PDB entry 1xq5

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HEADER    OXYGEN STORAGE/TRANSPORT                11-OCT-04   1XQ5              
TITLE     MET-PERCH HEMOGLOBIN AT 1.9A                                          
COMPND    MOL_ID: 1;                                                            
COMPND   2 MOLECULE: HEMOGLOBIN ALPHA-1 CHAIN;                                  
COMPND   3 CHAIN: A, C;                                                         
COMPND   4 MOL_ID: 2;                                                           
COMPND   5 MOLECULE: HEMOGLOBIN BETA-2 CHAIN;                                   
COMPND   6 CHAIN: B, D                                                          
SOURCE    MOL_ID: 1;                                                            
SOURCE   2 ORGANISM_SCIENTIFIC: PERCA FLAVESCENS;                               
SOURCE   3 ORGANISM_COMMON: YELLOW PERCH;                                       
SOURCE   4 MOL_ID: 2;                                                           
SOURCE   5 ORGANISM_SCIENTIFIC: PERCA FLAVESCENS;                               
SOURCE   6 ORGANISM_COMMON: YELLOW PERCH                                        
KEYWDS    FISH HEMOGLOBIN, RAPID OXIDATION, STRUCTURAL GENOMICS,                
KEYWDS   2 PROTEIN STRUCTURE INITIATIVE, PSI, CESG, CENTER FOR                  
KEYWDS   3 EUKARYOTIC STRUCTURAL GENOMICS                                       
EXPDTA    X-RAY DIFFRACTION                                                     
AUTHOR    CENTER FOR EUKARYOTIC STRUCTURAL GENOMICS (CESG)                      
REVDAT   2   01-FEB-05 1XQ5    1       KEYWDS REMARK                            
REVDAT   1   16-NOV-04 1XQ5    0                                                
JRNL        AUTH   CENTER FOR EUKARYOTIC STRUCTURAL GENOMICS (CESG)             
JRNL        TITL   MET-PERCH HEMOGLOBIN AT 1.9A                                 
JRNL        REF    TO BE PUBLISHED                                              
JRNL        REFN                                                                
REMARK   1                                                                      
REMARK   2                                                                      
REMARK   2 RESOLUTION. 1.90 ANGSTROMS.                                          
REMARK   3                                                                      
REMARK   3 REFINEMENT.                                                          
REMARK   3   PROGRAM     : REFMAC 5.2.0005                                      
REMARK   3   AUTHORS     : MURSHUDOV,VAGIN,DODSON                               
REMARK   3                                                                      
REMARK   3    REFINEMENT TARGET : MAXIMUM LIKELIHOOD                            
REMARK   3                                                                      
REMARK   3  DATA USED IN REFINEMENT.                                            
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.90                           
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 69.01                          
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.000                          
REMARK   3   COMPLETENESS FOR RANGE        (%) : 96.3                           
REMARK   3   NUMBER OF REFLECTIONS             : 43714                          
REMARK   3                                                                      
REMARK   3  FIT TO DATA USED IN REFINEMENT.                                     
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT                      
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM                          
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.246                           
REMARK   3   R VALUE            (WORKING SET) : 0.243                           
REMARK   3   FREE R VALUE                     : 0.297                           
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.100                           
REMARK   3   FREE R VALUE TEST SET COUNT      : 2327                            
REMARK   3                                                                      
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.                                  
REMARK   3   TOTAL NUMBER OF BINS USED           : 20                           
REMARK   3   BIN RESOLUTION RANGE HIGH           : 1.90                         
REMARK   3   BIN RESOLUTION RANGE LOW            : 1.95                         
REMARK   3   REFLECTION IN BIN     (WORKING SET) : 3128                         
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 94.99                        
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2530                       
REMARK   3   BIN FREE R VALUE SET COUNT          : 168                          
REMARK   3   BIN FREE R VALUE                    : 0.3220                       
REMARK   3                                                                      
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.                    
