALA C   107                                                      
REMARK 465     PRO C   108                                                      
REMARK 465     PRO C   415                                                      
REMARK 465     VAL C   416                                                      
REMARK 465     ASP C   417                                                      
REMARK 465     ILE C   418                                                      
REMARK 465     CYS C   419                                                      
REMARK 465     SER C   420                                                      
REMARK 465     ASN C   421                                                      
REMARK 465     GLU C   422                                                      
REMARK 465     GLU C   423                                                      
REMARK 465     GLU C   424                                                      
REMARK 465     GLN C   425                                                      
REMARK 465     LEU C   426                                                      
REMARK 465     ARG C   483                                                      
REMARK 465     LEU C   484                                                      
REMARK 465     LEU C   485                                                      
REMARK 465     GLU C   486                                                      
REMARK 465     ILE C   487                                                      
REMARK 465     GLY C   488                                                      
REMARK 465     ASN C   489                                                      
REMARK 465     ASN C   490                                                      
REMARK 465     ASN C   491                                                      
REMARK 465     VAL C   492                                                      
REMARK 465     SER C   493                                                      
REMARK 465     VAL C   494                                                      
REMARK 465     PRO C   495                                                      
REMARK 465     GLU C   496                                                      
REMARK 465     ARG C   497                                                      
REMARK 465     HIS C   498                                                      
REMARK 465     ILE C   499                                                      
REMARK 465     PRO C   500                                                      
REMARK 465     SER C   501                                                      
REMARK 465     HIS C   502                                                      
REMARK 465     PHE C   503                                                      
REMARK 465     GLN C   504                                                      
REMARK 465     LYS C   505                                                      
REMARK 465     PHE C   506                                                      
REMARK 465     ARG C   507                                                      
REMARK 465     ARG C   508                                                      
REMARK 465     SER C   509                                                      
REMARK 465     SER C   510                                                      
REMARK 465     ALA C   511                                                      
REMARK 465     SER C   512                                                      
REMARK 465     GLU C   513                                                      
REMARK 465     MET C   514                                                      
REMARK 465     TYR C   515                                                      
REMARK 465     PRO C   516                                                      
REMARK 465     LYS C   517                                                      
REMARK 465     THR C   518                                                      
REMARK 465     THR C   519                                                      
REMARK 465     GLU C   520                                                      
REMARK 465     GLU C   521                                                      
REMARK 465     THR C   522                                                      
REMARK 465     VAL C   523                                                      
REMARK 465     ILE C   524                                                      
REMARK 465     ARG C   525                                                      
REMARK 465     PRO C   526                                                      
REMARK 465     GLU C   527                                                      
REMARK 465     ASP C   528                                                      
REMARK 465     PHE C   529                                                      
REMARK 465     PRO C   530                                                      
REMARK 465     LYS C   531                                                      
REMARK 465     PHE C   532                                                      
REMARK 465     LYS C   544                                                      
REMARK 465     ASN C   545                                                      
REMARK 465     PHE C   546                                                      
REMARK 465     ALA C   547                                                      
REMARK 465     CYS C   548                                                      
REMARK 465     CYS C   549                                                      
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT                     
REMARK 500                                                                      
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.                            
