PDB entry 1xwd

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HEADER    HORMONE/GROWTH FACTOR                   30-OCT-04   1XWD              
TITLE     CRYSTAL STRUCTURE OF HUMAN FOLLICLE STIMULATING HORMONE               
TITLE    2 COMPLEXED WITH ITS RECEPTOR                                          
COMPND    MOL_ID: 1;                                                            
COMPND   2 MOLECULE: GLYCOPROTEIN HORMONES ALPHA CHAIN;                         
COMPND   3 CHAIN: A, D;                                                         
COMPND   4 SYNONYM: FOLLITROPIN ALPHA CHAIN, FOLLICLE-STIMULATING               
COMPND   5 HORMONE ALPHA CHAIN, FSH-ALPHA, LUTROPIN ALPHA CHAIN,                
COMPND   6 LUTEINIZING HORMONE ALPHA CHAIN, LSH-ALPHA, THYROTROPIN              
COMPND   7 ALPHA CHAIN, THYROID-STIMULATING HORMONE ALPHA CHAIN, TSH-           
COMPND   8 ALPHA, CHORIOGONADOTROPIN ALPHA CHAIN, CHORIONIC                     
COMPND   9 GONADOTROPHIN ALPHA SUBUNIT, CG-ALPHA;                               
COMPND  10 ENGINEERED: YES;                                                     
COMPND  11 MOL_ID: 2;                                                           
COMPND  12 MOLECULE: FOLLITROPIN BETA CHAIN;                                    
COMPND  13 CHAIN: B, E;                                                         
COMPND  14 SYNONYM: FOLLICLE-STIMULATING HORMONE BETA SUBUNIT, FSH-             
COMPND  15 BETA, FSH-B;                                                         
COMPND  16 ENGINEERED: YES;                                                     
COMPND  17 MOL_ID: 3;                                                           
COMPND  18 MOLECULE: FOLLICLE STIMULATING HORMONE RECEPTOR;                     
COMPND  19 CHAIN: C, F;                                                         
COMPND  20 FRAGMENT: EXTRACELLULAR DOMAIN;                                      
COMPND  21 SYNONYM: FSH-R, FOLLITROPIN RECEPTOR;                                
COMPND  22 ENGINEERED: YES                                                      
SOURCE    MOL_ID: 1;                                                            
SOURCE   2 ORGANISM_SCIENTIFIC: HOMO SAPIENS;                                   
SOURCE   3 ORGANISM_COMMON: HUMAN;                                              
SOURCE   4 GENE: CGA;                                                           
SOURCE   5 EXPRESSION_SYSTEM: TRICHOPLUSIA NI;                                  
SOURCE   6 EXPRESSION_SYSTEM_COMMON: CABBAGE LOOPER;                            
SOURCE   7 EXPRESSION_SYSTEM_CELL_LINE: HIGH 5;                                 
SOURCE   8 EXPRESSION_SYSTEM_VECTOR_TYPE: BACULOVIRUS;                          
SOURCE   9 EXPRESSION_SYSTEM_PLASMID: PFASTBAC DUAL;                            
SOURCE  10 MOL_ID: 2;                                                           
SOURCE  11 ORGANISM_SCIENTIFIC: HOMO SAPIENS;                                   
SOURCE  12 ORGANISM_COMMON: HUMAN;                                              
SOURCE  13 GENE: FSHB;                                                          
SOURCE  14 EXPRESSION_SYSTEM: TRICHOPLUSIA NI;                                  
SOURCE  15 EXPRESSION_SYSTEM_COMMON: CABBAGE LOOPER;                            
SOURCE  16 EXPRESSION_SYSTEM_CELL_LINE: HIGH 5;                                 
SOURCE  17 EXPRESSION_SYSTEM_VECTOR_TYPE: BACULOVIRUS;                          
SOURCE  18 EXPRESSION_SYSTEM_PLASMID: PFASTBAC DUAL;                            
SOURCE  19 MOL_ID: 3;                                                           
SOURCE  20 ORGANISM_SCIENTIFIC: HOMO SAPIENS;                                   
SOURCE  21 ORGANISM_COMMON: HUMAN;                                              
SOURCE  22 GENE: FSHR;                                                          
SOURCE  23 EXPRESSION_SYSTEM: TRICHOPLUSIA NI;                                  
SOURCE  24 EXPRESSION_SYSTEM_COMMON: CABBAGE LOOPER;                            
SOURCE  25 EXPRESSION_SYSTEM_CELL_LINE: HIGH 5;                                 
SOURCE  26 EXPRESSION_SYSTEM_VECTOR_TYPE: BACULOVIRUS;                          
SOURCE  27 EXPRESSION_SYSTEM_PLASMID: PFASTBAC DUAL                             
KEYWDS    HORMONE-RECEPTOR COMPLEX, LEUCINE-RICH REPEATS, CYSTEINE-             
KEYWDS   2 KNOT MOTIF                                                           
EXPDTA    X-RAY DIFFRACTION                                                     
AUTHOR    Q.R.FAN,W.A.HENDRICKSON                                               
REVDAT   1   25-JAN-05 1XWD    0                                                
JRNL        AUTH   Q.R.FAN,W.A.HENDRICKSON                                      
JRNL        TITL   STRUCTURE OF HUMAN FOLLICLE-STIMULATING HORMONE IN           
JRNL        TITL 2 COMPLEX WITH ITS RECEPTOR.                                   
JRNL        REF    NATURE                        V. 433   269 2005              
JRNL        REFN   ASTM NATUAS  UK ISSN 0028-0836                               
REMARK   1                                                                      
REMARK   2                                                                      
REMARK   2 RESOLUTION. 2.92 ANGSTROMS.                                          
REMARK   3                                                                      
REMARK   3 REFINEMENT.                                                          
REMARK   3   PROGRAM     : REFMAC 5.2.0003                                      
REMARK   3   AUTHORS     : MURSHUDOV,VAGIN,DODSON                               
REMARK   3                                                                      
REMARK   3    REFINEMENT TARGET : MAXIMUM LIKELIHOOD                            
REMARK   3                                                                      
REMARK   3  DATA USED IN REFINEMENT.                                            
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.92                           
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 25.00                          
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.000                          
REMARK   3   COMPLETENESS FOR RANGE        (%) : 98.0                           
REMARK   3   NUMBER OF REFLECTIONS             : 25282                          
REMARK   3                                                                      
REMARK   3  FIT TO DATA USED IN REFINEMENT.                                     
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT                      
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM                          
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.221                           
REMARK   3   R VALUE            (WORKING SET) : 0.219                           
REMARK   3   FREE R VALUE                     : 0.259                           
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.000                           
REMARK   3   FREE R VALUE TEST SET COUNT      : 1262                            
REMARK   3                                                                      
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.                                  
