PDB entry 1um5

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HEADER    IMMUNE SYSTEM                           23-SEP-03   1UM5              
TITLE     CATALYTIC ANTIBODY 21H3 WITH ALCOHOL SUBSTRATE                        
COMPND    MOL_ID: 1;                                                            
COMPND   2 MOLECULE: ANTIBODY 21H3 L CHAIN;                                     
COMPND   3 CHAIN: L;                                                            
COMPND   4 ENGINEERED: YES;                                                     
COMPND   5 MOL_ID: 2;                                                           
COMPND   6 MOLECULE: ANTIBODY 21H3 H CHAIN;                                     
COMPND   7 CHAIN: H;                                                            
COMPND   8 ENGINEERED: YES                                                      
SOURCE    MOL_ID: 1;                                                            
SOURCE   2 ORGANISM_SCIENTIFIC: MUS MUSCULUS;                                   
SOURCE   3 ORGANISM_COMMON: MOUSE;                                              
SOURCE   4 EXPRESSION_SYSTEM: ESCHERICHIA COLI;                                 
SOURCE   5 EXPRESSION_SYSTEM_COMMON: BACTERIA;                                  
SOURCE   6 MOL_ID: 2;                                                           
SOURCE   7 ORGANISM_SCIENTIFIC: MUS MUSCULUS;                                   
SOURCE   8 ORGANISM_COMMON: MOUSE;                                              
SOURCE   9 EXPRESSION_SYSTEM: ESCHERICHIA COLI;                                 
SOURCE  10 EXPRESSION_SYSTEM_COMMON: BACTERIA                                   
KEYWDS    CATALYTIC ANTIBODY LIGAND TRANSESTERIFICATION                         
EXPDTA    X-RAY DIFFRACTION                                                     
AUTHOR    A.E.BEUSCHER IV,J.REUTER,A.J.OLSON,F.E.ROMESBERG,                     
AUTHOR   2 P.G.SCHULTZ,P.WIRSCHING,K.D.JANDA,R.A.LERNER,R.C.STEVENS             
REVDAT   1   05-OCT-04 1UM5    0                                                
JRNL        AUTH   A.E.BEUSCHER IV,J.REUTER,A.J.OLSON,F.E.ROMESBERG,            
JRNL        AUTH 2 P.G.SCHULTZ,P.WIRSCHING,K.D.JANDA,R.A.LERNER,                
JRNL        AUTH 3 R.C.STEVENS                                                  
JRNL        TITL   STRUCTURAL STUDIES OF AN EFFICIENT CATALYTIC                 
JRNL        TITL 2 ANTIBODY OPERATING BY PING-PONG AND INDUCED FIT              
JRNL        TITL 3 MECHANISMS                                                   
JRNL        REF    TO BE PUBLISHED                                              
JRNL        REFN                                                                
REMARK   1                                                                      
REMARK   2                                                                      
REMARK   2 RESOLUTION. 1.60 ANGSTROMS.                                          
REMARK   3                                                                      
REMARK   3 REFINEMENT.                                                          
REMARK   3   PROGRAM     : CNS 1.1                                              
REMARK   3   AUTHORS     : BRUNGER,ADAMS,CLORE,DELANO,GROS,GROSSE-              
REMARK   3               : KUNSTLEVE,JIANG,KUSZEWSKI,NILGES, PANNU,             
REMARK   3               : READ,RICE,SIMONSON,WARREN                            
REMARK   3                                                                      
REMARK   3  REFINEMENT TARGET : ENGH & HUBER                                    
REMARK   3                                                                      
REMARK   3  DATA USED IN REFINEMENT.                                            
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.60                           
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 19.96                          
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 2.000                          
REMARK   3   DATA CUTOFF HIGH         (ABS(F)) : NULL                           
REMARK   3   DATA CUTOFF LOW          (ABS(F)) : NULL                           
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : 98.2                           
REMARK   3   NUMBER OF REFLECTIONS             : 55779                          
REMARK   3                                                                      
REMARK   3  FIT TO DATA USED IN REFINEMENT.                                     