REMARK 500                                                                      
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI                             
REMARK 500   O    ASP B   469     N    GLN B   471              2.15            
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND LENGTHS                                      
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,2(A3,1X,A1,I4,A1,1X,A4,3X),F6.3)                  
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   RES CSSEQI ATM2   DEVIATION                     
REMARK 500    ASP A  67   C     ASP A  67   O     -0.054                        
REMARK 500    LYS A  69   CB    LYS A  69   CG     0.060                        
REMARK 500    MET A 116   SD    MET A 116   CE    -0.077                        
REMARK 500    GLN A 163   CD    GLN A 163   OE1    0.067                        
REMARK 500    GLN A 163   CD    GLN A 163   NE2    0.065                        
REMARK 500    MET A 166   SD    MET A 166   CE    -0.101                        
REMARK 500    MET A 185   SD    MET A 185   CE    -0.078                        
REMARK 500    MET A 186   SD    MET A 186   CE    -0.046                        
REMARK 500    MET A 225   SD    MET A 225   CE    -0.097                        
REMARK 500    MET B   1   SD    MET B   1   CE     0.069                        
REMARK 500    MET B  47   CG    MET B  47   SD     0.048                        
REMARK 500    MET B 234   SD    MET B 234   CE    -0.051                        
REMARK 500    GLU C 134   C     GLU C 134   O      0.053                        
REMARK 500    MET C 210   SD    MET C 210   CE    -0.044                        
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES                                       
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)              
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3                                     
REMARK 500    GLY A  68   N   -  CA  -  C   ANGL. DEV. = 29.1 DEGREES           
REMARK 500    LYS A  69   N   -  CA  -  C   ANGL. DEV. = 17.5 DEGREES           
REMARK 500    GLY A  68   CA  -  C   -  N   ANGL. DEV. =-11.0 DEGREES           
REMARK 500    LYS A  69   C   -  N   -  CA  ANGL. DEV. =  9.7 DEGREES           
REMARK 500    ILE A  70   N   -  CA  -  C   ANGL. DEV. = 13.0 DEGREES           
REMARK 500    VAL A 153   N   -  CA  -  C   ANGL. DEV. = 10.7 DEGREES           
REMARK 500    GLU A 190   N   -  CA  -  C   ANGL. DEV. =-11.1 DEGREES           
REMARK 500    THR B  37   N   -  CA  -  C   ANGL. DEV. = -8.7 DEGREES           
REMARK 500    VAL B 105   N   -  CA  -  C   ANGL. DEV. =  9.2 DEGREES           
REMARK 500    LEU B 155   N   -  CA  -  C   ANGL. DEV. = -8.8 DEGREES           
REMARK 500    LEU B 157   N   -  CA  -  C   ANGL. DEV. = -8.9 DEGREES           
REMARK 500    SER B 222   N   -  CA  -  C   ANGL. DEV. =-11.7 DEGREES           
REMARK 500    ARG B 267   N   -  CA  -  C   ANGL. DEV. =-11.1 DEGREES           
REMARK 500    CYS B 268   N   -  CA  -  C   ANGL. DEV. = 10.4 DEGREES           
REMARK 500    LEU B 269   N   -  CA  -  C   ANGL. DEV. =-11.9 DEGREES           
REMARK 500    THR B 271   N   -  CA  -  C   ANGL. DEV. = -8.7 DEGREES           
REMARK 500    GLY B 359   N   -  CA  -  C   ANGL. DEV. = -8.7 DEGREES           
REMARK 500    ASP B 428   N   -  CA  -  C   ANGL. DEV. = -9.1 DEGREES           