REMARK   3   TOTAL NUMBER OF BINS USED           : 10                           
REMARK   3   BIN RESOLUTION RANGE HIGH           : 2.92                         
REMARK   3   BIN RESOLUTION RANGE LOW            : 3.08                         
REMARK   3   REFLECTION IN BIN     (WORKING SET) : 3271                         
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 93.95                        
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2970                       
REMARK   3   BIN FREE R VALUE SET COUNT          : 192                          
REMARK   3   BIN FREE R VALUE                    : 0.3560                       
REMARK   3                                                                      
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.                    
REMARK   3   ALL ATOMS                : 7031                                    
REMARK   3                                                                      
REMARK   3  B VALUES.                                                           
REMARK   3   FROM WILSON PLOT           (A**2) : 45.20                          
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 56.30                          
REMARK   3   OVERALL ANISOTROPIC B VALUE.                                       
REMARK   3    B11 (A**2) : 2.72000                                              
REMARK   3    B22 (A**2) : -0.37000                                             
REMARK   3    B33 (A**2) : -2.72000                                             
REMARK   3    B12 (A**2) : 0.00000                                              
REMARK   3    B13 (A**2) : -1.18000                                             
REMARK   3    B23 (A**2) : 0.00000                                              
REMARK   3                                                                      
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.                                 
REMARK   3   ESU BASED ON R VALUE                            (A): NULL          
REMARK   3   ESU BASED ON FREE R VALUE                       (A): 0.419         
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): 0.333         
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): 37.017        
REMARK   3                                                                      
REMARK   3 CORRELATION COEFFICIENTS.                                            
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.922                         
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.888                         
REMARK   3                                                                      
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES        COUNT    RMS    WEIGHT      
REMARK   3   BOND LENGTHS REFINED ATOMS        (A):  7160 ; 0.010 ; 0.021       
REMARK   3   BOND LENGTHS OTHERS               (A):  NULL ;  NULL ;  NULL       
REMARK   3   BOND ANGLES REFINED ATOMS   (DEGREES):  9737 ; 1.427 ; 1.976       
REMARK   3   BOND ANGLES OTHERS          (DEGREES):  NULL ;  NULL ;  NULL       
REMARK   3   TORSION ANGLES, PERIOD 1    (DEGREES):   856 ; 5.304 ; 5.000       
REMARK   3   TORSION ANGLES, PERIOD 2    (DEGREES):   322 ;40.573 ;24.472       
REMARK   3   TORSION ANGLES, PERIOD 3    (DEGREES):  1214 ;19.924 ;15.000       
REMARK   3   TORSION ANGLES, PERIOD 4    (DEGREES):    38 ;16.710 ;15.000       
REMARK   3   CHIRAL-CENTER RESTRAINTS       (A**3):  1120 ; 0.089 ; 0.200       
REMARK   3   GENERAL PLANES REFINED ATOMS      (A):  5292 ; 0.004 ; 0.020       
REMARK   3   GENERAL PLANES OTHERS             (A):  NULL ;  NULL ;  NULL       
REMARK   3   NON-BONDED CONTACTS REFINED ATOMS (A):  2955 ; 0.255 ; 0.200       
REMARK   3   NON-BONDED CONTACTS OTHERS        (A):  NULL ;  NULL ;  NULL       
REMARK   3   NON-BONDED TORSION REFINED ATOMS  (A):  4675 ; 0.323 ; 0.200       
REMARK   3   NON-BONDED TORSION OTHERS         (A):  NULL ;  NULL ;  NULL       
REMARK   3   H-BOND (X...Y) REFINED ATOMS      (A):   214 ; 0.180 ; 0.200       
REMARK   3   H-BOND (X...Y) OTHERS             (A):  NULL ;  NULL ;  NULL       
REMARK   3   POTENTIAL METAL-ION REFINED ATOMS (A):  NULL ;  NULL ;  NULL       
REMARK   3   POTENTIAL METAL-ION OTHERS        (A):  NULL ;  NULL ;  NULL       
REMARK   3   SYMMETRY VDW REFINED ATOMS        (A):    79 ; 0.255 ; 0.200       
REMARK   3   SYMMETRY VDW OTHERS               (A):  NULL ;  NULL ;  NULL       
REMARK   3   SYMMETRY H-BOND REFINED ATOMS     (A):     2 ; 0.064 ; 0.200       
REMARK   3   SYMMETRY H-BOND OTHERS            (A):  NULL ;  NULL ;  NULL       
REMARK   3   SYMMETRY METAL-ION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL       
REMARK   3   SYMMETRY METAL-ION OTHERS         (A):  NULL ;  NULL ;  NULL       
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.     COUNT   RMS    WEIGHT      
REMARK   3   MAIN-CHAIN BOND REFINED ATOMS  (A**2):  4405 ; 1.948 ; 5.000       
REMARK   3   MAIN-CHAIN BOND OTHER ATOMS    (A**2):  NULL ;  NULL ;  NULL       
REMARK   3   MAIN-CHAIN ANGLE REFINED ATOMS (A**2):  7023 ; 2.884 ; 6.000       
REMARK   3   SIDE-CHAIN BOND REFINED ATOMS  (A**2):  3004 ; 2.901 ; 6.000       
REMARK   3   SIDE-CHAIN ANGLE REFINED ATOMS (A**2):  2714 ; 4.234 ; 7.500       
REMARK   3                                                                      
REMARK   3 ANISOTROPIC THERMAL FACTOR RESTRAINTS.    COUNT   RMS   WEIGHT       
REMARK   3   RIGID-BOND RESTRAINTS          (A**2):  NULL ;  NULL ;  NULL       
REMARK   3   SPHERICITY; FREE ATOMS         (A**2):  NULL ;  NULL ;  NULL       
REMARK   3   SPHERICITY; BONDED ATOMS       (A**2):  NULL ;  NULL ;  NULL       
REMARK   3                                                                      
REMARK   3  NCS RESTRAINTS STATISTICS                                           
REMARK   3   NUMBER OF DIFFERENT NCS GROUPS : 3                                 
REMARK   3                                                                      
REMARK   3  NCS GROUP NUMBER               : 1                                  
REMARK   3     CHAIN NAMES                    : C F                             
REMARK   3     NUMBER OF COMPONENTS NCS GROUP : 11                              
REMARK   3       COMPONENT C  SSSEQI  TO  C   SSSEQI   CODE                     
REMARK   3           1     C     18       C      27      6                      
REMARK   3           1     F     18       F      27      6                      
REMARK   3           2     C     28       C      66      1                      
REMARK   3           2     F     28       F      66      1                      
REMARK   3           3     C     67       C      68      6                      
REMARK   3           3     F     67       F      68      6                      
REMARK   3           4     C     69       C      91      1                      
REMARK   3           4     F     69       F      91      1                      
REMARK   3           5     C     92       C      93      6                      
REMARK   3           5     F     92       F      93      6                      
REMARK   3           6     C     94       C     166      1                      
REMARK   3           6     F     94       F     166      1                      
REMARK   3           7     C    167       C     168      6                      
REMARK   3           7     F    167       F     168      6                      
REMARK   3           8     C    169       C     215      1                      
REMARK   3           8     F    169       F     215      1                      
REMARK   3           9     C    216       C     217      6                      
REMARK   3           9     F    216       F     217      6                      
REMARK   3          10     C    218       C     244      1                      
REMARK   3          10     F    218       F     244      1                      
REMARK   3          11     C    245       C     250      6                      
REMARK   3          11     F    245       F     250      6                      
REMARK   3                   GROUP CHAIN        COUNT   RMS     WEIGHT          
REMARK   3   TIGHT POSITIONAL   1    C    (A):   1694 ;  0.37 ;  0.32           
REMARK   3   LOOSE POSITIONAL   1    C    (A):    178 ;  2.94 ; 10.00           
REMARK   3   TIGHT THERMAL      1    C (A**2):   1694 ;  3.66 ;  3.16           
REMARK   3   LOOSE THERMAL      1    C (A**2):    178 ; 16.04 ; 99.90           
REMARK   3                                                                      
REMARK   3  NCS GROUP NUMBER               : 2                                  
REMARK   3     CHAIN NAMES                    : A D                             