REMARK   3   ALL ATOMS                : 4867                                    
REMARK   3                                                                      
REMARK   3  B VALUES.                                                           
REMARK   3   FROM WILSON PLOT           (A**2) : NULL                           
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 33.05                          
REMARK   3   OVERALL ANISOTROPIC B VALUE.                                       
REMARK   3    B11 (A**2) : 1.54000                                              
REMARK   3    B22 (A**2) : -1.58000                                             
REMARK   3    B33 (A**2) : 0.04000                                              
REMARK   3    B12 (A**2) : 0.00000                                              
REMARK   3    B13 (A**2) : 0.00000                                              
REMARK   3    B23 (A**2) : 0.00000                                              
REMARK   3                                                                      
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.                                 
REMARK   3   ESU BASED ON R VALUE                            (A): 0.213         
REMARK   3   ESU BASED ON FREE R VALUE                       (A): 0.194         
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): 0.141         
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): 4.633         
REMARK   3                                                                      
REMARK   3 CORRELATION COEFFICIENTS.                                            
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.927                         
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.893                         
REMARK   3                                                                      
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES        COUNT    RMS    WEIGHT      
REMARK   3   BOND LENGTHS REFINED ATOMS        (A):  4777 ; 0.018 ; 0.022       
REMARK   3   BOND LENGTHS OTHERS               (A):     1 ; 0.001 ; 0.020       
REMARK   3   BOND ANGLES REFINED ATOMS   (DEGREES):  6533 ; 1.809 ; 2.068       
REMARK   3   BOND ANGLES OTHERS          (DEGREES):     2 ; 0.807 ; 3.000       
REMARK   3   TORSION ANGLES, PERIOD 1    (DEGREES):   572 ; 6.887 ; 5.000       
REMARK   3   TORSION ANGLES, PERIOD 2    (DEGREES):   184 ;41.365 ;24.022       
REMARK   3   TORSION ANGLES, PERIOD 3    (DEGREES):   773 ;18.947 ;15.000       
REMARK   3   TORSION ANGLES, PERIOD 4    (DEGREES):    16 ;16.963 ;15.000       
REMARK   3   CHIRAL-CENTER RESTRAINTS       (A**3):   712 ; 0.138 ; 0.200       
REMARK   3   GENERAL PLANES REFINED ATOMS      (A):  3565 ; 0.007 ; 0.020       
REMARK   3   GENERAL PLANES OTHERS             (A):     1 ; 0.001 ; 0.020       
REMARK   3   NON-BONDED CONTACTS REFINED ATOMS (A):  2545 ; 0.244 ; 0.200       
REMARK   3   NON-BONDED CONTACTS OTHERS        (A):     1 ; 0.311 ; 0.200       
REMARK   3   NON-BONDED TORSION REFINED ATOMS  (A):  3340 ; 0.316 ; 0.200       
REMARK   3   NON-BONDED TORSION OTHERS         (A):     2 ; 0.341 ; 0.200       
REMARK   3   H-BOND (X...Y) REFINED ATOMS      (A):   233 ; 0.175 ; 0.200       
REMARK   3   H-BOND (X...Y) OTHERS             (A):  NULL ;  NULL ;  NULL       
REMARK   3   POTENTIAL METAL-ION REFINED ATOMS (A):  NULL ;  NULL ;  NULL       
REMARK   3   POTENTIAL METAL-ION OTHERS        (A):  NULL ;  NULL ;  NULL       
REMARK   3   SYMMETRY VDW REFINED ATOMS        (A):    53 ; 0.260 ; 0.200       
REMARK   3   SYMMETRY VDW OTHERS               (A):  NULL ;  NULL ;  NULL       
REMARK   3   SYMMETRY H-BOND REFINED ATOMS     (A):     5 ; 0.169 ; 0.200       
REMARK   3   SYMMETRY H-BOND OTHERS            (A):  NULL ;  NULL ;  NULL       
REMARK   3   SYMMETRY METAL-ION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL       
REMARK   3   SYMMETRY METAL-ION OTHERS         (A):  NULL ;  NULL ;  NULL       
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.     COUNT   RMS    WEIGHT      
REMARK   3   MAIN-CHAIN BOND REFINED ATOMS  (A**2):  2934 ; 1.780 ; 2.000       
REMARK   3   MAIN-CHAIN BOND OTHER ATOMS    (A**2):     1 ; 0.080 ; 2.000       