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT                      
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM                          
REMARK   3   R VALUE            (WORKING SET) : 0.205                           
REMARK   3   FREE R VALUE                     : 0.246                           
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 3.400                           
REMARK   3   FREE R VALUE TEST SET COUNT      : 1919                            
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : 0.006                           
REMARK   3                                                                      
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.                                  
REMARK   3   TOTAL NUMBER OF BINS USED           : 8                            
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 1.60                         
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 1.67                         
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 95.00                        
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : 6434                         
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2190                       
REMARK   3   BIN FREE R VALUE                    : 0.2710                       
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : 3.20                         
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : 216                          
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : 0.018                        
REMARK   3                                                                      
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.                    
REMARK   3   PROTEIN ATOMS            : 3275                                    
REMARK   3   NUCLEIC ACID ATOMS       : 0                                       
REMARK   3   HETEROGEN ATOMS          : 9                                       
REMARK   3   SOLVENT ATOMS            : 398                                     
REMARK   3                                                                      
REMARK   3  B VALUES.                                                           
REMARK   3   FROM WILSON PLOT           (A**2) : 15.50                          
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 17.75                          
REMARK   3   OVERALL ANISOTROPIC B VALUE.                                       
REMARK   3    B11 (A**2) : 0.39000                                              
REMARK   3    B22 (A**2) : 2.71000                                              
REMARK   3    B33 (A**2) : -3.10000                                             
REMARK   3    B12 (A**2) : 0.00000                                              
REMARK   3    B13 (A**2) : 0.00000                                              
REMARK   3    B23 (A**2) : 0.00000                                              
REMARK   3                                                                      
REMARK   3  ESTIMATED COORDINATE ERROR.                                         
REMARK   3   ESD FROM LUZZATI PLOT        (A) : 0.17                            
REMARK   3   ESD FROM SIGMAA              (A) : 0.04                            
REMARK   3   LOW RESOLUTION CUTOFF        (A) : 5.00                            
REMARK   3                                                                      
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.                         
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : 0.21                            
REMARK   3   ESD FROM C-V SIGMAA          (A) : 0.07                            
REMARK   3                                                                      
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.                                   
REMARK   3   BOND LENGTHS                 (A) : 0.023                           
REMARK   3   BOND ANGLES            (DEGREES) : 2.20                            
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : NULL                            
REMARK   3   IMPROPER ANGLES        (DEGREES) : NULL                            
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL MODEL : NULL                                      
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA                
REMARK   3   MAIN-CHAIN BOND              (A**2) : NULL  ; NULL                 
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : NULL  ; NULL                 
REMARK   3   SIDE-CHAIN BOND              (A**2) : NULL  ; NULL                 
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : NULL  ; NULL                 
REMARK   3                                                                      
REMARK   3  BULK SOLVENT MODELING.                                              
REMARK   3   METHOD USED : CNS BULK SOLVENT MODEL USED                          
REMARK   3   KSOL        : 0.34                                                 
REMARK   3   BSOL        : 38.49                                                
REMARK   3                                                                      
REMARK   3  NCS MODEL : NULL                                                    
REMARK   3                                                                      
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT          
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL                 
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL                 
REMARK   3                                                                      
REMARK   3  PARAMETER FILE  1  : PROTEIN_REP.PARAM                              
REMARK   3  PARAMETER FILE  2  : CARBOHYDRATE.PARAM                             
REMARK   3  PARAMETER FILE  3  : WATER_REP.PARAM                                
REMARK   3  PARAMETER FILE  4  : SSU.PARAM                                      
REMARK   3  PARAMETER FILE  5  : NULL                                           
REMARK   3  TOPOLOGY FILE  1   : PROTEIN.TOP                                    
REMARK   3  TOPOLOGY FILE  2   : CARBOHYDRATE.TOP                               
REMARK   3  TOPOLOGY FILE  3   : WATER.TOP                                      
REMARK   3  TOPOLOGY FILE  4   : SSU.TOP                                        
REMARK   3  TOPOLOGY FILE  5   : NULL                                           
REMARK   3                                                                      
REMARK   3  OTHER REFINEMENT REMARKS: NULL                                      
REMARK   4                                                                      
REMARK   4 1UM5 COMPLIES WITH FORMAT V. 3.0, 1-DEC-2006                         
REMARK   4                                                                      
REMARK   4 THIS IS THE REMEDIATED VERSION OF THIS PDB ENTRY.                    