REMARK   3     NUMBER OF COMPONENTS NCS GROUP : 10                              
REMARK   3       COMPONENT C  SSSEQI  TO  C   SSSEQI   CODE                     
REMARK   3           1     A      6       A      16      4                      
REMARK   3           1     D      6       D      16      4                      
REMARK   3           2     A     17       A      23      6                      
REMARK   3           2     D     17       D      23      6                      
REMARK   3           3     A     24       A      29      4                      
REMARK   3           3     D     24       D      29      4                      
REMARK   3           4     A     30       A      40      1                      
REMARK   3           4     D     30       D      40      1                      
REMARK   3           5     A     41       A      53      4                      
REMARK   3           5     D     41       D      53      4                      
REMARK   3           6     A     54       A      58      1                      
REMARK   3           6     D     54       D      58      1                      
REMARK   3           7     A     59       A      67      4                      
REMARK   3           7     D     59       D      67      4                      
REMARK   3           8     A     68       A      89      1                      
REMARK   3           8     D     68       D      89      1                      
REMARK   3           9     A     90       A      91      4                      
REMARK   3           9     D     90       D      91      4                      
REMARK   3          10     A     92       A      92      6                      
REMARK   3          10     D     92       D      92      6                      
REMARK   3                   GROUP CHAIN        COUNT   RMS     WEIGHT          
REMARK   3   TIGHT POSITIONAL   2    A    (A):    285 ;  0.40 ;  0.32           
REMARK   3   MEDIUM POSITIONAL  2    A    (A):    325 ;  0.57 ;  0.63           
REMARK   3   LOOSE POSITIONAL   2    A    (A):     60 ;  0.74 ; 10.00           
REMARK   3   TIGHT THERMAL      2    A (A**2):    285 ;  6.64 ;  3.16           
REMARK   3   MEDIUM THERMAL     2    A (A**2):    325 ;  5.23 ;  6.32           
REMARK   3   LOOSE THERMAL      2    A (A**2):     60 ;  4.79 ; 99.90           
REMARK   3                                                                      
REMARK   3  NCS GROUP NUMBER               : 3                                  
REMARK   3     CHAIN NAMES                    : B E                             
REMARK   3     NUMBER OF COMPONENTS NCS GROUP : 14                              
REMARK   3       COMPONENT C  SSSEQI  TO  C   SSSEQI   CODE                     
REMARK   3           1     B      3       B       4      4                      
REMARK   3           1     E      3       E       4      4                      
REMARK   3           2     B      5       B      12      1                      
REMARK   3           2     E      5       E      12      1                      
REMARK   3           3     B     13       B      15      4                      
REMARK   3           3     E     13       E      15      4                      
REMARK   3           4     B     16       B      35      1                      
REMARK   3           4     E     16       E      35      1                      
REMARK   3           5     B     36       B      37      4                      
REMARK   3           5     E     36       E      37      4                      
REMARK   3           6     B     38       B      42      6                      
REMARK   3           6     E     38       E      42      6                      
REMARK   3           7     B     43       B      47      4                      
REMARK   3           7     E     43       E      47      4                      
REMARK   3           8     B     48       B      55      1                      
REMARK   3           8     E     48       E      55      1                      
REMARK   3           9     B     56       B      75      4                      
REMARK   3           9     E     56       E      75      4                      
REMARK   3          10     B     76       B      93      1                      
REMARK   3          10     E     76       E      93      1                      
REMARK   3          11     B     94       B      99      4                      
REMARK   3          11     E     94       E      99      4                      
REMARK   3          12     B    100       B     101      6                      
REMARK   3          12     E    100       E     101      6                      
REMARK   3          13     B    102       B     106      1                      
REMARK   3          13     E    102       E     106      1                      
REMARK   3          14     B    107       B     107      6                      
REMARK   3          14     E    107       E     107      6                      
REMARK   3                   GROUP CHAIN        COUNT   RMS     WEIGHT          
REMARK   3   TIGHT POSITIONAL   3    B    (A):    462 ;  0.39 ;  0.32           
REMARK   3   MEDIUM POSITIONAL  3    B    (A):    297 ;  1.05 ;  0.63           
REMARK   3   LOOSE POSITIONAL   3    B    (A):     59 ;  1.11 ; 10.00           
REMARK   3   TIGHT THERMAL      3    B (A**2):    462 ; 10.01 ;  3.16           
REMARK   3   MEDIUM THERMAL     3    B (A**2):    297 ; 16.02 ;  6.32           
REMARK   3   LOOSE THERMAL      3    B (A**2):     59 ;  9.41 ; 99.90           
REMARK   3                                                                      
REMARK   3  TLS DETAILS                                                         
REMARK   3   NUMBER OF TLS GROUPS  : 2                                          
REMARK   3                                                                      
REMARK   3   TLS GROUP : 1                                                      
REMARK   3    NUMBER OF COMPONENTS GROUP : 3                                    
REMARK   3    COMPONENTS        C SSSEQI   TO  C SSSEQI                         
REMARK   3    RESIDUE RANGE :   A     3        A    92                          
REMARK   3    RESIDUE RANGE :   B     3        B   107                          
REMARK   3    RESIDUE RANGE :   C    18        C   259                          
REMARK   3    ORIGIN FOR THE GROUP (A):  15.6897  -8.8262  51.2914              
REMARK   3    T TENSOR                                                          
REMARK   3      T11:  -0.0210 T22:  -0.0824                                     
REMARK   3      T33:  -0.0387 T12:  -0.0261                                     
REMARK   3      T13:  -0.0480 T23:  -0.0136                                     
REMARK   3    L TENSOR                                                          
REMARK   3      L11:   0.4631 L22:   0.8966                                     
REMARK   3      L33:   1.8794 L12:  -0.2297                                     
REMARK   3      L13:   0.4158 L23:   0.0986                                     
REMARK   3    S TENSOR                                                          
REMARK   3      S11:   0.0367 S12:   0.0033 S13:  -0.0242                       
REMARK   3      S21:   0.0644 S22:   0.0248 S23:  -0.0550                       
REMARK   3      S31:   0.1184 S32:   0.1811 S33:  -0.0615                       
REMARK   3                                                                      
REMARK   3   TLS GROUP : 2                                                      
REMARK   3    NUMBER OF COMPONENTS GROUP : 3                                    
REMARK   3    COMPONENTS        C SSSEQI   TO  C SSSEQI                         
REMARK   3    RESIDUE RANGE :   D     6        D    92                          
REMARK   3    RESIDUE RANGE :   E     3        E   107                          
REMARK   3    RESIDUE RANGE :   F    18        F   250                          
REMARK   3    ORIGIN FOR THE GROUP (A):  19.4870 -43.3557  20.0938              
REMARK   3    T TENSOR                                                          
REMARK   3      T11:  -0.1665 T22:  -0.0380                                     
REMARK   3      T33:  -0.0448 T12:   0.0035                                     
REMARK   3      T13:   0.0194 T23:  -0.0058                                     
REMARK   3    L TENSOR                                                          
REMARK   3      L11:   1.0599 L22:   1.9714                                     
REMARK   3      L33:   2.4145 L12:  -0.1860                                     
REMARK   3      L13:  -0.4899 L23:   1.1100                                     
