PDB entry 1v75

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HEADER    OXYGEN STORAGE/TRANSPORT                12-DEC-03   1V75              
TITLE     CRYSTAL STRUCTURE OF HEMOGLOBIN D FROM THE ALDABRA GIANT              
TITLE    2 TORTOISE (GEOCHELONE GIGANTEA) AT 2.0 A RESOLUTION                   
COMPND    MOL_ID: 1;                                                            
COMPND   2 MOLECULE: HEMOGLOBIN D ALPHA CHAIN;                                  
COMPND   3 CHAIN: A;                                                            
COMPND   4 SYNONYM: HEMOGLOBIN ALPHA-D CHAIN;                                   
COMPND   5 MOL_ID: 2;                                                           
COMPND   6 MOLECULE: HEMOGLOBIN A AND D BETA CHAIN;                             
COMPND   7 CHAIN: B;                                                            
COMPND   8 SYNONYM: HEMOGLOBIN BETA CHAIN                                       
SOURCE    MOL_ID: 1;                                                            
SOURCE   2 ORGANISM_SCIENTIFIC: ALDABRACHELYS ELEPHANTINA;                      
SOURCE   3 ORGANISM_COMMON: ALDABRA GIANT TORTOISE;                             
SOURCE   4 TISSUE: RED BLOOD CELL;                                              
SOURCE   5 MOL_ID: 2;                                                           
SOURCE   6 ORGANISM_SCIENTIFIC: ALDABRACHELYS ELEPHANTINA;                      
SOURCE   7 ORGANISM_COMMON: ALDABRA GIANT TORTOISE;                             
SOURCE   8 TISSUE: RED BLOOD CELL                                               
KEYWDS    HEMOGLOBIN D, REPTILIA, THE ALDABRA GIANT TORTOISE,                   
KEYWDS   2 GEOCHELONE GIGANTEA                                                  
EXPDTA    X-RAY DIFFRACTION                                                     
AUTHOR    T.KUWADA,T.HASEGAWA,I.SATOH,K.ISHIKAWA,F.SHISHIKURA                   
REVDAT   1   30-DEC-03 1V75    0                                                
JRNL        AUTH   T.KUWADA,T.HASEGAWA,I.SATOH,K.ISHIKAWA,F.SHISHIKURA          
JRNL        TITL   CRYSTALLIZATION AND PRELIMINARY X-RAY DIFFRACTION            
JRNL        TITL 2 STUDY OF HEMOGLOBIN D FROM THE ALDABRA GIANT                 
JRNL        TITL 3 TORTOISE, GEOCHELONE GIGANTEA.                               
JRNL        REF    PROTEIN PEPT.LETT.            V.  10   422 2003              
JRNL        REFN   ASTM PPELEN  NE ISSN 0929-8665                               
REMARK   1                                                                      
REMARK   2                                                                      
REMARK   2 RESOLUTION. 2.02 ANGSTROMS.                                          
REMARK   3                                                                      
REMARK   3 REFINEMENT.                                                          
REMARK   3   PROGRAM     : CNX 2000.1                                           
REMARK   3   AUTHORS     : BRUNGER,ADAMS,CLORE,DELANO,GROS,GROSSE-              
REMARK   3               : KUNSTLEVE,JIANG,KUSZEWSKI,NILGES,PANNU,              
REMARK   3               : READ,RICE,SIMONSON,WARREN,MOLECULAR                  
REMARK   3               : SIMULATIONS (BADGER,BERARD,KUMAR,SZALMA,             
REMARK   3               : YIP,DZAKULA)                                         
REMARK   3                                                                      
REMARK   3  DATA USED IN REFINEMENT.                                            
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.02                           
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 33.96                          
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.000                          
REMARK   3   DATA CUTOFF HIGH         (ABS(F)) : 411817.340                     
REMARK   3   DATA CUTOFF LOW          (ABS(F)) : 0.0000                         
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : 98.7                           
REMARK   3   NUMBER OF REFLECTIONS             : 22778                          
REMARK   3                                                                      
REMARK   3  FIT TO DATA USED IN REFINEMENT.                                     
