PDB entry 1y8k

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HEADER    OXYGEN STORAGE/TRANSPORT                13-DEC-04   1Y8K              
TITLE     HORSE METHEMOGLOBIN LOW SALT, PH 7.0 (88% RELATIVE HUMIDITY)          
COMPND    MOL_ID: 1;                                                            
COMPND   2 MOLECULE: HEMOGLOBIN ALPHA CHAINS;                                   
COMPND   3 CHAIN: A, C;                                                         
COMPND   4 SYNONYM: SLOW AND FAST;                                              
COMPND   5 OTHER_DETAILS: PH 7.0 AQUOMET STRUCTURE;                             
COMPND   6 MOL_ID: 2;                                                           
COMPND   7 MOLECULE: HEMOGLOBIN BETA CHAIN;                                     
COMPND   8 CHAIN: B, D                                                          
SOURCE    MOL_ID: 1;                                                            
SOURCE   2 ORGANISM_SCIENTIFIC: EQUUS CABALLUS;                                 
SOURCE   3 ORGANISM_COMMON: HORSE;                                              
SOURCE   4 MOL_ID: 2;                                                           
SOURCE   5 ORGANISM_SCIENTIFIC: EQUUS CABALLUS;                                 
SOURCE   6 ORGANISM_COMMON: HORSE                                               
KEYWDS    AQUO METHEMOGLOBIN, QUARTERNARY ASSOCIATION, ALLOSTERIC               
KEYWDS   2 TRANSITION, PROTEIN HYDRATION                                        
EXPDTA    X-RAY DIFFRACTION                                                     
AUTHOR    R.SANKARANARAYANAN,B.K.BISWAL,M.VIJAYAN                               
REVDAT   1   26-JUL-05 1Y8K    0                                                
JRNL        AUTH   R.SANKARANARAYANAN,B.K.BISWAL,M.VIJAYAN                      
JRNL        TITL   A NEW RELAXED STATE IN HORSE METHEMOGLOBIN                   
JRNL        TITL 2 CHARACTERIZED BY CRYSTALLOGRAPHIC STUDIES.                   
JRNL        REF    PROTEINS                      V.  60   547 2005              
JRNL        REFN   ASTM PSFGEY  US ISSN 0887-3585                               
REMARK   1                                                                      
REMARK   1 REFERENCE 1                                                          
REMARK   1  AUTH   B.K.BISWAL,M.VIJAYAN                                         
REMARK   1  TITL   STRUCTURE OF HUMAN METHAEMOGLOBIN: THE VARIATION             
REMARK   1  TITL 2 OF A THEME                                                   
REMARK   1  REF    CURR.SCI.                     V.  81  1100 2001              
REMARK   1  REFN   ASTM CUSCAM  II ISSN 0011-3891                               
REMARK   1 REFERENCE 2                                                          
REMARK   1  AUTH   B.K.BISWAL,M.VIJAYAN                                         
REMARK   1  TITL   STRUCTURES OF HUMAN OXY- AND DEOXYHAEMOGLOBIN AT             
REMARK   1  TITL 2 DIFFERENT LEVELS OF HUMIDITY: VARIABILITY IN THE T           
REMARK   1  TITL 3 STATE                                                        
REMARK   1  REF    ACTA CRYSTALLOGR.,SECT.D      V.  58  1155 2002              
REMARK   1  REFN   ASTM ABCRE6  DK ISSN 0907-4449                               
REMARK   2                                                                      
REMARK   2 RESOLUTION. 2.30 ANGSTROMS.                                          
REMARK   3                                                                      
REMARK   3 REFINEMENT.                                                          
REMARK   3   PROGRAM     : CNS 0.4                                              
REMARK   3   AUTHORS     : BRUNGER,ADAMS,CLORE,DELANO,GROS,GROSSE-              
REMARK   3               : KUNSTLEVE,JIANG,KUSZEWSKI,NILGES, PANNU,             
REMARK   3               : READ,RICE,SIMONSON,WARREN                            
REMARK   3                                                                      
REMARK   3  REFINEMENT TARGET : NULL                                            
REMARK   3                                                                      
REMARK   3  DATA USED IN REFINEMENT.                                            
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.30                           
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 20.00                          
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.000                          
REMARK   3   DATA CUTOFF HIGH         (ABS(F)) : 372154.060                     
REMARK   3   DATA CUTOFF LOW          (ABS(F)) : 0.0000                         
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : 92.0                           
REMARK   3   NUMBER OF REFLECTIONS             : 23480                          
REMARK   3                                                                      
REMARK   3  FIT TO DATA USED IN REFINEMENT.                                     