REMARK   3   MAIN-CHAIN ANGLE REFINED ATOMS (A**2):  4603 ; 3.036 ; 4.000       
REMARK   3   SIDE-CHAIN BOND REFINED ATOMS  (A**2):  2126 ; 5.027 ; 6.000       
REMARK   3   SIDE-CHAIN ANGLE REFINED ATOMS (A**2):  1922 ; 6.581 ; 8.000       
REMARK   3                                                                      
REMARK   3 ANISOTROPIC THERMAL FACTOR RESTRAINTS.    COUNT   RMS   WEIGHT       
REMARK   3   RIGID-BOND RESTRAINTS          (A**2):  NULL ;  NULL ;  NULL       
REMARK   3   SPHERICITY; FREE ATOMS         (A**2):  NULL ;  NULL ;  NULL       
REMARK   3   SPHERICITY; BONDED ATOMS       (A**2):  NULL ;  NULL ;  NULL       
REMARK   3                                                                      
REMARK   3  NCS RESTRAINTS STATISTICS                                           
REMARK   3   NUMBER OF DIFFERENT NCS GROUPS : 2                                 
REMARK   3                                                                      
REMARK   3  NCS GROUP NUMBER               : 1                                  
REMARK   3     CHAIN NAMES                    : A C                             
REMARK   3     NUMBER OF COMPONENTS NCS GROUP : 1                               
REMARK   3       COMPONENT C  SSSEQI  TO  C   SSSEQI   CODE                     
REMARK   3           1     A      1       A     142      2                      
REMARK   3           1     C      1       C     142      2                      
REMARK   3                   GROUP CHAIN        COUNT   RMS     WEIGHT          
REMARK   3   TIGHT POSITIONAL   1    A    (A):    564 ;  0.08 ;  0.05           
REMARK   3   MEDIUM POSITIONAL  1    A    (A):    532 ;  0.53 ;  0.50           
REMARK   3   TIGHT THERMAL      1    A (A**2):    564 ;  0.38 ;  0.50           
REMARK   3   MEDIUM THERMAL     1    A (A**2):    532 ;  1.72 ;  2.00           
REMARK   3                                                                      
REMARK   3  NCS GROUP NUMBER               : 2                                  
REMARK   3     CHAIN NAMES                    : B D                             
REMARK   3     NUMBER OF COMPONENTS NCS GROUP : 1                               
REMARK   3       COMPONENT C  SSSEQI  TO  C   SSSEQI   CODE                     
REMARK   3           1     B     16       B     129      2                      
REMARK   3           1     D     16       D     129      2                      
REMARK   3                   GROUP CHAIN        COUNT   RMS     WEIGHT          
REMARK   3   TIGHT POSITIONAL   2    B    (A):    456 ;  0.07 ;  0.05           
REMARK   3   MEDIUM POSITIONAL  2    B    (A):    413 ;  0.37 ;  0.50           
REMARK   3   TIGHT THERMAL      2    B (A**2):    456 ;  0.34 ;  0.50           
REMARK   3   MEDIUM THERMAL     2    B (A**2):    413 ;  1.79 ;  2.00           
REMARK   3                                                                      
REMARK   3  TLS DETAILS                                                         
REMARK   3   NUMBER OF TLS GROUPS  : 0                                          
REMARK   3                                                                      
REMARK   3  BULK SOLVENT MODELLING.                                             
REMARK   3   METHOD USED : BABINET MODEL WITH MASK                              
REMARK   3   PARAMETERS FOR MASK CALCULATION                                    
REMARK   3   VDW PROBE RADIUS   : 1.20                                          
REMARK   3   ION PROBE RADIUS   : 0.80                                          
REMARK   3   SHRINKAGE RADIUS   : 0.80                                          
REMARK   3                                                                      
REMARK   3  OTHER REFINEMENT REMARKS: HYDROGENS HAVE BEEN ADDED IN THE          
REMARK   3  RIDING POSITIONS                                                    
REMARK   4                                                                      
REMARK   4 1XQ5 COMPLIES WITH FORMAT V. 3.0, 1-DEC-2006                         
REMARK   4                                                                      
REMARK   4 THIS IS THE REMEDIATED VERSION OF THIS PDB ENTRY.                    