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT                      
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM                          
REMARK   3   R VALUE     (WORKING + TEST SET) : NULL                            
REMARK   3   R VALUE            (WORKING SET) : 0.188                           
REMARK   3   FREE R VALUE                     : 0.233                           
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 9.900                           
REMARK   3   FREE R VALUE TEST SET COUNT      : 2260                            
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : 0.005                           
REMARK   3                                                                      
REMARK   3  FIT/AGREEMENT OF MODEL WITH ALL DATA.                               
REMARK   3   R VALUE     (WORKING + TEST SET, NO CUTOFF) : NULL                 
REMARK   3   R VALUE            (WORKING SET, NO CUTOFF) : NULL                 
REMARK   3   FREE R VALUE                    (NO CUTOFF) : NULL                 
REMARK   3   FREE R VALUE TEST SET SIZE   (%, NO CUTOFF) : NULL                 
REMARK   3   FREE R VALUE TEST SET COUNT     (NO CUTOFF) : NULL                 
REMARK   3   ESTIMATED ERROR OF FREE R VALUE (NO CUTOFF) : NULL                 
REMARK   3   TOTAL NUMBER OF REFLECTIONS     (NO CUTOFF) : NULL                 
REMARK   3                                                                      
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.                                  
REMARK   3   TOTAL NUMBER OF BINS USED           : 6                            
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 2.02                         
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 2.15                         
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 99.90                        
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : 3419                         
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2610                       
REMARK   3   BIN FREE R VALUE                    : 0.2630                       
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : 10.20                        
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : 388                          
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : 0.013                        
REMARK   3                                                                      
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.                    
REMARK   3   PROTEIN ATOMS            : 2212                                    
REMARK   3   NUCLEIC ACID ATOMS       : 0                                       
REMARK   3   HETEROGEN ATOMS          : 86                                      
REMARK   3   SOLVENT ATOMS            : 412                                     
REMARK   3                                                                      
REMARK   3  B VALUES.                                                           
REMARK   3   FROM WILSON PLOT           (A**2) : 22.70                          
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 35.20                          
REMARK   3   OVERALL ANISOTROPIC B VALUE.                                       
REMARK   3    B11 (A**2) : -6.85000                                             
REMARK   3    B22 (A**2) : 16.59000                                             
REMARK   3    B33 (A**2) : -9.74000                                             
REMARK   3    B12 (A**2) : 0.00000                                              
REMARK   3    B13 (A**2) : -0.71000                                             
REMARK   3    B23 (A**2) : 0.00000                                              
REMARK   3                                                                      
REMARK   3  ESTIMATED COORDINATE ERROR.                                         
REMARK   3   ESD FROM LUZZATI PLOT        (A) : 0.22                            
REMARK   3   ESD FROM SIGMAA              (A) : 0.23                            
REMARK   3   LOW RESOLUTION CUTOFF        (A) : 5.00                            
REMARK   3                                                                      
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.                         
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : 0.27                            
REMARK   3   ESD FROM C-V SIGMAA          (A) : 0.22                            
REMARK   3                                                                      
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.                                   