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT                      
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM                          
REMARK   3   R VALUE            (WORKING SET) : 0.183                           
REMARK   3   FREE R VALUE                     : 0.228                           
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 4.900                           
REMARK   3   FREE R VALUE TEST SET COUNT      : 1150                            
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : 0.007                           
REMARK   3                                                                      
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.                                  
REMARK   3   TOTAL NUMBER OF BINS USED           : 6                            
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 2.30                         
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 2.44                         
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 86.30                        
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : 3436                         
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2300                       
REMARK   3   BIN FREE R VALUE                    : 0.2850                       
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : 4.70                         
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : 169                          
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : 0.022                        
REMARK   3                                                                      
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.                    
REMARK   3   PROTEIN ATOMS            : 4406                                    
REMARK   3   NUCLEIC ACID ATOMS       : 0                                       
REMARK   3   HETEROGEN ATOMS          : 172                                     
REMARK   3   SOLVENT ATOMS            : 386                                     
REMARK   3                                                                      
REMARK   3  B VALUES.                                                           
REMARK   3   FROM WILSON PLOT           (A**2) : 31.50                          
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 32.10                          
REMARK   3   OVERALL ANISOTROPIC B VALUE.                                       
REMARK   3    B11 (A**2) : -0.90000                                             
REMARK   3    B22 (A**2) : -0.20000                                             
REMARK   3    B33 (A**2) : 1.10000                                              
REMARK   3    B12 (A**2) : 0.00000                                              
REMARK   3    B13 (A**2) : 0.00000                                              
REMARK   3    B23 (A**2) : 0.00000                                              
REMARK   3                                                                      
REMARK   3  ESTIMATED COORDINATE ERROR.                                         
REMARK   3   ESD FROM LUZZATI PLOT        (A) : 0.25                            
REMARK   3   ESD FROM SIGMAA              (A) : 0.19                            
REMARK   3   LOW RESOLUTION CUTOFF        (A) : 20.00                           
REMARK   3                                                                      
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.                         
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : 0.32                            
REMARK   3   ESD FROM C-V SIGMAA          (A) : 0.27                            
REMARK   3                                                                      
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.                                   
REMARK   3   BOND LENGTHS                 (A) : 0.008                           
REMARK   3   BOND ANGLES            (DEGREES) : 1.20                            
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : 19.70                           
REMARK   3   IMPROPER ANGLES        (DEGREES) : 1.02                            
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL MODEL : RESTRAINED                                
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA                
REMARK   3   MAIN-CHAIN BOND              (A**2) : NULL  ; NULL                 
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : NULL  ; NULL                 
REMARK   3   SIDE-CHAIN BOND              (A**2) : NULL  ; NULL                 
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : NULL  ; NULL                 
REMARK   3                                                                      
REMARK   3  BULK SOLVENT MODELING.                                              
REMARK   3   METHOD USED : FLAT MODEL                                           
REMARK   3   KSOL        : 0.31                                                 
REMARK   3   BSOL        : 49.34                                                
REMARK   3                                                                      
REMARK   3  NCS MODEL : NULL                                                    
REMARK   3                                                                      
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT          
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL                 
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL                 
REMARK   3                                                                      
REMARK   3  PARAMETER FILE  1  : PROTEIN_REP.PARAM                              
REMARK   3  PARAMETER FILE  2  : PARAM19X.HEME                                  
REMARK   3  PARAMETER FILE  3  : WATER_REP.PARAM                                
REMARK   3  PARAMETER FILE  4  : NULL                                           
REMARK   3  TOPOLOGY FILE  1   : PROTEIN.TOP                                    
REMARK   3  TOPOLOGY FILE  2   : TOPH19X.HEME                                   
REMARK   3  TOPOLOGY FILE  3   : WATER.TOP                                      
REMARK   3  TOPOLOGY FILE  4   : NULL                                           
REMARK   3                                                                      
REMARK   3  OTHER REFINEMENT REMARKS: NULL                                      
REMARK   4                                                                      
REMARK   4 1Y8K COMPLIES WITH FORMAT V. 3.0, 1-DEC-2006                         
REMARK   4                                                                      
REMARK   4 THIS IS THE REMEDIATED VERSION OF THIS PDB ENTRY.                    