REMARK   4 REMEDIATED DATA FILE REVISION 3.100 (2007-03-17)                     
REMARK 100                                                                      
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBJ.                               
REMARK 100 THE RCSB ID CODE IS RCSB005979.                                      
REMARK 200                                                                      
REMARK 200 EXPERIMENTAL DETAILS                                                 
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION                  
REMARK 200  DATE OF DATA COLLECTION        : 01-JAN-1999                        
REMARK 200  TEMPERATURE           (KELVIN) : 100.0                              
REMARK 200  PH                             : 4.25                               
REMARK 200  NUMBER OF CRYSTALS USED        : 1                                  
REMARK 200                                                                      
REMARK 200  SYNCHROTRON              (Y/N) : Y                                  
REMARK 200  RADIATION SOURCE               : SSRL                               
REMARK 200  BEAMLINE                       : BL9-2                              
REMARK 200  X-RAY GENERATOR MODEL          : NULL                               
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M                                  
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.0                                
REMARK 200  MONOCHROMATOR                  : NULL                               
REMARK 200  OPTICS                         : NULL                               
REMARK 200                                                                      
REMARK 200  DETECTOR TYPE                  : CCD                                
REMARK 200  DETECTOR MANUFACTURER          : ADSC QUANTUM 4                     
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : DENZO                              
REMARK 200  DATA SCALING SOFTWARE          : SCALEPACK                          
REMARK 200                                                                      
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 55779                              
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.600                              
REMARK 200  RESOLUTION RANGE LOW       (A) : 47.280                             
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 0.000                              
REMARK 200                                                                      
REMARK 200 OVERALL.                                                             
REMARK 200  COMPLETENESS FOR RANGE     (%) : 98.3                               
REMARK 200  DATA REDUNDANCY                : NULL                               
REMARK 200  R MERGE                    (I) : NULL                               
REMARK 200  R SYM                      (I) : NULL                               
REMARK 200   FOR THE DATA SET  : NULL                               
REMARK 200                                                                      
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.                                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.60                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.66                     
REMARK 200  COMPLETENESS FOR SHELL     (%) : 99.3                               
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL                               
REMARK 200  R MERGE FOR SHELL          (I) : NULL                               
REMARK 200  R SYM FOR SHELL            (I) : NULL                               
REMARK 200   FOR SHELL         : NULL                               
REMARK 200                                                                      
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH                              
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT        
REMARK 200 SOFTWARE USED: EPMR                                                  
REMARK 200 STARTING MODEL: NULL                                                 
REMARK 200                                                                      
REMARK 200 REMARK: NULL                                                         
REMARK 280                                                                      
REMARK 280 CRYSTAL                                                              
REMARK 280 SOLVENT CONTENT, VS   (%): 39.48                                     
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.06                     
REMARK 280                                                                      
REMARK 280 CRYSTALLIZATION CONDITIONS: PEG4000, LITHIUM SULFATE, PH 4.25,       
REMARK 280  VAPOR DIFFUSION, HANGING DROP, TEMPERATURE 277K                     
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY                                            
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 21                       
REMARK 290                                                                      
REMARK 290      SYMOP   SYMMETRY                                                
REMARK 290     NNNMMM   OPERATOR                                                
REMARK 290       1555   X,Y,Z                                                   
REMARK 290       2555   1/2-X,-Y,1/2+Z                                          
REMARK 290       3555   -X,1/2+Y,1/2-Z                                          
REMARK 290       4555   1/2+X,1/2-Y,-Z                                          
REMARK 290                                                                      
REMARK 290     WHERE NNN -> OPERATOR NUMBER                                     
REMARK 290           MMM -> TRANSLATION VECTOR                                  
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS                            
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM             
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY                
REMARK 290 RELATED MOLECULES.                                                   
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000       23.56400            
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       69.25500            
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       32.35300            
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000       69.25500            
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       23.56400            
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       32.35300            
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000            
REMARK 290                                                                      
REMARK 290 REMARK: NULL                                                         
REMARK 300                                                                      
REMARK 300 BIOMOLECULE: 1                                                       
REMARK 300 THIS ENTRY CONTAINS THE CRYSTALLOGRAPHIC ASYMMETRIC UNIT             
REMARK 300 WHICH CONSISTS OF 2 CHAIN(S). SEE REMARK 350 FOR                     
REMARK 300 INFORMATION ON GENERATING THE BIOLOGICAL MOLECULE(S).                