REMARK   3   BOND LENGTHS                 (A) : 0.005                           
REMARK   3   BOND ANGLES            (DEGREES) : 1.10                            
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : 16.00                           
REMARK   3   IMPROPER ANGLES        (DEGREES) : 0.71                            
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL MODEL : OVERALL                                   
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA                
REMARK   3   MAIN-CHAIN BOND              (A**2) : NULL  ; NULL                 
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : NULL  ; NULL                 
REMARK   3   SIDE-CHAIN BOND              (A**2) : NULL  ; NULL                 
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : NULL  ; NULL                 
REMARK   3                                                                      
REMARK   3  BULK SOLVENT MODELING.                                              
REMARK   3   METHOD USED : FLAT MODEL                                           
REMARK   3   KSOL        : 0.40                                                 
REMARK   3   BSOL        : 91.57                                                
REMARK   3                                                                      
REMARK   3  NCS MODEL : NULL                                                    
REMARK   3                                                                      
REMARK   3  PARAMETER FILE  1  : PROTEIN_REP.PARAM                              
REMARK   3  PARAMETER FILE  2  : ION.PARAM                                      
REMARK   3  PARAMETER FILE  3  : WATER_REP.PARAM                                
REMARK   3  PARAMETER FILE  4  : HEM2.PARAM                                     
REMARK   3  TOPOLOGY FILE  1   : PROTEIN.TOP                                    
REMARK   3  TOPOLOGY FILE  2   : ION.TOP                                        
REMARK   3  TOPOLOGY FILE  3   : WATER.TOP                                      
REMARK   3  TOPOLOGY FILE  4   : HEM2.TOP                                       
REMARK   3                                                                      
REMARK   3  OTHER REFINEMENT REMARKS: NULL                                      
REMARK   4                                                                      
REMARK   4 1V75 COMPLIES WITH FORMAT V. 3.0, 1-DEC-2006                         
REMARK   4                                                                      
REMARK   4 THIS IS THE REMEDIATED VERSION OF THIS PDB ENTRY.                    
REMARK   4 REMEDIATED DATA FILE REVISION 3.100 (2007-03-17)                     
REMARK 100                                                                      
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBJ .                              
REMARK 100 THE RCSB ID CODE IS RCSB006290.                                      
REMARK 200                                                                      
REMARK 200 EXPERIMENTAL DETAILS                                                 
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION                  
REMARK 200  DATE OF DATA COLLECTION        : 15-OCT-2002                        
REMARK 200  TEMPERATURE           (KELVIN) : 100.0                              
REMARK 200  PH                             : 7.50                               
REMARK 200  NUMBER OF CRYSTALS USED        : 1                                  
REMARK 200                                                                      
REMARK 200  SYNCHROTRON              (Y/N) : N                                  
REMARK 200  RADIATION SOURCE               : ROTATING ANODE                     
REMARK 200  BEAMLINE                       : NULL                               
REMARK 200  X-RAY GENERATOR MODEL          : RIGAKU ULTRAX 18                   
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M                                  
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.5418                             
REMARK 200  MONOCHROMATOR                  : CONFORCAL MAX-FLUX                 
REMARK 200  OPTICS                         : NULL                               
REMARK 200                                                                      
REMARK 200  DETECTOR TYPE                  : IMAGE PLATE                        
REMARK 200  DETECTOR MANUFACTURER          : RIGAKU RAXIS IV++                  
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : CRYSTALCLEAR                       
REMARK 200  DATA SCALING SOFTWARE          : CRYSTALCLEAR                       
REMARK 200                                                                      
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 22787                              
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.020                              
REMARK 200  RESOLUTION RANGE LOW       (A) : 53.650                             
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL                               
REMARK 200                                                                      
REMARK 200 OVERALL.                                                             
REMARK 200  COMPLETENESS FOR RANGE     (%) : 98.8                               
REMARK 200  DATA REDUNDANCY                : 2.780                              
REMARK 200  R MERGE                    (I) : 0.04000                            
REMARK 200  R SYM                      (I) : NULL                               
REMARK 200   FOR THE DATA SET  : 11.8000                            
REMARK 200                                                                      
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.                                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.02                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.09                     