REMARK   4 REMEDIATED DATA FILE REVISION 3.101 (2007-05-29)                     
REMARK 100                                                                      
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB .                              
REMARK 100 THE RCSB ID CODE IS RCSB031251.                                      
REMARK 200                                                                      
REMARK 200 EXPERIMENTAL DETAILS                                                 
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION                  
REMARK 200  DATE OF DATA COLLECTION        : 28-AUG-2002                        
REMARK 200  TEMPERATURE           (KELVIN) : 293.0                              
REMARK 200  PH                             : 7.00                               
REMARK 200  NUMBER OF CRYSTALS USED        : 1                                  
REMARK 200                                                                      
REMARK 200  SYNCHROTRON              (Y/N) : N                                  
REMARK 200  RADIATION SOURCE               : ROTATING ANODE                     
REMARK 200  BEAMLINE                       : NULL                               
REMARK 200  X-RAY GENERATOR MODEL          : RIGAKU RU200                       
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M                                  
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.5418                             
REMARK 200  MONOCHROMATOR                  : GRAPHITE                           
REMARK 200  OPTICS                         : OSMIC MIRROR                       
REMARK 200                                                                      
REMARK 200  DETECTOR TYPE                  : IMAGE PLATE                        
REMARK 200  DETECTOR MANUFACTURER          : MARRESEARCH                        
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : DENZO                              
REMARK 200  DATA SCALING SOFTWARE          : SCALEPACK                          
REMARK 200                                                                      
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 24502                              
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.300                              
REMARK 200  RESOLUTION RANGE LOW       (A) : 20.000                             
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL                               
REMARK 200                                                                      
REMARK 200 OVERALL.                                                             
REMARK 200  COMPLETENESS FOR RANGE     (%) : 96.0                               
REMARK 200  DATA REDUNDANCY                : NULL                               
REMARK 200  R MERGE                    (I) : 0.11200                            
REMARK 200  R SYM                      (I) : NULL                               
REMARK 200   FOR THE DATA SET  : NULL                               
REMARK 200                                                                      
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.                                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.30                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.38                     
REMARK 200  COMPLETENESS FOR SHELL     (%) : 94.9                               
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL                               
REMARK 200  R MERGE FOR SHELL          (I) : 0.52300                            
REMARK 200  R SYM FOR SHELL            (I) : NULL                               
REMARK 200   FOR SHELL         : NULL                               
REMARK 200                                                                      
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH                              
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT        
REMARK 200 SOFTWARE USED: AMORE                                                 