REMARK 350                                                                      
REMARK 350 GENERATING THE BIOMOLECULE                                           
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN           
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE                
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS          
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND                          
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.                               
REMARK 350                                                                      
REMARK 350 BIOMOLECULE: 1                                                       
REMARK 350 APPLY THE FOLLOWING TO CHAINS: L, H                                  
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 465                                                                      
REMARK 465 MISSING RESIDUES                                                     
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE                       
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)                
REMARK 465                                                                      
REMARK 465   M RES C SSSEQI                                                     
REMARK 465     ALA L   218                                                      
REMARK 465     GLY L   219                                                      
REMARK 465     ALA L   220                                                      
REMARK 470                                                                      
REMARK 470 MISSING ATOM                                                         
REMARK 470 THE FOLLOWING RESIDUES HAVE MISSING ATOMS(M=MODEL NUMBER;            
REMARK 470 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;          
REMARK 470 I=INSERTION CODE):                                                   
REMARK 470   M RES CSSEQI  ATOMS                                                
REMARK 470     LEU L   2    CG    CD1   CD2                                     
REMARK 470     CYS L 216    CB    SG                                            
REMARK 470     VAL H   3    CG1   CG2                                           
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: CLOSE CONTACTS                                             
REMARK 500                                                                      
REMARK 500 THE FOLLOWING ATOMS THAT ARE RELATED BY CRYSTALLOGRAPHIC             
REMARK 500 SYMMETRY ARE IN CLOSE CONTACT.  AN ATOM LOCATED WITHIN 0.15          
REMARK 500 ANGSTROMS OF A SYMMETRY RELATED ATOM IS ASSUMED TO BE ON A           
REMARK 500 SPECIAL POSITION AND IS, THEREFORE, LISTED IN REMARK 375             
REMARK 500 INSTEAD OF REMARK 500.  ATOMS WITH NON-BLANK ALTERNATE               
REMARK 500 LOCATION INDICATORS ARE NOT INCLUDED IN THE CALCULATIONS.            
REMARK 500                                                                      
REMARK 500 DISTANCE CUTOFF:                                                     
REMARK 500 2.2 ANGSTROMS FOR CONTACTS NOT INVOLVING HYDROGEN ATOMS              
REMARK 500 1.6 ANGSTROMS FOR CONTACTS INVOLVING HYDROGEN ATOMS                  
REMARK 500                                                                      
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI  SSYMOP   DISTANCE          
REMARK 500   O    HOH     303     O    HOH     304     4466     2.11            
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND LENGTHS                                      
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,2(A3,1X,A1,I4,A1,1X,A4,3X),F6.3)                  