REMARK   4 REMEDIATED DATA FILE REVISION 3.101 (2007-05-29)                     
REMARK 100                                                                      
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB .                              
REMARK 100 THE RCSB ID CODE IS RCSB030635.                                      
REMARK 200                                                                      
REMARK 200 EXPERIMENTAL DETAILS                                                 
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION                  
REMARK 200  DATE OF DATA COLLECTION        : 31-JUL-2004                        
REMARK 200  TEMPERATURE           (KELVIN) : 110.0                              
REMARK 200  PH                             : 7.00                               
REMARK 200  NUMBER OF CRYSTALS USED        : 1                                  
REMARK 200                                                                      
REMARK 200  SYNCHROTRON              (Y/N) : Y                                  
REMARK 200  RADIATION SOURCE               : APS                                
REMARK 200  BEAMLINE                       : 19-BM                              
REMARK 200  X-RAY GENERATOR MODEL          : NULL                               
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M                                  
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.97934                            
REMARK 200  MONOCHROMATOR                  : NULL                               
REMARK 200  OPTICS                         : NULL                               
REMARK 200                                                                      
REMARK 200  DETECTOR TYPE                  : CCD                                
REMARK 200  DETECTOR MANUFACTURER          : APS-1                              
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : HKL-2000                           
REMARK 200  DATA SCALING SOFTWARE          : SCALEPACK                          
REMARK 200                                                                      
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 43714                              
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.898                              
REMARK 200  RESOLUTION RANGE LOW       (A) : 69.007                             
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : -1.000                             
REMARK 200                                                                      
REMARK 200 OVERALL.                                                             
REMARK 200  COMPLETENESS FOR RANGE     (%) : 96.3                               
REMARK 200  DATA REDUNDANCY                : 6.500                              
REMARK 200  R MERGE                    (I) : 0.08400                            
REMARK 200  R SYM                      (I) : NULL                               
REMARK 200   FOR THE DATA SET  : 15.0960                            
REMARK 200                                                                      
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.                                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.90                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.97                     
REMARK 200  COMPLETENESS FOR SHELL     (%) : 96.1                               
REMARK 200  DATA REDUNDANCY IN SHELL       : 5.90                               
REMARK 200  R MERGE FOR SHELL          (I) : 0.52100                            
REMARK 200  R SYM FOR SHELL            (I) : NULL                               
REMARK 200   FOR SHELL         : 4.027                              
REMARK 200                                                                      
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH                              
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT        
REMARK 200 SOFTWARE USED: AMORE                                                 
REMARK 200 STARTING MODEL: PDB ENTRY 1OUU                                       
REMARK 200                                                                      
REMARK 200 REMARK: NULL                                                         
REMARK 280                                                                      
REMARK 280 CRYSTAL                                                              
REMARK 280 SOLVENT CONTENT, VS   (%): 46.00                                     
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.16                     
REMARK 280                                                                      
REMARK 280 CRYSTALLIZATION CONDITIONS: PEG 4K, HEPES, PH 7.0, BATCH,            
REMARK 280  TEMPERATURE 296K                                                    
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY                                            
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 21                       
REMARK 290                                                                      
REMARK 290      SYMOP   SYMMETRY                                                
REMARK 290     NNNMMM   OPERATOR                                                
REMARK 290       1555   X,Y,Z                                                   
REMARK 290       2555   1/2-X,-Y,1/2+Z                                          