REMARK 200 STARTING MODEL: HORSE METHEMOGLOBIN, PDB ENTRY 2MHB                  
REMARK 200                                                                      
REMARK 200 REMARK: NULL                                                         
REMARK 280                                                                      
REMARK 280 CRYSTAL                                                              
REMARK 280 SOLVENT CONTENT, VS   (%): 43.10                                     
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.20                     
REMARK 280                                                                      
REMARK 280 CRYSTALLIZATION CONDITIONS: 0.01M PHOSPHATE BUFFER, 30% PEG          
REMARK 280  3350, PH 7.0, LIQUID DIFFUSION, TEMPERATURE 293K                    
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY                                            
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 21                       
REMARK 290                                                                      
REMARK 290      SYMOP   SYMMETRY                                                
REMARK 290     NNNMMM   OPERATOR                                                
REMARK 290       1555   X,Y,Z                                                   
REMARK 290       2555   1/2-X,-Y,1/2+Z                                          
REMARK 290       3555   -X,1/2+Y,1/2-Z                                          
REMARK 290       4555   1/2+X,1/2-Y,-Z                                          
REMARK 290                                                                      
REMARK 290     WHERE NNN -> OPERATOR NUMBER                                     
REMARK 290           MMM -> TRANSLATION VECTOR                                  
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS                            
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM             
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY                
REMARK 290 RELATED MOLECULES.                                                   
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000       31.25950            
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       55.65750            
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       40.07000            
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000       55.65750            
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       31.25950            
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       40.07000            
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000            
REMARK 290                                                                      
REMARK 290 REMARK: NULL                                                         
REMARK 300                                                                      
REMARK 300 BIOMOLECULE: 1                                                       
REMARK 300 THIS ENTRY CONTAINS THE CRYSTALLOGRAPHIC ASYMMETRIC UNIT             
REMARK 300 WHICH CONSISTS OF 4 CHAIN(S). SEE REMARK 350 FOR                     
REMARK 300 INFORMATION ON GENERATING THE BIOLOGICAL MOLECULE(S).                
REMARK 350                                                                      
REMARK 350 GENERATING THE BIOMOLECULE                                           
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN           
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE                
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS          
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND                          
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.                               
REMARK 350                                                                      
REMARK 350 BIOMOLECULE: 1                                                       
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B, C, D                            
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT                     