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   RES CSSEQI ATM2   DEVIATION                     
REMARK 500    ILE L 108   CG1   ILE L 108   CD1   -0.137                        
REMARK 500    ILE H  52   CG1   ILE H  52   CD1   -0.219                        
REMARK 500    LEU H 114   CG    LEU H 114   CD2   -0.162                        
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES                                       
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)              
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3                                     
REMARK 500    LEU L  35   CA  -  CB  -  CG  ANGL. DEV. =-16.2 DEGREES           
REMARK 500    GLY L 217   N   -  CA  -  C   ANGL. DEV. =-14.2 DEGREES           
REMARK 500    LYS H  68   CD  -  CE  -  NZ  ANGL. DEV. = 13.7 DEGREES           
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: TORSION ANGLES                                             
REMARK 500                                                                      
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:            
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;               
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                             
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)                    
REMARK 500                                                                      
REMARK 500  M RES CSSEQI        PSI       PHI                                   
REMARK 500    CYS L 216     -128.48     90.28                                   
REMARK 900                                                                      
REMARK 900 RELATED ENTRIES                                                      
REMARK 900 RELATED ID: 1UM4   RELATED DB: PDB                                   
REMARK 900 THE SAME PROTEIN COMPLEXED WITH SH4                                  
REMARK 900 RELATED ID: 1UM6   RELATED DB: PDB                                   
REMARK 900 THE SAME PROTEIN                                                     
SEQRES   1 L  219  LEU ASP ILE GLN MET THR GLN SER PRO SER SER LEU SER          
SEQRES   2 L  219  ALA SER LEU GLY GLU ARG VAL SER LEU THR CYS ARG ALA          
SEQRES   3 L  219  SER GLN GLU ILE SER GLY TYR LEU TYR TRP LEU GLN GLN          
SEQRES   4 L  219  LYS PRO ASP GLY THR ILE LYS ARG LEU ILE TYR ALA GLY          
SEQRES   5 L  219  SER THR LEU ASP SER GLY VAL PRO LYS ARG PHE SER GLY          
SEQRES   6 L  219  SER ARG SER GLY SER ASP TYR SER LEU THR ILE SER SER          
SEQRES   7 L  219  LEU GLU SER GLU ASP PHE ALA ASP TYR TYR CYS LEU GLN          
SEQRES   8 L  219  TYR ALA SER TYR PRO ARG THR PHE GLY GLY GLY THR LYS          
SEQRES   9 L  219  VAL GLU ILE LYS ARG THR VAL ALA ALA PRO SER VAL PHE          
SEQRES  10 L  219  ILE PHE PRO PRO SER ASP GLU GLN LEU LYS SER GLY THR          
SEQRES  11 L  219  ALA SER VAL VAL CYS LEU LEU ASN ASN PHE TYR PRO ARG          
SEQRES  12 L  219  GLU ALA LYS VAL GLN TRP LYS VAL ASP ASN ALA LEU GLN          
SEQRES  13 L  219  SER GLY ASN SER GLN GLU SER VAL THR GLU GLN ASP SER          
SEQRES  14 L  219  LYS ASP SER THR TYR SER LEU SER SER THR LEU THR LEU          
SEQRES  15 L  219  SER LYS ALA ASP TYR GLU LYS HIS LYS VAL TYR ALA CYS          
SEQRES  16 L  219  GLU VAL THR HIS GLN GLY LEU SER SER PRO VAL THR LYS          
SEQRES  17 L  219  SER PHE ASN ARG GLY GLU CYS GLY ALA GLY ALA                  
SEQRES   1 H  217  VAL GLN LEU GLN GLN SER GLY PRO VAL LEU VAL LYS PRO          
SEQRES   2 H  217  GLY GLY SER VAL LYS MET SER CYS LYS ALA SER GLU TYR          
SEQRES   3 H  217  THR LEU THR SER TYR LEU PHE GLN TRP VAL LYS GLN LYS          