REMARK 500    ILE B 479   N   -  CA  -  C   ANGL. DEV. = -8.8 DEGREES           
REMARK 500    LEU B 480   N   -  CA  -  C   ANGL. DEV. =-10.3 DEGREES           
REMARK 500    LEU C 120   CA  -  CB  -  CG  ANGL. DEV. =  9.0 DEGREES           
REMARK 500    GLU C 134   CB  -  CA  -  C   ANGL. DEV. = 10.9 DEGREES           
REMARK 500    ARG C 267   N   -  CA  -  C   ANGL. DEV. = -8.9 DEGREES           
REMARK 500    LEU C 269   N   -  CA  -  C   ANGL. DEV. =-13.6 DEGREES           
REMARK 500    PRO C 308   N   -  CA  -  C   ANGL. DEV. =  9.2 DEGREES           
REMARK 500    GLU C 341   N   -  CA  -  C   ANGL. DEV. =-10.6 DEGREES           
REMARK 500    PRO C 342   N   -  CA  -  C   ANGL. DEV. = 10.0 DEGREES           
REMARK 500    PRO C 342   C   -  N   -  CA  ANGL. DEV. = 16.4 DEGREES           
REMARK 500    PRO C 342   C   -  N   -  CD  ANGL. DEV. =-13.6 DEGREES           
REMARK 500    LEU C 535   N   -  CA  -  C   ANGL. DEV. =-12.8 DEGREES           
DBREF  2A5Y A   48   251  UNP    P41958   CED9_CAEEL      48    251             
DBREF  2A5Y B    1   549  UNP    P30429   CED4_CAEEL       1    549             
DBREF  2A5Y C    1   549  UNP    P30429   CED4_CAEEL       1    549             
SEQADV 2A5Y SER A  107  UNP  P41958    CYS   107 ENGINEERED                     
SEQADV 2A5Y SER A  135  UNP  P41958    CYS   135 ENGINEERED                     
SEQADV 2A5Y SER A  164  UNP  P41958    CYS   164 ENGINEERED                     
SEQRES   1 A  204  LEU PRO SER PRO SER ARG GLN ALA SER THR ARG ARG MET          
SEQRES   2 A  204  SER ILE GLY GLU SER ILE ASP GLY LYS ILE ASN ASP TRP          
SEQRES   3 A  204  GLU GLU PRO ARG LEU ASP ILE GLU GLY PHE VAL VAL ASP          
SEQRES   4 A  204  TYR PHE THR HIS ARG ILE ARG GLN ASN GLY MET GLU TRP          
SEQRES   5 A  204  PHE GLY ALA PRO GLY LEU PRO SER GLY VAL GLN PRO GLU          
SEQRES   6 A  204  HIS GLU MET MET ARG VAL MET GLY THR ILE PHE GLU LYS          
SEQRES   7 A  204  LYS HIS ALA GLU ASN PHE GLU THR PHE SER GLU GLN LEU          
SEQRES   8 A  204  LEU ALA VAL PRO ARG ILE SER PHE SER LEU TYR GLN ASP          
SEQRES   9 A  204  VAL VAL ARG THR VAL GLY ASN ALA GLN THR ASP GLN SER          
SEQRES  10 A  204  PRO MET SER TYR GLY ARG LEU ILE GLY LEU ILE SER PHE          
SEQRES  11 A  204  GLY GLY PHE VAL ALA ALA LYS MET MET GLU SER VAL GLU          
SEQRES  12 A  204  LEU GLN GLY GLN VAL ARG ASN LEU PHE VAL TYR THR SER          
SEQRES  13 A  204  LEU PHE ILE LYS THR ARG ILE ARG ASN ASN TRP LYS GLU          
SEQRES  14 A  204  HIS ASN ARG SER TRP ASP ASP PHE MET THR LEU GLY LYS          
SEQRES  15 A  204  GLN MET LYS GLU ASP TYR GLU ARG ALA GLU ALA GLU LYS          
SEQRES  16 A  204  VAL GLY ARG ARG LYS GLN ASN ARG ARG                          
SEQRES   1 B  549  MET LEU CYS GLU ILE GLU CYS ARG ALA LEU SER THR ALA          
SEQRES   2 B  549  HIS THR ARG LEU ILE HIS ASP PHE GLU PRO ARG ASP ALA          
SEQRES   3 B  549  LEU THR TYR LEU GLU GLY LYS ASN ILE PHE THR GLU ASP          
SEQRES   4 B  549  HIS SER GLU LEU ILE SER LYS MET SER THR ARG LEU GLU          
SEQRES   5 B  549  ARG ILE ALA ASN PHE LEU ARG ILE TYR ARG ARG GLN ALA          
SEQRES   6 B  549  SER GLU LEU GLY PRO LEU ILE ASP PHE PHE ASN TYR ASN          
SEQRES   7 B  549  ASN GLN SER HIS LEU ALA ASP PHE LEU GLU ASP TYR ILE          
SEQRES   8 B  549  ASP PHE ALA ILE ASN GLU PRO ASP LEU LEU ARG PRO VAL          