REMARK   3    S TENSOR                                                          
REMARK   3      S11:   0.0470 S12:   0.2397 S13:  -0.0193                       
REMARK   3      S21:   0.1828 S22:  -0.1388 S23:   0.1307                       
REMARK   3      S31:   0.0829 S32:  -0.1850 S33:   0.0918                       
REMARK   3                                                                      
REMARK   3  BULK SOLVENT MODELLING.                                             
REMARK   3   METHOD USED : BABINET MODEL WITH MASK                              
REMARK   3   PARAMETERS FOR MASK CALCULATION                                    
REMARK   3   VDW PROBE RADIUS   : 1.20                                          
REMARK   3   ION PROBE RADIUS   : 0.80                                          
REMARK   3   SHRINKAGE RADIUS   : 0.80                                          
REMARK   3                                                                      
REMARK   3  OTHER REFINEMENT REMARKS: NULL                                      
REMARK   4                                                                      
REMARK   4 1XWD COMPLIES WITH FORMAT V. 3.0, 1-DEC-2006                         
REMARK   4                                                                      
REMARK   4 THIS IS THE REMEDIATED VERSION OF THIS PDB ENTRY.                    
REMARK   4 REMEDIATED DATA FILE REVISION 3.101 (2007-05-29)                     
REMARK 100                                                                      
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB .                              
REMARK 100 THE RCSB ID CODE IS RCSB030842.                                      
REMARK 200                                                                      
REMARK 200 EXPERIMENTAL DETAILS                                                 
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION                  
REMARK 200  DATE OF DATA COLLECTION        : 26-MAR-2003                        
REMARK 200  TEMPERATURE           (KELVIN) : 100.0                              
REMARK 200  PH                             : 6.40                               
REMARK 200  NUMBER OF CRYSTALS USED        : 1                                  
REMARK 200                                                                      
REMARK 200  SYNCHROTRON              (Y/N) : Y                                  
REMARK 200  RADIATION SOURCE               : APS                                
REMARK 200  BEAMLINE                       : 31-ID                              
REMARK 200  X-RAY GENERATOR MODEL          : NULL                               
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M                                  
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.98                               
REMARK 200  MONOCHROMATOR                  : SI(111)                            
REMARK 200  OPTICS                         : NULL                               
REMARK 200                                                                      
REMARK 200  DETECTOR TYPE                  : CCD                                
REMARK 200  DETECTOR MANUFACTURER          : MARRESEARCH                        
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : DENZO                              
REMARK 200  DATA SCALING SOFTWARE          : SCALEPACK                          
REMARK 200                                                                      
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 25282                              
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.900                              
REMARK 200  RESOLUTION RANGE LOW       (A) : 25.000                             
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : -3.000                             
REMARK 200                                                                      
REMARK 200 OVERALL.                                                             
REMARK 200  COMPLETENESS FOR RANGE     (%) : 98.6                               
REMARK 200  DATA REDUNDANCY                : 3.000                              
REMARK 200  R MERGE                    (I) : 0.06900                            
REMARK 200  R SYM                      (I) : 0.06900                            
REMARK 200   FOR THE DATA SET  : 12.4000                            
REMARK 200                                                                      
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.                                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.90                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 3.00                     
REMARK 200  COMPLETENESS FOR SHELL     (%) : 97.7                               
REMARK 200  DATA REDUNDANCY IN SHELL       : 3.00                               
REMARK 200  R MERGE FOR SHELL          (I) : 0.38800                            
REMARK 200  R SYM FOR SHELL            (I) : 0.38800                            
REMARK 200   FOR SHELL         : 3.400                              
REMARK 200                                                                      
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH                              
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT        
REMARK 200 SOFTWARE USED: AMORE                                                 
REMARK 200 STARTING MODEL: PDB ENTRY 1FL7                                       
REMARK 200                                                                      
REMARK 200 REMARK: NULL                                                         
REMARK 280                                                                      
REMARK 280 CRYSTAL                                                              
REMARK 280 SOLVENT CONTENT, VS   (%): 59.00                                     
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 3.03                     
REMARK 280                                                                      
REMARK 280 CRYSTALLIZATION CONDITIONS: PEG 3350, LITHIUM SULFATE, PH 6.4,       
REMARK 280  VAPOR DIFFUSION, HANGING DROP, TEMPERATURE 293K                     
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY                                            
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: C 1 2 1                          
REMARK 290                                                                      
REMARK 290      SYMOP   SYMMETRY                                                
REMARK 290     NNNMMM   OPERATOR                                                
REMARK 290       1555   X,Y,Z                                                   
REMARK 290       2555   -X,Y,-Z                                                 
REMARK 290       3555   1/2+X,1/2+Y,Z                                           
REMARK 290       4555   1/2-X,1/2+Y,-Z                                          
REMARK 290                                                                      
REMARK 290     WHERE NNN -> OPERATOR NUMBER                                     
REMARK 290           MMM -> TRANSLATION VECTOR                                  
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS                            
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM             
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY                
REMARK 290 RELATED MOLECULES.                                                   
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   2  0.000000  1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   2  0.000000  0.000000 -1.000000        0.00000            
REMARK 290   SMTRY1   3  1.000000  0.000000  0.000000       60.66500            
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       33.46900            
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   4 -1.000000  0.000000  0.000000       60.66500            
REMARK 290   SMTRY2   4  0.000000  1.000000  0.000000       33.46900            
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000            
REMARK 290                                                                      
REMARK 290 REMARK: NULL                                                         
REMARK 300                                                                      
REMARK 300 BIOMOLECULE: 1, 2                                                    
REMARK 300 THIS ENTRY CONTAINS THE CRYSTALLOGRAPHIC ASYMMETRIC UNIT             
REMARK 300 WHICH CONSISTS OF 6 CHAIN(S). SEE REMARK 350 FOR                     
REMARK 300 INFORMATION ON GENERATING THE BIOLOGICAL MOLECULE(S).                