REMARK 290       3555   -X,1/2+Y,1/2-Z                                          
REMARK 290       4555   1/2+X,1/2-Y,-Z                                          
REMARK 290                                                                      
REMARK 290     WHERE NNN -> OPERATOR NUMBER                                     
REMARK 290           MMM -> TRANSLATION VECTOR                                  
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS                            
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM             
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY                
REMARK 290 RELATED MOLECULES.                                                   
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000       28.65100            
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       60.88950            
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       42.49300            
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000       60.88950            
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       28.65100            
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       42.49300            
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000            
REMARK 290                                                                      
REMARK 290 REMARK: NULL                                                         
REMARK 300                                                                      
REMARK 300 BIOMOLECULE: 1                                                       
REMARK 300 THIS ENTRY CONTAINS THE CRYSTALLOGRAPHIC ASYMMETRIC UNIT             
REMARK 300 WHICH CONSISTS OF 4 CHAIN(S). SEE REMARK 350 FOR                     
REMARK 300 INFORMATION ON GENERATING THE BIOLOGICAL MOLECULE(S).                
REMARK 350                                                                      
REMARK 350 GENERATING THE BIOMOLECULE                                           
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN           
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE                
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS          
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND                          
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.                               
REMARK 350                                                                      
REMARK 350 BIOMOLECULE: 1                                                       
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B, C, D                            
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT                     
REMARK 500                                                                      
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.                            
REMARK 500                                                                      
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI                             
REMARK 500   OE1  GLU D   125     O    HOH     275              2.05            
REMARK 500  FE    HEM B   148     O    HOH     379              2.10            
REMARK 500  FE    HEM C   143     O    HOH     380              2.14            
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES                                       
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)              
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3                                     
REMARK 500    ASP A  48   C   -  N   -  CA  ANGL. DEV. = 14.1 DEGREES           
REMARK 500    VAL A  68   CB  -  CA  -  C   ANGL. DEV. = 11.0 DEGREES           
REMARK 500    LEU A  87   CA  -  CB  -  CG  ANGL. DEV. = 12.0 DEGREES           
REMARK 500    LEU A 133   CB  -  CG  -  CD2 ANGL. DEV. =-13.5 DEGREES           
REMARK 500    LEU B 141   N   -  CA  -  C   ANGL. DEV. = 12.2 DEGREES           
REMARK 500    ASP C   8   CB  -  CA  -  C   ANGL. DEV. = 10.6 DEGREES           
REMARK 500    ILE C  67   CG1 -  CB  -  CG2 ANGL. DEV. =-15.1 DEGREES           
REMARK 500    LEU C 133   CB  -  CG  -  CD2 ANGL. DEV. =-10.7 DEGREES           
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: TORSION ANGLES                                             
REMARK 500                                                                      
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:            
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;               
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                             
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)                    
REMARK 500                                                                      
REMARK 500  M RES CSSEQI        PSI       PHI                                   
REMARK 500    ASP A  48       62.92    121.92                                   
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: NON-CIS, NON-TRANS                                         
REMARK 500                                                                      
REMARK 500 THE FOLLOWING PEPTIDE BONDS DEVIATE SIGNIFICANTLY FROM BOTH          
REMARK 500 CIS AND TRANS CONFORMATION.  CIS BONDS, IF ANY, ARE LISTED           
REMARK 500 ON CISPEP RECORDS.  TRANS IS DEFINED AS 180 +/- 30 AND               
REMARK 500 CIS IS DEFINED AS 0 +/- 30 DEGREES.                                  