SEQRES   9 B  549  VAL ILE ALA PRO GLN PHE SER ARG GLN MET LEU ASP ARG          
SEQRES  10 B  549  LYS LEU LEU LEU GLY ASN VAL PRO LYS GLN MET THR CYS          
SEQRES  11 B  549  TYR ILE ARG GLU TYR HIS VAL ASP ARG VAL ILE LYS LYS          
SEQRES  12 B  549  LEU ASP GLU MET CYS ASP LEU ASP SER PHE PHE LEU PHE          
SEQRES  13 B  549  LEU HIS GLY ARG ALA GLY SER GLY LYS SER VAL ILE ALA          
SEQRES  14 B  549  SER GLN ALA LEU SER LYS SER ASP GLN LEU ILE GLY ILE          
SEQRES  15 B  549  ASN TYR ASP SER ILE VAL TRP LEU LYS ASP SER GLY THR          
SEQRES  16 B  549  ALA PRO LYS SER THR PHE ASP LEU PHE THR ASP ILE LEU          
SEQRES  17 B  549  LEU MET LEU LYS SER GLU ASP ASP LEU LEU ASN PHE PRO          
SEQRES  18 B  549  SER VAL GLU HIS VAL THR SER VAL VAL LEU LYS ARG MET          
SEQRES  19 B  549  ILE CYS ASN ALA LEU ILE ASP ARG PRO ASN THR LEU PHE          
SEQRES  20 B  549  VAL PHE ASP ASP VAL VAL GLN GLU GLU THR ILE ARG TRP          
SEQRES  21 B  549  ALA GLN GLU LEU ARG LEU ARG CYS LEU VAL THR THR ARG          
SEQRES  22 B  549  ASP VAL GLU ILE SER ASN ALA ALA SER GLN THR CYS GLU          
SEQRES  23 B  549  PHE ILE GLU VAL THR SER LEU GLU ILE ASP GLU CYS TYR          
SEQRES  24 B  549  ASP PHE LEU GLU ALA TYR GLY MET PRO MET PRO VAL GLY          
SEQRES  25 B  549  GLU LYS GLU GLU ASP VAL LEU ASN LYS THR ILE GLU LEU          
SEQRES  26 B  549  SER SER GLY ASN PRO ALA THR LEU MET MET PHE PHE LYS          
SEQRES  27 B  549  SER CYS GLU PRO LYS THR PHE GLU LYS MET ALA GLN LEU          
SEQRES  28 B  549  ASN ASN LYS LEU GLU SER ARG GLY LEU VAL GLY VAL GLU          
SEQRES  29 B  549  CYS ILE THR PRO TYR SER TYR LYS SER LEU ALA MET ALA          
SEQRES  30 B  549  LEU GLN ARG CYS VAL GLU VAL LEU SER ASP GLU ASP ARG          
SEQRES  31 B  549  SER ALA LEU ALA PHE ALA VAL VAL MET PRO PRO GLY VAL          
SEQRES  32 B  549  ASP ILE PRO VAL LYS LEU TRP SER CYS VAL ILE PRO VAL          
SEQRES  33 B  549  ASP ILE CYS SER ASN GLU GLU GLU GLN LEU ASP ASP GLU          
SEQRES  34 B  549  VAL ALA ASP ARG LEU LYS ARG LEU SER LYS ARG GLY ALA          
SEQRES  35 B  549  LEU LEU SER GLY LYS ARG MET PRO VAL LEU THR PHE LYS          
SEQRES  36 B  549  ILE ASP HIS ILE ILE HIS MET PHE LEU LYS HIS VAL VAL          
SEQRES  37 B  549  ASP ALA GLN THR ILE ALA ASN GLY ILE SER ILE LEU GLU          
SEQRES  38 B  549  GLN ARG LEU LEU GLU ILE GLY ASN ASN ASN VAL SER VAL          
SEQRES  39 B  549  PRO GLU ARG HIS ILE PRO SER HIS PHE GLN LYS PHE ARG          
SEQRES  40 B  549  ARG SER SER ALA SER GLU MET TYR PRO LYS THR THR GLU          
SEQRES  41 B  549  GLU THR VAL ILE ARG PRO GLU ASP PHE PRO LYS PHE MET          
SEQRES  42 B  549  GLN LEU HIS GLN LYS PHE TYR ASP SER LEU LYS ASN PHE          
SEQRES  43 B  549  ALA CYS CYS                                                  
SEQRES   1 C  549  MET LEU CYS GLU ILE GLU CYS ARG ALA LEU SER THR ALA          
SEQRES   2 C  549  HIS THR ARG LEU ILE HIS ASP PHE GLU PRO ARG ASP ALA          
SEQRES   3 C  549  LEU THR TYR LEU GLU GLY LYS ASN ILE PHE THR GLU ASP          
SEQRES   4 C  549  HIS SER GLU LEU ILE SER LYS MET SER THR ARG LEU GLU          
SEQRES   5 C  549  ARG ILE ALA ASN PHE LEU ARG ILE TYR ARG ARG GLN ALA          
SEQRES   6 C  549  SER GLU LEU GLY PRO LEU ILE ASP PHE PHE ASN TYR ASN          
SEQRES   7 C  549  ASN GLN SER HIS LEU ALA ASP PHE LEU GLU ASP TYR ILE          