SEQRES   4 H  217  SER GLY GLN GLY LEU GLU TRP ILE GLY TYR ILE TYR PRO          
SEQRES   5 H  217  TYR ASN GLY GLY THR ARG TYR ASN GLU LYS PHE ARG GLY          
SEQRES   6 H  217  LYS ALA THR LEU THR SER ASP LYS SER SER ASN THR ALA          
SEQRES   7 H  217  TYR LEU GLU LEU SER SER LEU THR SER GLU ASP SER ALA          
SEQRES   8 H  217  VAL TYR TYR CYS ALA ARG SER SER MET SER ASP PRO GLY          
SEQRES   9 H  217  ALA ASN TRP GLY PRO GLY THR LEU VAL THR VAL SER SER          
SEQRES  10 H  217  ALA SER THR LYS GLY PRO SER VAL PHE PRO LEU ALA PRO          
SEQRES  11 H  217  SER SER LYS SER THR SER GLY GLY THR ALA ALA LEU GLY          
SEQRES  12 H  217  CYS LEU VAL LYS ASP TYR PHE PRO GLU PRO VAL THR VAL          
SEQRES  13 H  217  SER TRP ASN SER GLY ALA LEU THR SER GLY VAL HIS THR          
SEQRES  14 H  217  PHE PRO ALA VAL LEU GLN SER SER GLY LEU TYR SER LEU          
SEQRES  15 H  217  SER SER VAL VAL THR VAL PRO SER SER SER LEU GLY THR          
SEQRES  16 H  217  GLN THR TYR ILE CYS ASN VAL ASN HIS LYS PRO SER ASN          
SEQRES  17 H  217  THR LYS VAL ASP LYS LYS VAL GLU PRO                          
HET    SS1   1001       9                                                       
HETNAM     SS1 1-PHENYLETHANOL                                                  
HETSYN     SS1 (1S)-1-PHENYL-ETHANOL                                            
FORMUL   3  SS1    C8 H10 O                                                     
FORMUL   4  HOH   *398(H2 O)                                                    
HELIX    1   1 GLU L   81  PHE L   85  5                                   5    
HELIX    2   2 SER L  123  LYS L  128  1                                   6    
HELIX    3   3 LYS L  185  HIS L  191  1                                   7    
HELIX    4   4 GLY L  214  CYS L  216  5                                   3    
HELIX    5   5 GLU H   63  ARG H   66  5                                   4    
HELIX    6   6 THR H   88  SER H   92  5                                   5    
HELIX    7   7 SER H  133  THR H  137  5                                   5    
HELIX    8   8 SER H  162  ALA H  164  5                                   3    
HELIX    9   9 SER H  193  LEU H  195  5                                   3    
HELIX   10  10 LYS H  207  ASN H  210  5                                   4    
SHEET    1   A 4 MET L   6  SER L   9  0                                        
SHEET    2   A 4 VAL L  21  ALA L  27 -1  O  THR L  24   N  SER L   9           
SHEET    3   A 4 ASP L  72  ILE L  77 -1  O  LEU L  75   N  LEU L  23           
SHEET    4   A 4 PHE L  64  SER L  69 -1  N  SER L  69   O  ASP L  72           
SHEET    1   B 6 SER L  12  SER L  16  0                                        
SHEET    2   B 6 THR L 104  LYS L 109  1  O  GLU L 107   N  LEU L  13           
SHEET    3   B 6 ALA L  86  GLN L  92 -1  N  ALA L  86   O  VAL L 106           
SHEET    4   B 6 LEU L  35  GLN L  40 -1  N  GLN L  40   O  ASP L  87           
SHEET    5   B 6 ILE L  46  TYR L  51 -1  O  LEU L  49   N  TRP L  37           
SHEET    6   B 6 THR L  55  LEU L  56 -1  O  THR L  55   N  TYR L  51           
SHEET    1   C 4 SER L  12  SER L  16  0                                        
SHEET    2   C 4 THR L 104  LYS L 109  1  O  GLU L 107   N  LEU L  13           
SHEET    3   C 4 ALA L  86  GLN L  92 -1  N  ALA L  86   O  VAL L 106           
SHEET    4   C 4 THR L  99  PHE L 100 -1  O  THR L  99   N  GLN L  92           
SHEET    1   D 4 SER L 116  PHE L 120  0                                        
SHEET    2   D 4 THR L 131  PHE L 141 -1  O  ASN L 139   N  SER L 116           
SHEET    3   D 4 TYR L 175  SER L 184 -1  O  LEU L 177   N  LEU L 138           