REMARK 500                                 MODEL     OMEGA                      
REMARK 500 LYS A   47    ASP A   48                  -49.52                     
REMARK 500 LEU B  141    GLY B  142                   78.06                     
REMARK 525                                                                      
REMARK 525 SOLVENT                                                              
REMARK 525 THE FOLLOWING SOLVENT MOLECULES LIE FARTHER THAN EXPECTED            
REMARK 525 FROM THE PROTEIN OR NUCLEIC ACID MOLECULE AND MAY BE                 
REMARK 525 ASSOCIATED WITH A SYMMETRY RELATED MOLECULE (M=MODEL                 
REMARK 525 NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE          
REMARK 525 NUMBER; I=INSERTION CODE):                                           
REMARK 525                                                                      
REMARK 525  M RES CSSEQI                                                        
REMARK 525    HOH   274        DISTANCE =  7.65 ANGSTROMS                       
REMARK 525    HOH   299        DISTANCE =  9.39 ANGSTROMS                       
REMARK 900                                                                      
REMARK 900 RELATED ENTRIES                                                      
REMARK 900 RELATED ID: PERCH HEMOGLOBIN   RELATED DB: TARGETDB                  
REMARK 900 RELATED ID: PERCH   RELATED DB: TARGETDB                             
REMARK 900 RELATED ID: HEMOGLOBIN   RELATED DB: TARGETDB                        
REMARK 999                                                                      
REMARK 999 SEQUENCE                                                             
REMARK 999 THE SEQUENCE OF THE PROTEIN WAS NOT DEPOSITED                        
REMARK 999 INTO ANY SEQUENCE DATABASE.                                          
SEQRES   1 A  142  SER LEU SER SER LYS ASP LYS ASP THR VAL LYS ALA LEU          
SEQRES   2 A  142  TRP GLY LYS ILE ALA ASP LYS ALA GLU GLU ILE GLY SER          
SEQRES   3 A  142  ASP ALA LEU SER ARG MET LEU ALA VAL TYR PRO GLN THR          
SEQRES   4 A  142  LYS THR TYR PHE SER HIS TRP LYS ASP LEU SER PRO GLY          
SEQRES   5 A  142  SER ALA PRO VAL ASN LYS HIS GLY LYS THR ILE MET GLY          
SEQRES   6 A  142  GLY ILE VAL ASP ALA VAL ALA SER ILE ASP ASP LEU ASN          
SEQRES   7 A  142  ALA GLY LEU LEU ALA LEU SER GLU LEU HIS ALA PHE THR          
SEQRES   8 A  142  LEU ARG VAL ASP PRO ALA ASN PHE LYS ILE LEU SER HIS          
SEQRES   9 A  142  CYS ILE LEU VAL LEU LEU ALA VAL LYS PHE PRO LYS ASP          
SEQRES  10 A  142  PHE THR PRO GLU VAL HIS ILE SER TYR ASP LYS PHE PHE          
SEQRES  11 A  142  SER ALA LEU ALA ARG ALA LEU ALA GLU LYS TYR ARG              
SEQRES   1 B  146  VAL VAL TRP THR ASP PHE GLU ARG ALA THR ILE ALA ASP          
SEQRES   2 B  146  ILE PHE SER LYS LEU ASP TYR GLU ALA VAL GLY GLY ALA          
SEQRES   3 B  146  THR LEU ALA ARG CYS LEU ILE VAL TYR PRO TRP THR GLN          
SEQRES   4 B  146  ARG TYR PHE GLY ASN PHE GLY ASN LEU TYR ASN ALA ALA          
SEQRES   5 B  146  ALA ILE MET GLY ASN PRO MET ILE ALA LYS HIS GLY THR          
SEQRES   6 B  146  THR ILE LEU HIS GLY LEU ASP ARG ALA VAL LYS ASN MET          
SEQRES   7 B  146  ASP ASN ILE LYS ALA THR TYR ALA GLU LEU SER VAL LEU          
SEQRES   8 B  146  HIS SER GLU LYS LEU HIS VAL ASP PRO ASP ASN PHE LYS          
SEQRES   9 B  146  LEU LEU SER ASP CYS LEU THR ILE VAL VAL ALA ALA GLN          
SEQRES  10 B  146  LEU GLY LYS ALA PHE SER GLY GLU VAL GLN ALA ALA PHE          
SEQRES  11 B  146  GLN LYS PHE LEU SER VAL VAL VAL SER ALA LEU GLY LYS          