REMARK 500                                                                      
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.                            
REMARK 500                                                                      
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI                             
REMARK 500  FE    HEM B   147     O    HOH     901              2.13            
REMARK 500  FE    HEM A   142     O    HOH     900              2.14            
REMARK 500  FE    HEM C   142     O    HOH     902              2.14            
REMARK 500  FE    HEM D   147     O    HOH     903              2.14            
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES                                       
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)              
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3                                     
REMARK 500    PHE A  46   N   -  CA  -  C   ANGL. DEV. =  7.0 DEGREES           
REMARK 500    ASP A  47   N   -  CA  -  C   ANGL. DEV. = -9.7 DEGREES           
REMARK 500    LEU A  68   CA  -  CB  -  CG  ANGL. DEV. =  7.3 DEGREES           
REMARK 500    LYS A 139   N   -  CA  -  C   ANGL. DEV. = -7.8 DEGREES           
REMARK 500    TYR A 140   N   -  CA  -  C   ANGL. DEV. =  8.3 DEGREES           
REMARK 500    GLN B   2   N   -  CA  -  C   ANGL. DEV. =  7.6 DEGREES           
REMARK 500    GLU B  22   N   -  CA  -  C   ANGL. DEV. =  7.3 DEGREES           
REMARK 500    PRO B  36   N   -  CA  -  C   ANGL. DEV. =  9.0 DEGREES           
REMARK 500    PHE B 118   N   -  CA  -  C   ANGL. DEV. =  7.3 DEGREES           
REMARK 500    PHE B 122   N   -  CA  -  C   ANGL. DEV. = -7.5 DEGREES           
REMARK 500    TYR B 145   N   -  CA  -  C   ANGL. DEV. = 12.4 DEGREES           
REMARK 500    ASP C  47   N   -  CA  -  C   ANGL. DEV. =-10.7 DEGREES           
REMARK 500    ALA C  88   N   -  CA  -  C   ANGL. DEV. =  7.4 DEGREES           
REMARK 500    GLN D   2   N   -  CA  -  C   ANGL. DEV. =  7.6 DEGREES           
REMARK 500    ASN D  19   N   -  CA  -  C   ANGL. DEV. = -7.9 DEGREES           
REMARK 500    PRO D  36   N   -  CA  -  C   ANGL. DEV. =  7.8 DEGREES           
REMARK 500    PHE D 122   N   -  CA  -  C   ANGL. DEV. = -7.5 DEGREES           
REMARK 500    TYR D 145   N   -  CA  -  C   ANGL. DEV. = -7.6 DEGREES           
REMARK 525                                                                      
REMARK 525 SOLVENT                                                              
REMARK 525 THE FOLLOWING SOLVENT MOLECULES LIE FARTHER THAN EXPECTED            
REMARK 525 FROM THE PROTEIN OR NUCLEIC ACID MOLECULE AND MAY BE                 
REMARK 525 ASSOCIATED WITH A SYMMETRY RELATED MOLECULE (M=MODEL                 
REMARK 525 NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE          
REMARK 525 NUMBER; I=INSERTION CODE):                                           
REMARK 525                                                                      
REMARK 525  M RES CSSEQI                                                        
REMARK 525    HOH   217        DISTANCE =  5.48 ANGSTROMS                       
REMARK 525    HOH   228        DISTANCE =  5.61 ANGSTROMS                       
REMARK 525    HOH   238        DISTANCE =  7.01 ANGSTROMS                       
REMARK 525    HOH   285        DISTANCE =  8.65 ANGSTROMS                       
REMARK 525    HOH   287        DISTANCE =  7.24 ANGSTROMS                       
REMARK 525    HOH   290        DISTANCE =  7.26 ANGSTROMS                       
REMARK 525    HOH   310        DISTANCE =  8.54 ANGSTROMS                       
REMARK 525    HOH   314        DISTANCE =  6.63 ANGSTROMS                       
REMARK 525    HOH   323        DISTANCE =  5.73 ANGSTROMS                       
REMARK 525    HOH   328        DISTANCE =  8.41 ANGSTROMS                       
REMARK 525    HOH   356        DISTANCE =  6.69 ANGSTROMS                       
REMARK 525    HOH   360        DISTANCE =  8.79 ANGSTROMS                       