REMARK 350                                                                      
REMARK 350 GENERATING THE BIOMOLECULE                                           
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN           
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE                
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS          
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND                          
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.                               
REMARK 350                                                                      
REMARK 350 BIOMOLECULE: 1                                                       
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B, C, H, I, J                      
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 350 BIOMOLECULE: 2                                                       
REMARK 350 APPLY THE FOLLOWING TO CHAINS: D, E, F, K, L, M                      
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 465                                                                      
REMARK 465 MISSING RESIDUES                                                     
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE                       
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)                
REMARK 465                                                                      
REMARK 465   M RES C SSSEQI                                                     
REMARK 465     ALA A     1                                                      
REMARK 465     PRO A     2                                                      
REMARK 465     ASN B     1                                                      
REMARK 465     SER B     2                                                      
REMARK 465     GLU B   108                                                      
REMARK 465     MET B   109                                                      
REMARK 465     LYS B   110                                                      
REMARK 465     GLU B   111                                                      
REMARK 465     GLY C    17                                                      
REMARK 465     LYS C   260                                                      
REMARK 465     LEU C   261                                                      
REMARK 465     VAL C   262                                                      
REMARK 465     ALA C   263                                                      
REMARK 465     LEU C   264                                                      
REMARK 465     MET C   265                                                      
REMARK 465     GLU C   266                                                      
REMARK 465     ALA C   267                                                      
REMARK 465     SER C   268                                                      
REMARK 465     ALA D     1                                                      
REMARK 465     PRO D     2                                                      
REMARK 465     ASP D     3                                                      
REMARK 465     VAL D     4                                                      
REMARK 465     GLN D     5                                                      
REMARK 465     ASN E     1                                                      
REMARK 465     SER E     2                                                      
REMARK 465     GLU E   108                                                      
REMARK 465     MET E   109                                                      
REMARK 465     LYS E   110                                                      
REMARK 465     GLU E   111                                                      
REMARK 465     GLY F    17                                                      
REMARK 465     ASN F   251                                                      
REMARK 465     LEU F   252                                                      
REMARK 465     LYS F   253                                                      
REMARK 465     LYS F   254                                                      
REMARK 465     LEU F   255                                                      
REMARK 465     PRO F   256                                                      
REMARK 465     THR F   257                                                      
REMARK 465     LEU F   258                                                      
REMARK 465     GLU F   259                                                      
REMARK 465     LYS F   260                                                      
REMARK 465     LEU F   261                                                      
REMARK 465     VAL F   262                                                      
REMARK 465     ALA F   263                                                      
REMARK 465     LEU F   264                                                      
REMARK 465     MET F   265                                                      
REMARK 465     GLU F   266                                                      
REMARK 465     ALA F   267                                                      
REMARK 465     SER F   268                                                      
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT                     
REMARK 500                                                                      
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.                            
REMARK 500                                                                      
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI                             
REMARK 500   ND2  ASN C   191     C2   NAG J   191              1.97            
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND LENGTHS                                      
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,2(A3,1X,A1,I4,A1,1X,A4,3X),F6.3)                  
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   RES CSSEQI ATM2   DEVIATION                     
REMARK 500    PRO B 101   CB    PRO B 101   CG    -0.061                        