SEQRES  12 B  146  GLN TYR HIS                                                  
SEQRES   1 C  142  SER LEU SER SER LYS ASP LYS ASP THR VAL LYS ALA LEU          
SEQRES   2 C  142  TRP GLY LYS ILE ALA ASP LYS ALA GLU GLU ILE GLY SER          
SEQRES   3 C  142  ASP ALA LEU SER ARG MET LEU ALA VAL TYR PRO GLN THR          
SEQRES   4 C  142  LYS THR TYR PHE SER HIS TRP LYS ASP LEU SER PRO GLY          
SEQRES   5 C  142  SER ALA PRO VAL ASN LYS HIS GLY LYS THR ILE MET GLY          
SEQRES   6 C  142  GLY ILE VAL ASP ALA VAL ALA SER ILE ASP ASP LEU ASN          
SEQRES   7 C  142  ALA GLY LEU LEU ALA LEU SER GLU LEU HIS ALA PHE THR          
SEQRES   8 C  142  LEU ARG VAL ASP PRO ALA ASN PHE LYS ILE LEU SER HIS          
SEQRES   9 C  142  CYS ILE LEU VAL LEU LEU ALA VAL LYS PHE PRO LYS ASP          
SEQRES  10 C  142  PHE THR PRO GLU VAL HIS ILE SER TYR ASP LYS PHE PHE          
SEQRES  11 C  142  SER ALA LEU ALA ARG ALA LEU ALA GLU LYS TYR ARG              
SEQRES   1 D  146  VAL VAL TRP THR ASP PHE GLU ARG ALA THR ILE ALA ASP          
SEQRES   2 D  146  ILE PHE SER LYS LEU ASP TYR GLU ALA VAL GLY GLY ALA          
SEQRES   3 D  146  THR LEU ALA ARG CYS LEU ILE VAL TYR PRO TRP THR GLN          
SEQRES   4 D  146  ARG TYR PHE GLY ASN PHE GLY ASN LEU TYR ASN ALA ALA          
SEQRES   5 D  146  ALA ILE MET GLY ASN PRO MET ILE ALA LYS HIS GLY THR          
SEQRES   6 D  146  THR ILE LEU HIS GLY LEU ASP ARG ALA VAL LYS ASN MET          
SEQRES   7 D  146  ASP ASN ILE LYS ALA THR TYR ALA GLU LEU SER VAL LEU          
SEQRES   8 D  146  HIS SER GLU LYS LEU HIS VAL ASP PRO ASP ASN PHE LYS          
SEQRES   9 D  146  LEU LEU SER ASP CYS LEU THR ILE VAL VAL ALA ALA GLN          
SEQRES  10 D  146  LEU GLY LYS ALA PHE SER GLY GLU VAL GLN ALA ALA PHE          
SEQRES  11 D  146  GLN LYS PHE LEU SER VAL VAL VAL SER ALA LEU GLY LYS          
SEQRES  12 D  146  GLN TYR HIS                                                  
HET    ACE  A   0       3                                                       
HET    ACE  C   0       3                                                       
HET    HEM  A 143      43                                                       
HET    HEM  B 148      43                                                       
HET    HEM  C 143      43                                                       
HET    HEM  D 148      43                                                       
HETNAM     ACE ACETYL GROUP                                                     
HETNAM     HEM PROTOPORPHYRIN IX CONTAINING FE                                  
HETSYN     HEM HEME                                                             
FORMUL   5  ACE    2(C2 H4 O)                                                   
FORMUL   7  HEM    4(C34 H32 FE N4 O4)                                          
FORMUL  11  HOH   *219(H2 O)                                                    
HELIX    1   1 SER A    3  ILE A   17  1                                  15    
HELIX    2   2 LYS A   20  TYR A   36  1                                  17    
HELIX    3   3 PRO A   37  SER A   44  5                                   8    
HELIX    4   4 SER A   53  SER A   73  1                                  21    
HELIX    5   5 ASP A   76  LEU A   81  1                                   6    
HELIX    6   6 LEU A   81  THR A   91  1                                  11    
HELIX    7   7 PRO A   96  PHE A  114  1                                  19    
HELIX    8   8 PRO A  115  PHE A  118  5                                   4    