REMARK 200  COMPLETENESS FOR SHELL     (%) : 100.0                              
REMARK 200  DATA REDUNDANCY IN SHELL       : 2.71                               
REMARK 200  R MERGE FOR SHELL          (I) : 0.13500                            
REMARK 200  R SYM FOR SHELL            (I) : NULL                               
REMARK 200   FOR SHELL         : 3.300                              
REMARK 200                                                                      
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH                              
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT        
REMARK 200 SOFTWARE USED: CNX 2000.1                                            
REMARK 200 STARTING MODEL: NULL                                                 
REMARK 200                                                                      
REMARK 200 REMARK: NULL                                                         
REMARK 280                                                                      
REMARK 280 CRYSTAL                                                              
REMARK 280 SOLVENT CONTENT, VS   (%): 46.89                                     
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.33                     
REMARK 280                                                                      
REMARK 280 CRYSTALLIZATION CONDITIONS: 10% PEG3350, 50MM HEPES-NA, PH 7.5,      
REMARK 280  VAPOR DIFFUSION, HANGING DROP, TEMPERATURE 293K                     
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY                                            
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: C 1 2 1                          
REMARK 290                                                                      
REMARK 290      SYMOP   SYMMETRY                                                
REMARK 290     NNNMMM   OPERATOR                                                
REMARK 290       1555   X,Y,Z                                                   
REMARK 290       2555   -X,Y,-Z                                                 
REMARK 290       3555   1/2+X,1/2+Y,Z                                           
REMARK 290       4555   1/2-X,1/2+Y,-Z                                          
REMARK 290                                                                      
REMARK 290     WHERE NNN -> OPERATOR NUMBER                                     
REMARK 290           MMM -> TRANSLATION VECTOR                                  
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS                            
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM             
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY                
REMARK 290 RELATED MOLECULES.                                                   
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   2  0.000000  1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   2  0.000000  0.000000 -1.000000        0.00000            
REMARK 290   SMTRY1   3  1.000000  0.000000  0.000000       56.07100            
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       31.18700            
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   4 -1.000000  0.000000  0.000000       56.07100            
REMARK 290   SMTRY2   4  0.000000  1.000000  0.000000       31.18700            
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000            
REMARK 290                                                                      
REMARK 290 REMARK: NULL                                                         
REMARK 300                                                                      
REMARK 300 BIOMOLECULE: 1                                                       
REMARK 300 THIS ENTRY CONTAINS THE CRYSTALLOGRAPHIC ASYMMETRIC UNIT             
REMARK 300 WHICH CONSISTS OF 2 CHAIN(S). SEE REMARK 350 FOR                     
REMARK 300 INFORMATION ON GENERATING THE BIOLOGICAL MOLECULE(S).                
REMARK 350                                                                      
REMARK 350 GENERATING THE BIOMOLECULE                                           
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN           
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE                
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS          
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND                          
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.                               
REMARK 350                                                                      
REMARK 350 BIOMOLECULE: 1                                                       
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B                                  
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 350   BIOMT1   2 -1.000000  0.000000  0.000000      112.14200            
REMARK 350   BIOMT2   2  0.000000  1.000000  0.000000        0.00000            
REMARK 350   BIOMT3   2  0.000000  0.000000 -1.000000        0.00000            
REMARK 375                                                                      
REMARK 375 SPECIAL POSITION                                                     
REMARK 375 THE FOLLOWING ATOMS ARE FOUND TO BE WITHIN 0.15 ANGSTROMS            
REMARK 375 OF A SYMMETRY RELATED ATOM AND ARE ASSUMED TO BE ON SPECIAL          
REMARK 375 POSITIONS.                                                           
REMARK 375                                                                      
REMARK 375 ATOM RES CSSEQI                                                      
REMARK 375      HOH   303   LIES ON A SPECIAL POSITION.                         
REMARK 375      HOH   372   LIES ON A SPECIAL POSITION.                         