REMARK 525    HOH   361        DISTANCE =  6.46 ANGSTROMS                       
REMARK 525    HOH   378        DISTANCE =  7.35 ANGSTROMS                       
REMARK 525    HOH   386        DISTANCE =  8.80 ANGSTROMS                       
REMARK 525    HOH   399        DISTANCE =  6.30 ANGSTROMS                       
REMARK 525    HOH   400        DISTANCE =  5.35 ANGSTROMS                       
REMARK 525    HOH   404        DISTANCE =  9.53 ANGSTROMS                       
REMARK 525    HOH   409        DISTANCE =  5.54 ANGSTROMS                       
REMARK 525    HOH   417        DISTANCE =  5.57 ANGSTROMS                       
REMARK 525    HOH   420        DISTANCE =  6.83 ANGSTROMS                       
REMARK 525    HOH   423        DISTANCE =  8.95 ANGSTROMS                       
REMARK 525    HOH   428        DISTANCE =  5.76 ANGSTROMS                       
REMARK 525    HOH   432        DISTANCE =  9.59 ANGSTROMS                       
REMARK 525    HOH   436        DISTANCE =  6.12 ANGSTROMS                       
REMARK 525    HOH   437        DISTANCE =  5.64 ANGSTROMS                       
REMARK 525    HOH   445        DISTANCE =  5.15 ANGSTROMS                       
REMARK 525    HOH   456        DISTANCE =  6.88 ANGSTROMS                       
REMARK 525    HOH   461        DISTANCE =  5.52 ANGSTROMS                       
REMARK 525    HOH   472        DISTANCE =  6.96 ANGSTROMS                       
REMARK 525    HOH   474        DISTANCE =  5.11 ANGSTROMS                       
REMARK 525    HOH   475        DISTANCE =  9.88 ANGSTROMS                       
REMARK 525    HOH   484        DISTANCE =  9.90 ANGSTROMS                       
REMARK 525    HOH   487        DISTANCE =  7.14 ANGSTROMS                       
REMARK 525    HOH   495        DISTANCE =  6.76 ANGSTROMS                       
REMARK 525    HOH   499        DISTANCE =  7.45 ANGSTROMS                       
REMARK 525    HOH   500        DISTANCE = 10.57 ANGSTROMS                       
REMARK 525    HOH   504        DISTANCE = 10.19 ANGSTROMS                       
REMARK 525    HOH   507        DISTANCE =  8.97 ANGSTROMS                       
REMARK 525    HOH   511        DISTANCE =  5.47 ANGSTROMS                       
REMARK 525    HOH   512        DISTANCE =  7.56 ANGSTROMS                       
REMARK 525    HOH   513        DISTANCE =  6.17 ANGSTROMS                       
REMARK 525    HOH   529        DISTANCE =  9.33 ANGSTROMS                       
REMARK 525    HOH   531        DISTANCE =  5.17 ANGSTROMS                       
REMARK 900                                                                      
REMARK 900 RELATED ENTRIES                                                      
REMARK 900 RELATED ID: 1JY7   RELATED DB: PDB                                   
REMARK 900 RELATED ID: 1LFQ   RELATED DB: PDB                                   
REMARK 900 RELATED ID: 1Y8H   RELATED DB: PDB                                   
REMARK 900 RELATED ID: 1Y8I   RELATED DB: PDB                                   
DBREF  1Y8K A    1   141  UNP    P01958   HBA_HORSE        1    141             
DBREF  1Y8K C    1   141  UNP    P01958   HBA_HORSE        1    141             
DBREF  1Y8K B    1   146  UNP    P02062   HBB_HORSE        1    146             
DBREF  1Y8K D    1   146  UNP    P02062   HBB_HORSE        1    146             
SEQADV 1Y8K ASP A   82  UNP  P01958    ASN    82 CONFLICT                       
SEQADV 1Y8K ASN A   85  UNP  P01958    ASP    85 CONFLICT                       
SEQADV 1Y8K ASP C   82  UNP  P01958    ASN    82 CONFLICT                       
SEQADV 1Y8K ASN C   85  UNP  P01958    ASP    85 CONFLICT                       
SEQRES   1 A  141  VAL LEU SER ALA ALA ASP LYS THR ASN VAL LYS ALA ALA          
SEQRES   2 A  141  TRP SER LYS VAL GLY GLY HIS ALA GLY GLU TYR GLY ALA          
SEQRES   3 A  141  GLU ALA LEU GLU ARG MET PHE LEU GLY PHE PRO THR THR          
SEQRES   4 A  141  LYS THR TYR PHE PRO HIS PHE ASP LEU SER HIS GLY SER          