SEQRES   8 C  549  ASP PHE ALA ILE ASN GLU PRO ASP LEU LEU ARG PRO VAL          
SEQRES   9 C  549  VAL ILE ALA PRO GLN PHE SER ARG GLN MET LEU ASP ARG          
SEQRES  10 C  549  LYS LEU LEU LEU GLY ASN VAL PRO LYS GLN MET THR CYS          
SEQRES  11 C  549  TYR ILE ARG GLU TYR HIS VAL ASP ARG VAL ILE LYS LYS          
SEQRES  12 C  549  LEU ASP GLU MET CYS ASP LEU ASP SER PHE PHE LEU PHE          
SEQRES  13 C  549  LEU HIS GLY ARG ALA GLY SER GLY LYS SER VAL ILE ALA          
SEQRES  14 C  549  SER GLN ALA LEU SER LYS SER ASP GLN LEU ILE GLY ILE          
SEQRES  15 C  549  ASN TYR ASP SER ILE VAL TRP LEU LYS ASP SER GLY THR          
SEQRES  16 C  549  ALA PRO LYS SER THR PHE ASP LEU PHE THR ASP ILE LEU          
SEQRES  17 C  549  LEU MET LEU LYS SER GLU ASP ASP LEU LEU ASN PHE PRO          
SEQRES  18 C  549  SER VAL GLU HIS VAL THR SER VAL VAL LEU LYS ARG MET          
SEQRES  19 C  549  ILE CYS ASN ALA LEU ILE ASP ARG PRO ASN THR LEU PHE          
SEQRES  20 C  549  VAL PHE ASP ASP VAL VAL GLN GLU GLU THR ILE ARG TRP          
SEQRES  21 C  549  ALA GLN GLU LEU ARG LEU ARG CYS LEU VAL THR THR ARG          
SEQRES  22 C  549  ASP VAL GLU ILE SER ASN ALA ALA SER GLN THR CYS GLU          
SEQRES  23 C  549  PHE ILE GLU VAL THR SER LEU GLU ILE ASP GLU CYS TYR          
SEQRES  24 C  549  ASP PHE LEU GLU ALA TYR GLY MET PRO MET PRO VAL GLY          
SEQRES  25 C  549  GLU LYS GLU GLU ASP VAL LEU ASN LYS THR ILE GLU LEU          
SEQRES  26 C  549  SER SER GLY ASN PRO ALA THR LEU MET MET PHE PHE LYS          
SEQRES  27 C  549  SER CYS GLU PRO LYS THR PHE GLU LYS MET ALA GLN LEU          
SEQRES  28 C  549  ASN ASN LYS LEU GLU SER ARG GLY LEU VAL GLY VAL GLU          
SEQRES  29 C  549  CYS ILE THR PRO TYR SER TYR LYS SER LEU ALA MET ALA          
SEQRES  30 C  549  LEU GLN ARG CYS VAL GLU VAL LEU SER ASP GLU ASP ARG          
SEQRES  31 C  549  SER ALA LEU ALA PHE ALA VAL VAL MET PRO PRO GLY VAL          
SEQRES  32 C  549  ASP ILE PRO VAL LYS LEU TRP SER CYS VAL ILE PRO VAL          
SEQRES  33 C  549  ASP ILE CYS SER ASN GLU GLU GLU GLN LEU ASP ASP GLU          
SEQRES  34 C  549  VAL ALA ASP ARG LEU LYS ARG LEU SER LYS ARG GLY ALA          
SEQRES  35 C  549  LEU LEU SER GLY LYS ARG MET PRO VAL LEU THR PHE LYS          
SEQRES  36 C  549  ILE ASP HIS ILE ILE HIS MET PHE LEU LYS HIS VAL VAL          
SEQRES  37 C  549  ASP ALA GLN THR ILE ALA ASN GLY ILE SER ILE LEU GLU          
SEQRES  38 C  549  GLN ARG LEU LEU GLU ILE GLY ASN ASN ASN VAL SER VAL          
SEQRES  39 C  549  PRO GLU ARG HIS ILE PRO SER HIS PHE GLN LYS PHE ARG          
SEQRES  40 C  549  ARG SER SER ALA SER GLU MET TYR PRO LYS THR THR GLU          
SEQRES  41 C  549  GLU THR VAL ILE ARG PRO GLU ASP PHE PRO LYS PHE MET          
SEQRES  42 C  549  GLN LEU HIS GLN LYS PHE TYR ASP SER LEU LYS ASN PHE          
SEQRES  43 C  549  ALA CYS CYS                                                  
HET     MG    501       1                                                       
HET     MG    502       1                                                       
HET    ATP    301      31                                                       
HET    ATP    401      31                                                       
HETNAM      MG MAGNESIUM ION                                                    
HETNAM     ATP ADENOSINE-5'-TRIPHOSPHATE                                        