SHEET    4   D 4 SER L 161  VAL L 165 -1  N  GLN L 162   O  THR L 180           
SHEET    1   E 4 ALA L 155  LEU L 156  0                                        
SHEET    2   E 4 LYS L 147  VAL L 152 -1  N  VAL L 152   O  ALA L 155           
SHEET    3   E 4 VAL L 193  THR L 199 -1  O  GLU L 197   N  GLN L 149           
SHEET    4   E 4 VAL L 207  ASN L 212 -1  O  VAL L 207   N  VAL L 198           
SHEET    1   F 4 LEU H   5  GLN H   7  0                                        
SHEET    2   F 4 VAL H  19  ALA H  25 -1  O  LYS H  24   N  GLN H   6           
SHEET    3   F 4 THR H  79  LEU H  84 -1  O  ALA H  80   N  CYS H  23           
SHEET    4   F 4 ALA H  69  ASP H  74 -1  N  THR H  70   O  GLU H  83           
SHEET    1   G 6 VAL H  11  VAL H  13  0                                        
SHEET    2   G 6 THR H 113  VAL H 117  1  O  THR H 116   N  VAL H  11           
SHEET    3   G 6 ALA H  93  SER H 101 -1  N  ALA H  93   O  VAL H 115           
SHEET    4   G 6 LEU H  34  GLN H  40 -1  N  VAL H  38   O  TYR H  96           
SHEET    5   G 6 LEU H  46  ILE H  52 -1  O  ILE H  49   N  TRP H  37           
SHEET    6   G 6 THR H  59  TYR H  61 -1  O  ARG H  60   N  TYR H  51           
SHEET    1   H 4 VAL H  11  VAL H  13  0                                        
SHEET    2   H 4 THR H 113  VAL H 117  1  O  THR H 116   N  VAL H  11           
SHEET    3   H 4 ALA H  93  SER H 101 -1  N  ALA H  93   O  VAL H 115           
SHEET    4   H 4 ASP H 104  TRP H 109 -1  O  ASP H 104   N  SER H 101           
SHEET    1   I 4 SER H 126  LEU H 130  0                                        
SHEET    2   I 4 THR H 141  TYR H 151 -1  O  LYS H 149   N  SER H 126           
SHEET    3   I 4 TYR H 182  PRO H 191 -1  O  LEU H 184   N  VAL H 148           
SHEET    4   I 4 VAL H 169  THR H 171 -1  N  HIS H 170   O  VAL H 187           
SHEET    1   J 4 SER H 126  LEU H 130  0                                        
SHEET    2   J 4 THR H 141  TYR H 151 -1  O  LYS H 149   N  SER H 126           
SHEET    3   J 4 TYR H 182  PRO H 191 -1  O  LEU H 184   N  VAL H 148           
SHEET    4   J 4 VAL H 175  LEU H 176 -1  N  VAL H 175   O  SER H 183           
SHEET    1   K 3 THR H 157  TRP H 160  0                                        
SHEET    2   K 3 ILE H 201  HIS H 206 -1  O  ASN H 203   N  SER H 159           
SHEET    3   K 3 THR H 211  LYS H 216 -1  O  VAL H 213   N  VAL H 204           
SSBOND   1 CYS L   25    CYS L   90                                             
SSBOND   2 CYS L  136    CYS L  196                                             
SSBOND   3 CYS H   23    CYS H   97                                             
SSBOND   4 CYS H  146    CYS H  202                                             
CISPEP   1 SER L    9    PRO L   10          0        -0.40                     
CISPEP   2 TYR L   96    PRO L   97          0         1.25                     
CISPEP   3 TYR L  142    PRO L  143          0        -1.53                     
CISPEP   4 PHE H  152    PRO H  153          0        -2.16                     
CISPEP   5 GLU H  154    PRO H  155          0         0.35                     
CRYST1   47.128   64.706  138.510  90.00  90.00  90.00 P 21 21 21    4          
ORIGX1      1.000000  0.000000  0.000000        0.00000                         
ORIGX2      0.000000  1.000000  0.000000        0.00000                         
ORIGX3      0.000000  0.000000  1.000000        0.00000                         
SCALE1      0.021219  0.000000  0.000000        0.00000                         
SCALE2      0.000000  0.015455  0.000000        0.00000                         
SCALE3      0.000000  0.000000  0.007220        0.00000