REMARK 500    THR C  56   CB    THR C  56   CG2   -0.057                        
REMARK 500    PRO F 120   CB    PRO F 120   CG    -0.072                        
REMARK 500    PRO F 136   CB    PRO F 136   CG    -0.093                        
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES                                       
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)              
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3                                     
REMARK 500    GLN A   5   N   -  CA  -  C   ANGL. DEV. = -9.2 DEGREES           
REMARK 500    ASP A   6   N   -  CA  -  C   ANGL. DEV. = 10.2 DEGREES           
REMARK 500    MET A  71   N   -  CA  -  C   ANGL. DEV. =  9.8 DEGREES           
REMARK 500    ASP B  41   N   -  CA  -  C   ANGL. DEV. =-10.8 DEGREES           
REMARK 500    LEU B  56   CA  -  CB  -  CG  ANGL. DEV. =  9.2 DEGREES           
REMARK 500    HIS C  20   N   -  CA  -  C   ANGL. DEV. =-14.8 DEGREES           
REMARK 500    LEU C 258   N   -  CA  -  C   ANGL. DEV. =-10.1 DEGREES           
DBREF  1XWD A    1    92  UNP    P01215   GLHA_HUMAN      25    116             
DBREF  1XWD D    1    92  UNP    P01215   GLHA_HUMAN      25    116             
DBREF  1XWD B    1   111  UNP    P01225   FSHB_HUMAN      19    129             
DBREF  1XWD E    1   111  UNP    P01225   FSHB_HUMAN      19    129             
DBREF  1XWD C   17   268  UNP    P23945   FSHR_HUMAN      17    268             
DBREF  1XWD F   17   268  UNP    P23945   FSHR_HUMAN      17    268             
SEQRES   1 A   92  ALA PRO ASP VAL GLN ASP CYS PRO GLU CYS THR LEU GLN          
SEQRES   2 A   92  GLU ASN PRO PHE PHE SER GLN PRO GLY ALA PRO ILE LEU          
SEQRES   3 A   92  GLN CYS MET GLY CYS CYS PHE SER ARG ALA TYR PRO THR          
SEQRES   4 A   92  PRO LEU ARG SER LYS LYS THR MET LEU VAL GLN LYS ASN          
SEQRES   5 A   92  VAL THR SER GLU SER THR CYS CYS VAL ALA LYS SER TYR          
SEQRES   6 A   92  ASN ARG VAL THR VAL MET GLY GLY PHE LYS VAL GLU ASN          
SEQRES   7 A   92  HIS THR ALA CYS HIS CYS SER THR CYS TYR TYR HIS LYS          
SEQRES   8 A   92  SER                                                          
SEQRES   1 B  111  ASN SER CYS GLU LEU THR ASN ILE THR ILE ALA ILE GLU          
SEQRES   2 B  111  LYS GLU GLU CYS ARG PHE CYS ILE SER ILE ASN THR THR          
SEQRES   3 B  111  TRP CYS ALA GLY TYR CYS TYR THR ARG ASP LEU VAL TYR          
SEQRES   4 B  111  LYS ASP PRO ALA ARG PRO LYS ILE GLN LYS THR CYS THR          
SEQRES   5 B  111  PHE LYS GLU LEU VAL TYR GLU THR VAL ARG VAL PRO GLY          
SEQRES   6 B  111  CYS ALA HIS HIS ALA ASP SER LEU TYR THR TYR PRO VAL          
SEQRES   7 B  111  ALA THR GLN CYS HIS CYS GLY LYS CYS ASP SER ASP SER          
SEQRES   8 B  111  THR ASP CYS THR VAL ARG GLY LEU GLY PRO SER TYR CYS          
SEQRES   9 B  111  SER PHE GLY GLU MET LYS GLU                                  
SEQRES   1 C  252  GLY CYS HIS HIS ARG ILE CYS HIS CYS SER ASN ARG VAL          
SEQRES   2 C  252  PHE LEU CYS GLN GLU SER LYS VAL THR GLU ILE PRO SER          
SEQRES   3 C  252  ASP LEU PRO ARG ASN ALA ILE GLU LEU ARG PHE VAL LEU          
SEQRES   4 C  252  THR LYS LEU ARG VAL ILE GLN LYS GLY ALA PHE SER GLY          
SEQRES   5 C  252  PHE GLY ASP LEU GLU LYS ILE GLU ILE SER GLN ASN ASP          
SEQRES   6 C  252  VAL LEU GLU VAL ILE GLU ALA ASP VAL PHE SER ASN LEU          
SEQRES   7 C  252  PRO LYS LEU HIS GLU ILE ARG ILE GLU LYS ALA ASN ASN          
SEQRES   8 C  252  LEU LEU TYR ILE ASN PRO GLU ALA PHE GLN ASN LEU PRO          
SEQRES   9 C  252  ASN LEU GLN TYR LEU LEU ILE SER ASN THR GLY ILE LYS          
SEQRES  10 C  252  HIS LEU PRO ASP VAL HIS LYS ILE HIS SER LEU GLN LYS          
SEQRES  11 C  252  VAL LEU LEU ASP ILE GLN ASP ASN ILE ASN ILE HIS THR          
SEQRES  12 C  252  ILE GLU ARG ASN SER PHE VAL GLY LEU SER PHE GLU SER          
SEQRES  13 C  252  VAL ILE LEU TRP LEU ASN LYS ASN GLY ILE GLN GLU ILE          
SEQRES  14 C  252  HIS ASN CYS ALA PHE ASN GLY THR GLN LEU ASP GLU LEU          
SEQRES  15 C  252  ASN LEU SER ASP ASN ASN ASN LEU GLU GLU LEU PRO ASN          
SEQRES  16 C  252  ASP VAL PHE HIS GLY ALA SER GLY PRO VAL ILE LEU ASP          
SEQRES  17 C  252  ILE SER ARG THR ARG ILE HIS SER LEU PRO SER TYR GLY          
SEQRES  18 C  252  LEU GLU ASN LEU LYS LYS LEU ARG ALA ARG SER THR TYR          
SEQRES  19 C  252  ASN LEU LYS LYS LEU PRO THR LEU GLU LYS LEU VAL ALA          
SEQRES  20 C  252  LEU MET GLU ALA SER                                          
SEQRES   1 D   92  ALA PRO ASP VAL GLN ASP CYS PRO GLU CYS THR LEU GLN          
SEQRES   2 D   92  GLU ASN PRO PHE PHE SER GLN PRO GLY ALA PRO ILE LEU          
SEQRES   3 D   92  GLN CYS MET GLY CYS CYS PHE SER ARG ALA TYR PRO THR          
SEQRES   4 D   92  PRO LEU ARG SER LYS LYS THR MET LEU VAL GLN LYS ASN          
SEQRES   5 D   92  VAL THR SER GLU SER THR CYS CYS VAL ALA LYS SER TYR          
SEQRES   6 D   92  ASN ARG VAL THR VAL MET GLY GLY PHE LYS VAL GLU ASN          
SEQRES   7 D   92  HIS THR ALA CYS HIS CYS SER THR CYS TYR TYR HIS LYS          
SEQRES   8 D   92  SER                                                          
SEQRES   1 E  111  ASN SER CYS GLU LEU THR ASN ILE THR ILE ALA ILE GLU          
SEQRES   2 E  111  LYS GLU GLU CYS ARG PHE CYS ILE SER ILE ASN THR THR          
SEQRES   3 E  111  TRP CYS ALA GLY TYR CYS TYR THR ARG ASP LEU VAL TYR          
SEQRES   4 E  111  LYS ASP PRO ALA ARG PRO LYS ILE GLN LYS THR CYS THR          
SEQRES   5 E  111  PHE LYS GLU LEU VAL TYR GLU THR VAL ARG VAL PRO GLY          
SEQRES   6 E  111  CYS ALA HIS HIS ALA ASP SER LEU TYR THR TYR PRO VAL          
SEQRES   7 E  111  ALA THR GLN CYS HIS CYS GLY LYS CYS ASP SER ASP SER          
SEQRES   8 E  111  THR ASP CYS THR VAL ARG GLY LEU GLY PRO SER TYR CYS          
SEQRES   9 E  111  SER PHE GLY GLU MET LYS GLU                                  
SEQRES   1 F  252  GLY CYS HIS HIS ARG ILE CYS HIS CYS SER ASN ARG VAL          
SEQRES   2 F  252  PHE LEU CYS GLN GLU SER LYS VAL THR GLU ILE PRO SER          
SEQRES   3 F  252  ASP LEU PRO ARG ASN ALA ILE GLU LEU ARG PHE VAL LEU          