SEQRES   5 A  141  ALA GLN VAL LYS ALA HIS GLY LYS LYS VAL GLY ASP ALA          
SEQRES   6 A  141  LEU THR LEU ALA VAL GLY HIS LEU ASP ASP LEU PRO GLY          
SEQRES   7 A  141  ALA LEU SER ASP LEU SER ASN LEU HIS ALA HIS LYS LEU          
SEQRES   8 A  141  ARG VAL ASP PRO VAL ASN PHE LYS LEU LEU SER HIS CYS          
SEQRES   9 A  141  LEU LEU SER THR LEU ALA VAL HIS LEU PRO ASN ASP PHE          
SEQRES  10 A  141  THR PRO ALA VAL HIS ALA SER LEU ASP LYS PHE LEU SER          
SEQRES  11 A  141  SER VAL SER THR VAL LEU THR SER LYS TYR ARG                  
SEQRES   1 B  146  VAL GLN LEU SER GLY GLU GLU LYS ALA ALA VAL LEU ALA          
SEQRES   2 B  146  LEU TRP ASP LYS VAL ASN GLU GLU GLU VAL GLY GLY GLU          
SEQRES   3 B  146  ALA LEU GLY ARG LEU LEU VAL VAL TYR PRO TRP THR GLN          
SEQRES   4 B  146  ARG PHE PHE ASP SER PHE GLY ASP LEU SER ASN PRO GLY          
SEQRES   5 B  146  ALA VAL MET GLY ASN PRO LYS VAL LYS ALA HIS GLY LYS          
SEQRES   6 B  146  LYS VAL LEU HIS SER PHE GLY GLU GLY VAL HIS HIS LEU          
SEQRES   7 B  146  ASP ASN LEU LYS GLY THR PHE ALA ALA LEU SER GLU LEU          
SEQRES   8 B  146  HIS CYS ASP LYS LEU HIS VAL ASP PRO GLU ASN PHE ARG          
SEQRES   9 B  146  LEU LEU GLY ASN VAL LEU VAL VAL VAL LEU ALA ARG HIS          
SEQRES  10 B  146  PHE GLY LYS ASP PHE THR PRO GLU LEU GLN ALA SER TYR          
SEQRES  11 B  146  GLN LYS VAL VAL ALA GLY VAL ALA ASN ALA LEU ALA HIS          
SEQRES  12 B  146  LYS TYR HIS                                                  
SEQRES   1 C  141  VAL LEU SER ALA ALA ASP LYS THR ASN VAL LYS ALA ALA          
SEQRES   2 C  141  TRP SER LYS VAL GLY GLY HIS ALA GLY GLU TYR GLY ALA          
SEQRES   3 C  141  GLU ALA LEU GLU ARG MET PHE LEU GLY PHE PRO THR THR          
SEQRES   4 C  141  LYS THR TYR PHE PRO HIS PHE ASP LEU SER HIS GLY SER          
SEQRES   5 C  141  ALA GLN VAL LYS ALA HIS GLY LYS LYS VAL GLY ASP ALA          
SEQRES   6 C  141  LEU THR LEU ALA VAL GLY HIS LEU ASP ASP LEU PRO GLY          
SEQRES   7 C  141  ALA LEU SER ASP LEU SER ASN LEU HIS ALA HIS LYS LEU          
SEQRES   8 C  141  ARG VAL ASP PRO VAL ASN PHE LYS LEU LEU SER HIS CYS          
SEQRES   9 C  141  LEU LEU SER THR LEU ALA VAL HIS LEU PRO ASN ASP PHE          
SEQRES  10 C  141  THR PRO ALA VAL HIS ALA SER LEU ASP LYS PHE LEU SER          
SEQRES  11 C  141  SER VAL SER THR VAL LEU THR SER LYS TYR ARG                  
SEQRES   1 D  146  VAL GLN LEU SER GLY GLU GLU LYS ALA ALA VAL LEU ALA          
SEQRES   2 D  146  LEU TRP ASP LYS VAL ASN GLU GLU GLU VAL GLY GLY GLU          
SEQRES   3 D  146  ALA LEU GLY ARG LEU LEU VAL VAL TYR PRO TRP THR GLN          
SEQRES   4 D  146  ARG PHE PHE ASP SER PHE GLY ASP LEU SER ASN PRO GLY          
SEQRES   5 D  146  ALA VAL MET GLY ASN PRO LYS VAL LYS ALA HIS GLY LYS          
SEQRES   6 D  146  LYS VAL LEU HIS SER PHE GLY GLU GLY VAL HIS HIS LEU          
SEQRES   7 D  146  ASP ASN LEU LYS GLY THR PHE ALA ALA LEU SER GLU LEU          
SEQRES   8 D  146  HIS CYS ASP LYS LEU HIS VAL ASP PRO GLU ASN PHE ARG          
SEQRES   9 D  146  LEU LEU GLY ASN VAL LEU VAL VAL VAL LEU ALA ARG HIS          
SEQRES  10 D  146  PHE GLY LYS ASP PHE THR PRO GLU LEU GLN ALA SER TYR          
SEQRES  11 D  146  GLN LYS VAL VAL ALA GLY VAL ALA ASN ALA LEU ALA HIS          
SEQRES  12 D  146  LYS TYR HIS                                                  
HET    HEM  A 142      43                                                       
HET    HEM  B 147      43                                                       
HET    HEM  C 142      43                                                       
HET    HEM  D 147      43                                                       
HETNAM     HEM PROTOPORPHYRIN IX CONTAINING FE                                  
HETSYN     HEM HEME                                                             
FORMUL   5  HEM    4(C34 H32 FE N4 O4)                                          
FORMUL   9  HOH   *386(H2 O)                                                    