REMARK 465                                                                      
REMARK 465 MISSING RESIDUES                                                     
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE                       
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)                
REMARK 465                                                                      
REMARK 465   M RES C SSSEQI                                                     
REMARK 465     VAL B     1                                                      
REMARK 465     GLY B   144                                                      
REMARK 465     TYR B   145                                                      
REMARK 465     HIS B   146                                                      
REMARK 470                                                                      
REMARK 470 MISSING ATOM                                                         
REMARK 470 THE FOLLOWING RESIDUES HAVE MISSING ATOMS(M=MODEL NUMBER;            
REMARK 470 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;          
REMARK 470 I=INSERTION CODE):                                                   
REMARK 470   M RES CSSEQI  ATOMS                                                
REMARK 470     HIS A  20    CG    ND1   CD2   CE1   NE2                         
REMARK 470     LYS A  71    CB    CG    CD    CE    NZ                          
REMARK 470     ALA A  78    CB                                                  
REMARK 470     ARG A 141    CG    CD    NE    CZ    NH1   NH2                   
REMARK 470     HIS B   2    CB    CG    ND1   CD2   CE1   NE2                   
REMARK 470     LYS B  83    CG    CD    CE    NZ                                
REMARK 470     GLN B  87    CG    CD    OE1   NE2                               
REMARK 470     GLU B  90    CG    CD    OE1   OE2                               
REMARK 470     GLU B  94    CG    CD    OE1   OE2                               
REMARK 470     LYS B 120    CG    CD    CE    NZ                                
REMARK 470     LEU B 143    CG    CD1   CD2                                     
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT                     
REMARK 500                                                                      
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.                            
REMARK 500                                                                      
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI                             
REMARK 500  FE    HEM B   201     O    HOH     412              2.01            
REMARK 500  FE    HEM A   201     O    HOH     411              2.18            
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: CLOSE CONTACTS                                             
REMARK 500                                                                      
REMARK 500 THE FOLLOWING ATOMS THAT ARE RELATED BY CRYSTALLOGRAPHIC             
REMARK 500 SYMMETRY ARE IN CLOSE CONTACT.  AN ATOM LOCATED WITHIN 0.15          
REMARK 500 ANGSTROMS OF A SYMMETRY RELATED ATOM IS ASSUMED TO BE ON A           
REMARK 500 SPECIAL POSITION AND IS, THEREFORE, LISTED IN REMARK 375             
REMARK 500 INSTEAD OF REMARK 500.  ATOMS WITH NON-BLANK ALTERNATE               
REMARK 500 LOCATION INDICATORS ARE NOT INCLUDED IN THE CALCULATIONS.            
REMARK 500                                                                      
REMARK 500 DISTANCE CUTOFF:                                                     
REMARK 500 2.2 ANGSTROMS FOR CONTACTS NOT INVOLVING HYDROGEN ATOMS              
REMARK 500 1.6 ANGSTROMS FOR CONTACTS INVOLVING HYDROGEN ATOMS                  
REMARK 500                                                                      
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI  SSYMOP   DISTANCE          
REMARK 500   O    HOH     281     O    HOH     281     2655     1.91            