SEQRES   4 F  252  THR LYS LEU ARG VAL ILE GLN LYS GLY ALA PHE SER GLY          
SEQRES   5 F  252  PHE GLY ASP LEU GLU LYS ILE GLU ILE SER GLN ASN ASP          
SEQRES   6 F  252  VAL LEU GLU VAL ILE GLU ALA ASP VAL PHE SER ASN LEU          
SEQRES   7 F  252  PRO LYS LEU HIS GLU ILE ARG ILE GLU LYS ALA ASN ASN          
SEQRES   8 F  252  LEU LEU TYR ILE ASN PRO GLU ALA PHE GLN ASN LEU PRO          
SEQRES   9 F  252  ASN LEU GLN TYR LEU LEU ILE SER ASN THR GLY ILE LYS          
SEQRES  10 F  252  HIS LEU PRO ASP VAL HIS LYS ILE HIS SER LEU GLN LYS          
SEQRES  11 F  252  VAL LEU LEU ASP ILE GLN ASP ASN ILE ASN ILE HIS THR          
SEQRES  12 F  252  ILE GLU ARG ASN SER PHE VAL GLY LEU SER PHE GLU SER          
SEQRES  13 F  252  VAL ILE LEU TRP LEU ASN LYS ASN GLY ILE GLN GLU ILE          
SEQRES  14 F  252  HIS ASN CYS ALA PHE ASN GLY THR GLN LEU ASP GLU LEU          
SEQRES  15 F  252  ASN LEU SER ASP ASN ASN ASN LEU GLU GLU LEU PRO ASN          
SEQRES  16 F  252  ASP VAL PHE HIS GLY ALA SER GLY PRO VAL ILE LEU ASP          
SEQRES  17 F  252  ILE SER ARG THR ARG ILE HIS SER LEU PRO SER TYR GLY          
SEQRES  18 F  252  LEU GLU ASN LEU LYS LYS LEU ARG ALA ARG SER THR TYR          
SEQRES  19 F  252  ASN LEU LYS LYS LEU PRO THR LEU GLU LYS LEU VAL ALA          
SEQRES  20 F  252  LEU MET GLU ALA SER                                          
MODRES 1XWD ASN A   52  ASN  GLYCOSYLATION SITE                                 
MODRES 1XWD ASN A   78  ASN  GLYCOSYLATION SITE                                 
MODRES 1XWD ASN B    7  ASN  GLYCOSYLATION SITE                                 
MODRES 1XWD ASN B   24  ASN  GLYCOSYLATION SITE                                 
MODRES 1XWD ASN C  191  ASN  GLYCOSYLATION SITE                                 
MODRES 1XWD ASN D   78  ASN  GLYCOSYLATION SITE                                 
MODRES 1XWD ASN E    7  ASN  GLYCOSYLATION SITE                                 
MODRES 1XWD ASN F  191  ASN  GLYCOSYLATION SITE                                 
HET    NAG  H  52      14                                                       
HET    NAG  H  78      14                                                       
HET    NAG  I   7      14                                                       
HET    NAG  I  24      14                                                       
HET    NAG  J 191      14                                                       
HET    NAG  J 192      14                                                       
HET    NAG  K  78      14                                                       
HET    NAG  L   7      14                                                       
HET    NAG  M 191      14                                                       
HET    NAG  M 192      14                                                       
HET    BMA  M 193      11                                                       
HET    SO4    401       5                                                       
HET    SO4    402       5                                                       
HET    SO4    403       5                                                       
HETNAM     NAG N-ACETYL-D-GLUCOSAMINE                                           
HETNAM     BMA BETA-D-MANNOSE                                                   
HETNAM     SO4 SULFATE ION                                                      
HETSYN     NAG NAG                                                              
FORMUL   7  NAG    10(C8 H15 N O6)                                              
FORMUL  14  BMA    C6 H12 O6                                                    
FORMUL  15  SO4    3(O4 S 2-)                                                   
FORMUL  18  HOH   *50(H2 O)                                                     
HELIX    1   1 PRO A   40  LYS A   45  1                                   6    
HELIX    2   2 ARG D   42  THR D   46  5                                   5    
HELIX    3   3 GLU E   15  ARG E   18  5                                   4    
SHEET    1   A 4 THR A  11  GLU A  14  0                                        
SHEET    2   A 4 LEU A  26  PRO A  38 -1  O  GLN A  27   N  GLN A  13           
SHEET    3   A 4 PHE B  19  ARG B  35 -1  O  THR B  34   N  GLY A  30           
SHEET    4   A 4 GLU B   4  LYS B  14 -1  N  THR B   6   O  TRP B  27           
SHEET    1   B 4 THR A  11  GLU A  14  0                                        
SHEET    2   B 4 LEU A  26  PRO A  38 -1  O  GLN A  27   N  GLN A  13           
SHEET    3   B 4 VAL A  53  SER A  57 -1  O  THR A  54   N  TYR A  37           
SHEET    4   B 4 THR B  92  THR B  95  1  O  ASP B  93   N  SER A  55           
SHEET    1   C 2 CYS A  59  VAL A  70  0                                        
SHEET    2   C 2 PHE A  74  SER A  85 -1  O  SER A  85   N  CYS A  59           
SHEET    1   D 2 THR B  50  VAL B  63  0                                        
SHEET    2   D 2 SER B  72  GLY B  85 -1  O  GLY B  85   N  THR B  50           
SHEET    1   E12 CYS C  23  CYS C  25  0                                        
SHEET    2   E12 VAL C  29  GLN C  33 -1  O  LEU C  31   N  HIS C  24           
SHEET    3   E12 GLU C  50  VAL C  54  1  O  ARG C  52   N  PHE C  30           
SHEET    4   E12 LYS C  74  SER C  78  1  O  GLU C  76   N  PHE C  53           
SHEET    5   E12 GLU C  99  GLU C 103  1  O  ARG C 101   N  ILE C  77           
SHEET    6   E12 TYR C 124  SER C 128  1  O  LEU C 126   N  ILE C 100           
SHEET    7   E12 VAL C 147  GLN C 152  1  O  ASP C 150   N  LEU C 125           
SHEET    8   E12 VAL C 173  TRP C 176  1  O  TRP C 176   N  LEU C 149           
SHEET    9   E12 LEU C 195  ASN C 199  1  O  GLU C 197   N  LEU C 175           
SHEET   10   E12 ILE C 222  ASP C 224  1  O  ASP C 224   N  LEU C 198           
SHEET   11   E12 LYS C 243  ARG C 245  1  O  LYS C 243   N  LEU C 223           
SHEET   12   E12 THR C 249  TYR C 250 -1  O  TYR C 250   N  LEU C 244           
SHEET    1   F 3 VAL C  60  ILE C  61  0                                        
SHEET    2   F 3 VAL C  85  ILE C  86  1  O  VAL C  85   N  ILE C  61           
SHEET    3   F 3 TYR C 110  ILE C 111  1  O  TYR C 110   N  ILE C  86           
SHEET    1   G 2 PHE C  91  SER C  92  0                                        