HELIX    9   9 THR A  119  GLU A  139  1                                  21    
HELIX   10  10 LYS A  140  ARG A  142  5                                   3    
HELIX   11  11 THR B    4  LEU B   18  1                                  15    
HELIX   12  12 ASP B   19  TYR B   35  1                                  17    
HELIX   13  13 PRO B   36  GLY B   43  5                                   8    
HELIX   14  14 ASN B   50  GLY B   56  1                                   7    
HELIX   15  15 ASN B   57  LYS B   76  1                                  20    
HELIX   16  16 ASN B   80  TYR B   85  1                                   6    
HELIX   17  17 TYR B   85  SER B   93  1                                   9    
HELIX   18  18 PRO B  100  GLY B  119  1                                  20    
HELIX   19  19 LYS B  120  PHE B  122  5                                   3    
HELIX   20  20 SER B  123  GLY B  142  1                                  20    
HELIX   21  21 SER C    3  ILE C   17  1                                  15    
HELIX   22  22 LYS C   20  TYR C   36  1                                  17    
HELIX   23  23 PRO C   37  SER C   44  5                                   8    
HELIX   24  24 SER C   53  SER C   73  1                                  21    
HELIX   25  25 ASP C   76  LEU C   81  1                                   6    
HELIX   26  26 LEU C   81  PHE C   90  1                                  10    
HELIX   27  27 ASP C   95  ALA C   97  5                                   3    
HELIX   28  28 ASN C   98  PHE C  114  1                                  17    
HELIX   29  29 PRO C  115  PHE C  118  5                                   4    
HELIX   30  30 THR C  119  GLU C  139  1                                  21    
HELIX   31  31 THR D    4  LEU D   18  1                                  15    
HELIX   32  32 ASP D   19  TYR D   35  1                                  17    
HELIX   33  33 PRO D   36  GLY D   43  5                                   8    
HELIX   34  34 ASN D   50  GLY D   56  1                                   7    
HELIX   35  35 ASN D   57  LYS D   76  1                                  20    
HELIX   36  36 ASN D   80  TYR D   85  1                                   6    
HELIX   37  37 TYR D   85  SER D   93  1                                   9    
HELIX   38  38 PRO D  100  GLY D  119  1                                  20    
HELIX   39  39 LYS D  120  PHE D  122  5                                   3    
HELIX   40  40 SER D  123  GLY D  142  1                                  20    
LINK        FE   HEM A 143                 NE2 HIS A  88                        
LINK        FE   HEM B 148                 NE2 HIS B  92                        
LINK        FE   HEM C 143                 NE2 HIS C  88                        
LINK        FE   HEM D 148                 NE2 HIS D  92                        
LINK         C   ACE A   0                 N   SER A   1                        
LINK         C   ACE C   0                 N   SER C   1                        
CRYST1   57.302   84.986  121.779  90.00  90.00  90.00 P 21 21 21    8          
ORIGX1      1.000000  0.000000  0.000000        0.00000                         
ORIGX2      0.000000  1.000000  0.000000        0.00000                         
ORIGX3      0.000000  0.000000  1.000000        0.00000                         
SCALE1      0.017451  0.000000  0.000000        0.00000                         
SCALE2      0.000000  0.011767  0.000000        0.00000                         
SCALE3      0.000000  0.000000  0.008212        0.00000