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND LENGTHS                                      
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,2(A3,1X,A1,I4,A1,1X,A4,3X),F6.3)                  
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   RES CSSEQI ATM2   DEVIATION                     
REMARK 500    THR B  38   CA    THR B  38   CB     0.038                        
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES                                       
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)              
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3                                     
REMARK 500    ASP A  47   N   -  CA  -  C   ANGL. DEV. =-14.9 DEGREES           
REMARK 500    ASP A  51   N   -  CA  -  C   ANGL. DEV. =  7.9 DEGREES           
REMARK 500    ASN B  19   N   -  CA  -  C   ANGL. DEV. =-10.3 DEGREES           
REMARK 500    PRO B  36   N   -  CA  -  C   ANGL. DEV. =  8.0 DEGREES           
REMARK 500    SER B  49   N   -  CA  -  C   ANGL. DEV. =  7.3 DEGREES           
REMARK 500    LEU B  78   N   -  CA  -  C   ANGL. DEV. =  7.2 DEGREES           
REMARK 525                                                                      
REMARK 525 SOLVENT                                                              
REMARK 525 THE FOLLOWING SOLVENT MOLECULES LIE FARTHER THAN EXPECTED            
REMARK 525 FROM THE PROTEIN OR NUCLEIC ACID MOLECULE AND MAY BE                 
REMARK 525 ASSOCIATED WITH A SYMMETRY RELATED MOLECULE (M=MODEL                 
REMARK 525 NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE          
REMARK 525 NUMBER; I=INSERTION CODE):                                           
REMARK 525                                                                      
REMARK 525  M RES CSSEQI                                                        
REMARK 525    HOH     3        DISTANCE =  6.49 ANGSTROMS                       
REMARK 525    HOH   175        DISTANCE =  5.21 ANGSTROMS                       
REMARK 525    HOH   178        DISTANCE =  6.07 ANGSTROMS                       
REMARK 525    HOH   220        DISTANCE =  6.08 ANGSTROMS                       
REMARK 525    HOH   221        DISTANCE =  6.28 ANGSTROMS                       
REMARK 525    HOH   247        DISTANCE =  6.39 ANGSTROMS                       
REMARK 525    HOH   249        DISTANCE =  8.43 ANGSTROMS                       
REMARK 525    HOH   250        DISTANCE =  8.45 ANGSTROMS                       
REMARK 525    HOH   251        DISTANCE =  9.12 ANGSTROMS                       
REMARK 525    HOH   261        DISTANCE =  6.69 ANGSTROMS                       
REMARK 525    HOH   264        DISTANCE = 12.67 ANGSTROMS                       
REMARK 525    HOH   270        DISTANCE =  9.79 ANGSTROMS                       
REMARK 525    HOH   271        DISTANCE =  5.16 ANGSTROMS                       
REMARK 525    HOH   272        DISTANCE =  7.66 ANGSTROMS                       
REMARK 525    HOH   274        DISTANCE =  6.35 ANGSTROMS                       
REMARK 525    HOH   277        DISTANCE =  5.26 ANGSTROMS                       
REMARK 525    HOH   279        DISTANCE =  5.82 ANGSTROMS                       
REMARK 525    HOH   283        DISTANCE =  9.60 ANGSTROMS                       
REMARK 525    HOH   286        DISTANCE = 12.93 ANGSTROMS                       
REMARK 525    HOH   287        DISTANCE =  5.87 ANGSTROMS                       
REMARK 525    HOH   288        DISTANCE =  7.50 ANGSTROMS                       
REMARK 525    HOH   296        DISTANCE = 11.07 ANGSTROMS                       
REMARK 525    HOH   297        DISTANCE =  6.94 ANGSTROMS                       
REMARK 525    HOH   298        DISTANCE = 13.23 ANGSTROMS                       
REMARK 525    HOH   299        DISTANCE = 10.01 ANGSTROMS                       
REMARK 525    HOH   300        DISTANCE = 16.38 ANGSTROMS                       
REMARK 525    HOH   301        DISTANCE = 14.52 ANGSTROMS                       