SHEET    2   G 2 PHE C 116  GLN C 117  1  O  GLN C 117   N  PHE C  91           
SHEET    1   H 2 THR C 159  ILE C 160  0                                        
SHEET    2   H 2 GLU C 184  ILE C 185  1  O  GLU C 184   N  ILE C 160           
SHEET    1   I 4 THR D  11  GLU D  14  0                                        
SHEET    2   I 4 LEU D  26  PRO D  38 -1  O  GLN D  27   N  GLN D  13           
SHEET    3   I 4 CYS E  20  ARG E  35 -1  O  THR E  34   N  GLY D  30           
SHEET    4   I 4 GLU E   4  GLU E  13 -1  N  THR E   6   O  TRP E  27           
SHEET    1   J 4 THR D  11  GLU D  14  0                                        
SHEET    2   J 4 LEU D  26  PRO D  38 -1  O  GLN D  27   N  GLN D  13           
SHEET    3   J 4 VAL D  53  SER D  57 -1  O  THR D  54   N  TYR D  37           
SHEET    4   J 4 THR E  92  THR E  95  1  O  THR E  95   N  SER D  55           
SHEET    1   K 2 CYS D  59  THR D  69  0                                        
SHEET    2   K 2 LYS D  75  SER D  85 -1  O  ASN D  78   N  ASN D  66           
SHEET    1   L 2 THR E  50  VAL E  63  0                                        
SHEET    2   L 2 SER E  72  GLY E  85 -1  O  GLN E  81   N  LYS E  54           
SHEET    1   M11 HIS F  24  CYS F  25  0                                        
SHEET    2   M11 VAL F  29  GLN F  33 -1  O  LEU F  31   N  HIS F  24           
SHEET    3   M11 GLU F  50  VAL F  54  1  O  ARG F  52   N  CYS F  32           
SHEET    4   M11 LYS F  74  SER F  78  1  O  GLU F  76   N  LEU F  51           
SHEET    5   M11 GLU F  99  ALA F 105  1  O  ARG F 101   N  ILE F  77           
SHEET    6   M11 TYR F 124  THR F 130  1  O  LEU F 126   N  ILE F 100           
SHEET    7   M11 VAL F 147  GLN F 152  1  O  ASP F 150   N  ILE F 127           
SHEET    8   M11 SER F 172  TRP F 176  1  O  ILE F 174   N  VAL F 147           
SHEET    9   M11 GLN F 194  ASN F 199  1  O  GLU F 197   N  LEU F 175           
SHEET   10   M11 ILE F 222  ASP F 224  1  O  ILE F 222   N  LEU F 198           
SHEET   11   M11 LYS F 243  ARG F 245  1  O  LYS F 243   N  LEU F 223           
SHEET    1   N 3 VAL F  60  ILE F  61  0                                        
SHEET    2   N 3 VAL F  85  ILE F  86  1  O  VAL F  85   N  ILE F  61           
SHEET    3   N 3 TYR F 110  ILE F 111  1  O  TYR F 110   N  ILE F  86           
SHEET    1   O 2 PHE F  91  SER F  92  0                                        
SHEET    2   O 2 PHE F 116  GLN F 117  1  O  GLN F 117   N  PHE F  91           
SHEET    1   P 2 THR F 159  ILE F 160  0                                        
SHEET    2   P 2 GLU F 184  ILE F 185  1  O  GLU F 184   N  ILE F 160           
SSBOND   1 CYS A    7    CYS A   31                                             
SSBOND   2 CYS A   10    CYS A   60                                             
SSBOND   3 CYS A   28    CYS A   82                                             
SSBOND   4 CYS A   32    CYS A   84                                             
SSBOND   5 CYS A   59    CYS A   87                                             
SSBOND   6 CYS B    3    CYS B   51                                             
SSBOND   7 CYS B   17    CYS B   66                                             
SSBOND   8 CYS B   20    CYS B  104                                             
SSBOND   9 CYS B   28    CYS B   82                                             
SSBOND  10 CYS B   32    CYS B   84                                             
SSBOND  11 CYS B   87    CYS B   94                                             
SSBOND  12 CYS C   18    CYS C   25                                             
SSBOND  13 CYS C   23    CYS C   32                                             
SSBOND  14 CYS D    7    CYS D   31                                             
SSBOND  15 CYS D   10    CYS D   60                                             
SSBOND  16 CYS D   28    CYS D   82                                             
SSBOND  17 CYS D   32    CYS D   84                                             
SSBOND  18 CYS D   59    CYS D   87                                             
SSBOND  19 CYS E    3    CYS E   51                                             
SSBOND  20 CYS E   17    CYS E   66                                             
SSBOND  21 CYS E   20    CYS E  104                                             
SSBOND  22 CYS E   28    CYS E   82                                             
SSBOND  23 CYS E   32    CYS E   84                                             
SSBOND  24 CYS E   87    CYS E   94                                             
SSBOND  25 CYS F   18    CYS F   25                                             
SSBOND  26 CYS F   23    CYS F   32                                             
LINK         C1  NAG H  52                 ND2 ASN A  52                        
LINK         C1  NAG H  78                 ND2 ASN A  78                        
LINK         C1  NAG I   7                 ND2 ASN B   7                        
LINK         C1  NAG I  24                 ND2 ASN B  24                        
LINK         C1  NAG J 191                 ND2 ASN C 191                        
LINK         C1  NAG K  78                 ND2 ASN D  78                        
LINK         C1  NAG L   7                 ND2 ASN E   7                        
LINK         C1  NAG M 191                 ND2 ASN F 191                        
LINK         O4  NAG J 191                 C1  NAG J 192                        
LINK         O4  NAG M 191                 C1  NAG M 192                        
LINK         O4  NAG M 192                 C1  BMA M 193                        
CRYST1  121.330   66.938  148.629  90.00  99.13  90.00 C 1 2 1       8          
ORIGX1      1.000000  0.000000  0.000000        0.00000                         
ORIGX2      0.000000  1.000000  0.000000        0.00000                         
ORIGX3      0.000000  0.000000  1.000000        0.00000                         
SCALE1      0.008242  0.000000  0.001325        0.00000                         
SCALE2      0.000000  0.014939  0.000000        0.00000                         
SCALE3      0.000000  0.000000  0.006814        0.00000