FORMUL   4   MG    2(MG 2+)                                                     
FORMUL   6  ATP    2(C10 H16 N5 O13 P3)                                         
FORMUL   8  HOH   *260(H2 O)                                                    
HELIX    1   1 ASN A   71  LEU A   78  5                                   8    
HELIX    2   2 ASP A   79  GLN A   94  1                                  16    
HELIX    3   3 GLN A  110  HIS A  127  1                                  18    
HELIX    4   4 ASN A  130  ALA A  140  1                                  11    
HELIX    5   5 SER A  145  ARG A  154  1                                  10    
HELIX    6   6 SER A  167  GLU A  187  1                                  21    
HELIX    7   7 GLN A  194  ARG A  211  1                                  18    
HELIX    8   8 SER A  220  GLU A  239  1                                  20    
HELIX    9   9 CYS B    3  PHE B   21  1                                  19    
HELIX   10  10 GLU B   22  ASP B   25  5                                   4    
HELIX   11  11 ALA B   26  LYS B   33  1                                   8    
HELIX   12  12 THR B   37  LYS B   46  1                                  10    
HELIX   13  13 THR B   49  ALA B   65  1                                  17    
HELIX   14  14 LEU B   68  ASN B   78  1                                  11    
HELIX   15  15 GLN B   80  GLU B   97  1                                  18    
HELIX   16  16 LEU B  100  VAL B  105  1                                   6    
HELIX   17  17 SER B  111  GLY B  122  1                                  12    
HELIX   18  18 ARG B  133  CYS B  148  1                                  16    
HELIX   19  19 GLY B  164  SER B  176  1                                  13    
HELIX   20  20 LYS B  198  LYS B  212  1                                  15    
HELIX   21  21 THR B  227  ILE B  240  1                                  14    
HELIX   22  22 GLN B  254  LEU B  264  1                                  11    
HELIX   23  23 ASP B  274  ALA B  281  5                                   8    
HELIX   24  24 GLU B  294  TYR B  305  1                                  12    
HELIX   25  25 GLU B  313  SER B  327  1                                  15    
HELIX   26  26 ASN B  329  LYS B  338  1                                  10    
HELIX   27  27 THR B  344  GLY B  359  1                                  16    
HELIX   28  28 SER B  373  VAL B  384  1                                  12    
HELIX   29  29 SER B  386  LEU B  393  1                                   8    
HELIX   30  30 ALA B  394  VAL B  398  5                                   5    
HELIX   31  31 VAL B  407  ILE B  414  1                                   8    
HELIX   32  32 ASP B  427  LEU B  437  1                                  11    
HELIX   33  33 ASP B  457  HIS B  466  1                                  10    
HELIX   34  34 ALA B  470  SER B  478  1                                   9    
HELIX   35  35 HIS B  536  ASP B  541  1                                   6    
HELIX   36  36 GLN C  109  GLY C  122  1                                  14    
HELIX   37  37 ARG C  133  CYS C  148  1                                  16    
HELIX   38  38 GLY C  164  LYS C  175  1                                  12    
HELIX   39  39 LYS C  198  LYS C  212  1                                  15    
HELIX   40  40 SER C  213  PHE C  220  1                                   8    