REMARK 525    HOH   304        DISTANCE =  5.51 ANGSTROMS                       
REMARK 525    HOH   306        DISTANCE = 12.51 ANGSTROMS                       
REMARK 525    HOH   309        DISTANCE =  5.67 ANGSTROMS                       
REMARK 525    HOH   310        DISTANCE =  9.22 ANGSTROMS                       
REMARK 525    HOH   313        DISTANCE = 15.08 ANGSTROMS                       
REMARK 525    HOH   315        DISTANCE =  6.99 ANGSTROMS                       
REMARK 525    HOH   316        DISTANCE =  6.79 ANGSTROMS                       
REMARK 525    HOH   318        DISTANCE =  6.76 ANGSTROMS                       
REMARK 525    HOH   319        DISTANCE =  8.89 ANGSTROMS                       
REMARK 525    HOH   320        DISTANCE =  9.69 ANGSTROMS                       
REMARK 525    HOH   323        DISTANCE = 12.89 ANGSTROMS                       
REMARK 525    HOH   325        DISTANCE = 15.00 ANGSTROMS                       
REMARK 525    HOH   326        DISTANCE = 11.87 ANGSTROMS                       
REMARK 525    HOH   331        DISTANCE = 15.06 ANGSTROMS                       
REMARK 525    HOH   332        DISTANCE =  7.89 ANGSTROMS                       
REMARK 525    HOH   334        DISTANCE = 22.62 ANGSTROMS                       
REMARK 525    HOH   335        DISTANCE = 11.01 ANGSTROMS                       
REMARK 525    HOH   342        DISTANCE =  8.71 ANGSTROMS                       
REMARK 525    HOH   344        DISTANCE = 10.85 ANGSTROMS                       
REMARK 525    HOH   347        DISTANCE =  9.93 ANGSTROMS                       
REMARK 525    HOH   351        DISTANCE = 20.20 ANGSTROMS                       
REMARK 525    HOH   354        DISTANCE = 10.11 ANGSTROMS                       
REMARK 525    HOH   357        DISTANCE =  8.68 ANGSTROMS                       
REMARK 525    HOH   358        DISTANCE =  7.41 ANGSTROMS                       
REMARK 525    HOH   360        DISTANCE =  7.18 ANGSTROMS                       
REMARK 525    HOH   369        DISTANCE =  6.26 ANGSTROMS                       
REMARK 525    HOH   372        DISTANCE =  6.59 ANGSTROMS                       
REMARK 525    HOH   374        DISTANCE =  8.10 ANGSTROMS                       
REMARK 525    HOH   375        DISTANCE = 10.80 ANGSTROMS                       
REMARK 525    HOH   376        DISTANCE = 18.79 ANGSTROMS                       
REMARK 525    HOH   378        DISTANCE = 21.96 ANGSTROMS                       
REMARK 525    HOH   379        DISTANCE =  8.98 ANGSTROMS                       
REMARK 525    HOH   381        DISTANCE =  8.03 ANGSTROMS                       
REMARK 525    HOH   386        DISTANCE =  7.58 ANGSTROMS                       
REMARK 525    HOH   387        DISTANCE =  7.22 ANGSTROMS                       
REMARK 525    HOH   392        DISTANCE =  5.13 ANGSTROMS                       
REMARK 525    HOH   396        DISTANCE =  7.97 ANGSTROMS                       
REMARK 525    HOH   397        DISTANCE = 21.41 ANGSTROMS                       
REMARK 525    HOH   399        DISTANCE = 18.77 ANGSTROMS                       
REMARK 525    HOH   401        DISTANCE =  7.09 ANGSTROMS                       
REMARK 525    HOH   404        DISTANCE =  5.10 ANGSTROMS                       
REMARK 525    HOH   405        DISTANCE =  7.82 ANGSTROMS                       
REMARK 525    HOH   409        DISTANCE =  9.02 ANGSTROMS                       
DBREF  1V75 A    1   141  UNP    P83134   HBAD_ALDEL       1    141             
DBREF  1V75 B    1   146  UNP    P83133   HBB_ALDEL        1    146             
SEQRES   1 A  141  MET LEU THR GLU ASP ASP LYS GLN LEU ILE GLN HIS VAL          
SEQRES   2 A  141  TRP GLU LYS VAL LEU GLU HIS GLN GLU ASP PHE GLY ALA          
SEQRES   3 A  141  GLU ALA LEU GLU ARG MET PHE ILE VAL TYR PRO SER THR          
SEQRES   4 A  141  LYS THR TYR PHE PRO HIS PHE ASP LEU HIS HIS ASP SER          
SEQRES   5 A  141  GLU GLN ILE ARG HIS HIS GLY LYS LYS VAL VAL GLY ALA          