HELIX    1   1 SER A    3  GLY A   18  1                                  16    
HELIX    2   2 HIS A   20  PHE A   36  1                                  17    
HELIX    3   3 PRO A   37  PHE A   43  5                                   7    
HELIX    4   4 SER A   52  GLY A   71  1                                  20    
HELIX    5   5 HIS A   72  ASP A   74  5                                   3    
HELIX    6   6 ASP A   75  LEU A   80  1                                   6    
HELIX    7   7 LEU A   80  HIS A   89  1                                  10    
HELIX    8   8 PRO A   95  LEU A  113  1                                  19    
HELIX    9   9 THR A  118  SER A  138  1                                  21    
HELIX   10  10 SER B    4  ASP B   16  1                                  13    
HELIX   11  11 GLU B   21  TYR B   35  1                                  15    
HELIX   12  12 PRO B   36  GLY B   46  5                                  11    
HELIX   13  13 ASN B   50  ASN B   57  1                                   8    
HELIX   14  14 ASN B   57  HIS B   76  1                                  20    
HELIX   15  15 HIS B   77  ASP B   79  5                                   3    
HELIX   16  16 ASN B   80  PHE B   85  1                                   6    
HELIX   17  17 PHE B   85  LYS B   95  1                                  11    
HELIX   18  18 ASP B   99  GLY B  119  1                                  21    
HELIX   19  19 LYS B  120  PHE B  122  5                                   3    
HELIX   20  20 THR B  123  LYS B  144  1                                  22    
HELIX   21  21 SER C    3  GLY C   18  1                                  16    
HELIX   22  22 HIS C   20  PHE C   36  1                                  17    
HELIX   23  23 PRO C   37  PHE C   43  5                                   7    
HELIX   24  24 SER C   52  GLY C   71  1                                  20    
HELIX   25  25 HIS C   72  ASP C   74  5                                   3    
HELIX   26  26 ASP C   75  LEU C   80  1                                   6    
HELIX   27  27 LEU C   80  HIS C   89  1                                  10    
HELIX   28  28 PRO C   95  LEU C  113  1                                  19    
HELIX   29  29 THR C  118  SER C  138  1                                  21    
HELIX   30  30 SER D    4  ASP D   16  1                                  13    
HELIX   31  31 ASN D   19  TYR D   35  1                                  17    
HELIX   32  32 PRO D   36  GLY D   46  5                                  11    
HELIX   33  33 ASN D   50  GLY D   56  1                                   7    
HELIX   34  34 ASN D   57  HIS D   76  1                                  20    
HELIX   35  35 HIS D   77  ASP D   79  5                                   3    
HELIX   36  36 ASN D   80  PHE D   85  1                                   6    
HELIX   37  37 PHE D   85  LYS D   95  1                                  11    
HELIX   38  38 PRO D  100  GLY D  119  1                                  20    
HELIX   39  39 LYS D  120  PHE D  122  5                                   3    
HELIX   40  40 THR D  123  ALA D  142  1                                  20    
LINK         NE2 HIS A  87                FE   HEM A 142                        
LINK         NE2 HIS C  87                FE   HEM C 142                        
LINK         NE2 HIS B  92                FE   HEM B 147                        
LINK         NE2 HIS D  92                FE   HEM D 147                        
CRYST1   62.519   80.140  111.315  90.00  90.00  90.00 P 21 21 21    8          
ORIGX1      1.000000  0.000000  0.000000        0.00000                         
ORIGX2      0.000000  1.000000  0.000000        0.00000                         
ORIGX3      0.000000  0.000000  1.000000        0.00000                         
SCALE1      0.015995  0.000000  0.000000        0.00000                         
SCALE2      0.000000  0.012478  0.000000        0.00000                         
SCALE3      0.000000  0.000000  0.008984        0.00000