HELIX   41  41 THR C  227  LEU C  239  1                                  13    
HELIX   42  42 GLN C  254  LEU C  264  1                                  11    
HELIX   43  43 ASP C  274  ASN C  279  5                                   6    
HELIX   44  44 GLU C  294  TYR C  305  1                                  12    
HELIX   45  45 GLY C  312  SER C  326  1                                  15    
HELIX   46  46 ASN C  329  CYS C  340  1                                  12    
HELIX   47  47 THR C  344  GLY C  359  1                                  16    
HELIX   48  48 LEU C  360  GLU C  364  5                                   5    
HELIX   49  49 SER C  373  GLU C  383  1                                  11    
HELIX   50  50 GLU C  388  LEU C  393  1                                   6    
HELIX   51  51 VAL C  407  CYS C  412  1                                   6    
HELIX   52  52 LYS C  435  ARG C  440  1                                   6    
HELIX   53  53 ASP C  457  HIS C  466  1                                  10    
HELIX   54  54 GLN C  471  ILE C  479  1                                   9    
HELIX   55  55 MET C  533  LEU C  535  5                                   3    
HELIX   56  56 HIS C  536  LEU C  543  1                                   8    
SHEET    1   A 5 SER B 186  LYS B 191  0                                        
SHEET    2   A 5 THR B 245  VAL B 252  1  O  VAL B 248   N  VAL B 188           
SHEET    3   A 5 ARG B 267  THR B 272  1  O  ARG B 267   N  PHE B 247           
SHEET    4   A 5 PHE B 153  HIS B 158  1  N  LEU B 157   O  VAL B 270           
SHEET    5   A 5 CYS B 285  GLU B 289  1  O  GLU B 286   N  PHE B 156           
SHEET    1   B 3 ILE B 405  PRO B 406  0                                        
SHEET    2   B 3 THR B 453  LYS B 455 -1  O  PHE B 454   N  ILE B 405           
SHEET    3   B 3 SER B 445  LYS B 447 -1  N  SER B 445   O  LYS B 455           
SHEET    1   C 5 SER C 186  LYS C 191  0                                        
SHEET    2   C 5 THR C 245  ASP C 250  1  O  VAL C 248   N  VAL C 188           
SHEET    3   C 5 ARG C 267  THR C 272  1  O  ARG C 267   N  PHE C 247           
SHEET    4   C 5 PHE C 154  GLY C 159  1  N  LEU C 157   O  VAL C 270           
SHEET    5   C 5 GLU C 286  VAL C 290  1  O  VAL C 290   N  HIS C 158           
SHEET    1   D 3 ASP C 404  PRO C 406  0                                        
SHEET    2   D 3 THR C 453  LYS C 455 -1  O  PHE C 454   N  ILE C 405           
SHEET    3   D 3 SER C 445  LYS C 447 -1  N  LYS C 447   O  THR C 453           
CISPEP   1 MET B  449    PRO B  450          0        -0.01                     
CISPEP   2 GLU C  341    PRO C  342          0         2.61                     
CISPEP   3 MET C  449    PRO C  450          0         0.22                     
CRYST1  128.898  128.898  209.928  90.00  90.00  90.00 P 41 21 2    16          
ORIGX1      1.000000  0.000000  0.000000        0.00000                         
ORIGX2      0.000000  1.000000  0.000000        0.00000                         
ORIGX3      0.000000  0.000000  1.000000        0.00000                         
SCALE1      0.007758  0.000000  0.000000        0.00000                         
SCALE2      0.000000  0.007758  0.000000        0.00000                         
SCALE3      0.000000  0.000000  0.004764        0.00000