SEQRES   6 A  141  LEU GLY ASP ALA VAL LYS HIS ILE ASP ASN LEU SER ALA          
SEQRES   7 A  141  THR LEU SER GLU LEU SER ASN LEU HIS ALA TYR ASN LEU          
SEQRES   8 A  141  ARG VAL ASP PRO VAL ASN PHE LYS LEU LEU SER HIS CYS          
SEQRES   9 A  141  PHE GLN VAL VAL LEU GLY ALA HIS LEU GLY ARG GLU TYR          
SEQRES  10 A  141  THR PRO GLN VAL GLN VAL ALA TYR ASP LYS PHE LEU ALA          
SEQRES  11 A  141  ALA VAL SER ALA VAL LEU ALA GLU LYS TYR ARG                  
SEQRES   1 B  146  VAL HIS TRP THR SER GLU GLU LYS GLN TYR ILE THR SER          
SEQRES   2 B  146  LEU TRP ALA LYS VAL ASN VAL GLY GLU VAL GLY GLY GLU          
SEQRES   3 B  146  ALA LEU ALA ARG LEU LEU ILE VAL TYR PRO TRP THR GLN          
SEQRES   4 B  146  ARG PHE PHE ALA SER PHE GLY ASN LEU SER SER ALA ASN          
SEQRES   5 B  146  ALA ILE LEU HIS ASN ALA LYS VAL LEU ALA HIS GLY GLN          
SEQRES   6 B  146  LYS VAL LEU THR SER PHE GLY GLU ALA VAL LYS ASN LEU          
SEQRES   7 B  146  ASP ASN ILE LYS LYS THR PHE ALA GLN LEU SER GLU LEU          
SEQRES   8 B  146  HIS CYS GLU LYS LEU HIS VAL ASP PRO GLU ASN PHE LYS          
SEQRES   9 B  146  LEU LEU GLY ASN ILE LEU ILE ILE VAL LEU ALA THR HIS          
SEQRES  10 B  146  PHE PRO LYS GLU PHE THR PRO ALA SER GLN ALA ALA TRP          
SEQRES  11 B  146  THR LYS LEU VAL ASN ALA VAL ALA HIS ALA LEU ALA LEU          
SEQRES  12 B  146  GLY TYR HIS                                                  
HET    HEM  A 201      43                                                       
HET    HEM  B 201      43                                                       
HETNAM     HEM PROTOPORPHYRIN IX CONTAINING FE                                  
HETSYN     HEM HEME                                                             
FORMUL   3  HEM    2(C34 H32 FE N4 O4)                                          
FORMUL   5  HOH   *412(H2 O)                                                    
HELIX    1   1 THR A    3  LEU A   18  1                                  16    
HELIX    2   2 HIS A   20  TYR A   36  1                                  17    
HELIX    3   3 PRO A   37  PHE A   43  5                                   7    
HELIX    4   4 SER A   52  HIS A   72  1                                  21    
HELIX    5   5 ASN A   75  ASN A   90  1                                  16    
HELIX    6   6 ASP A   94  GLY A  114  1                                  21    
HELIX    7   7 ARG A  115  TYR A  117  5                                   3    
HELIX    8   8 THR A  118  GLU A  138  1                                  21    
HELIX    9   9 LYS A  139  ARG A  141  5                                   3    
HELIX   10  10 THR B    4  ALA B   16  1                                  13    
HELIX   11  11 ASN B   19  TYR B   35  1                                  17    
HELIX   12  12 PRO B   36  GLY B   46  5                                  11    
HELIX   13  13 SER B   50  HIS B   56  1                                   7    
HELIX   14  14 ASN B   57  LYS B   76  1                                  20    
HELIX   15  15 ASN B   80  GLU B   94  1                                  15    
HELIX   16  16 PRO B  100  PHE B  118  1                                  19    
HELIX   17  17 THR B  123  LEU B  143  1                                  21    
LINK        FE   HEM A 201                 NE2 HIS A  87                        
LINK        FE   HEM B 201                 NE2 HIS B  92                        
CRYST1  112.142   62.374   53.980  90.00 110.26  90.00 C 1 2 1       4          
ORIGX1      1.000000  0.000000  0.000000        0.00000                         
ORIGX2      0.000000  1.000000  0.000000        0.00000                         
ORIGX3      0.000000  0.000000  1.000000        0.00000                         
SCALE1      0.008917  0.000000  0.003292        0.00000                         
SCALE2      0.000000  0.016032  0.000000        0.00000                         
SCALE3      0.000000  0.000000  0.019747        0.00000