PDB entry 1sq0

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HEADER    BLOOD CLOTTING                          17-MAR-04   1SQ0              
TITLE     CRYSTAL STRUCTURE OF THE COMPLEX OF THE WILD-TYPE VON                 
TITLE    2 WILLEBRAND FACTOR A1 DOMAIN AND GLYCOPROTEIN IB ALPHA AT             
TITLE    3 2.6 ANGSTROM RESOLUTION                                              
COMPND    MOL_ID: 1;                                                            
COMPND   2 MOLECULE: VON WILLEBRAND FACTOR (VWF) [CONTAINS: VON                 
COMPND   3 WILLEBRAND ANTIGEN II];                                              
COMPND   4 CHAIN: A;                                                            
COMPND   5 FRAGMENT: A1;                                                        
COMPND   6 ENGINEERED: YES;                                                     
COMPND   7 MOL_ID: 2;                                                           
COMPND   8 MOLECULE: PLATELET GLYCOPROTEIN IB ALPHA CHAIN                       
COMPND   9 (GLYCOPROTEIN IBALPHA) (GP-IB ALPHA) (GPIBA) (GPIB-ALPHA)            
COMPND  10 (CD42B-ALPHA) (CD42B) [CONTAINS: GLYCOCALICIN];                      
COMPND  11 CHAIN: B;                                                            
COMPND  12 FRAGMENT: GLYCOPROTEIN IB-(ALPHA);                                   
COMPND  13 ENGINEERED: YES;                                                     
COMPND  14 MUTATION: YES                                                        
SOURCE    MOL_ID: 1;                                                            
SOURCE   2 ORGANISM_SCIENTIFIC: HOMO SAPIENS;                                   
SOURCE   3 ORGANISM_COMMON: HUMAN;                                              
SOURCE   4 GENE: VWF, F8VWF;                                                    
SOURCE   5 EXPRESSION_SYSTEM: ESCHERICHIA COLI;                                 
SOURCE   6 EXPRESSION_SYSTEM_COMMON: BACTERIA;                                  
SOURCE   7 EXPRESSION_SYSTEM_STRAIN: BL21DE3;                                   
SOURCE   8 EXPRESSION_SYSTEM_VECTOR_TYPE: PLASMID;                              
SOURCE   9 MOL_ID: 2;                                                           
SOURCE  10 ORGANISM_SCIENTIFIC: HOMO SAPIENS;                                   
SOURCE  11 ORGANISM_COMMON: HUMAN;                                              
SOURCE  12 GENE: GP1BA;                                                         
SOURCE  13 EXPRESSION_SYSTEM: CRICETULUS GRISEUS;                               
SOURCE  14 EXPRESSION_SYSTEM_COMMON: CHINESE HAMSTER;                           
SOURCE  15 EXPRESSION_SYSTEM_CELL: CHO                                          
KEYWDS    LEUCINE RICH REPEAT (LRR), RIGHT-HANDED BETA-ALPHA                    
KEYWDS   2 SUPERHELIX); INTEGRIN A (OR I) DOMAIN FOLD                           
EXPDTA    X-RAY DIFFRACTION                                                     
AUTHOR    J.J.DUMAS,R.KUMAR,T.MCDONAGH,F.SULLIVAN,M.L.STAHL,                    
AUTHOR   2 W.S.SOMERS,L.MOSYAK                                                  
REVDAT   2   31-AUG-04 1SQ0    1       JRNL                                     
REVDAT   1   13-APR-04 1SQ0    0                                                
JRNL        AUTH   J.J.DUMAS,R.KUMAR,T.MCDONAGH,F.SULLIVAN,M.L.STAHL,           
JRNL        AUTH 2 W.S.SOMERS,L.MOSYAK                                          
JRNL        TITL   CRYSTAL STRUCTURE OF THE WILD-TYPE VON WILLEBRAND            
JRNL        TITL 2 FACTOR A1-GLYCOPROTEIN IBALPHA COMPLEX REVEALS               
JRNL        TITL 3 CONFORMATION DIFFERENCES WITH A COMPLEX BEARING              
JRNL        TITL 4 VON WILLEBRAND DISEASE MUTATIONS                             
JRNL        REF    J.BIOL.CHEM.                  V. 279 23327 2004              
JRNL        REFN   ASTM JBCHA3  US ISSN 0021-9258                               
REMARK   1                                                                      
REMARK   1 REFERENCE 1                                                          
REMARK   1  AUTH   J.J.DUMAS,R.KUMAR,J.SEEHRA,W.S.SOMERS,L.MOSYAK               
REMARK   1  TITL   CRYSTAL STRUCTURE OF THE GPIB(ALPHA)-THROMBIN                
REMARK   1  TITL 2 COMPLEX ESSENTIAL FOR PLATELET AGGREGATION                   
REMARK   1  REF    SCIENCE                       V. 301   222 2003              
REMARK   1  REFN   ASTM SCIEAS  US ISSN 0036-8075                               
REMARK   2                                                                      
REMARK   2 RESOLUTION. 2.60 ANGSTROMS.                                          
REMARK   3                                                                      
REMARK   3 REFINEMENT.                                                          
REMARK   3   PROGRAM     : CNS 1.1                                              
REMARK   3   AUTHORS     : BRUNGER,ADAMS,CLORE,DELANO,GROS,GROSSE-              
REMARK   3               : KUNSTLEVE,JIANG,KUSZEWSKI,NILGES, PANNU,             
REMARK   3               : READ,RICE,SIMONSON,WARREN                            
REMARK   3                                                                      
REMARK   3  REFINEMENT TARGET : NULL                                            
REMARK   3                                                                      
REMARK   3  DATA USED IN REFINEMENT.                                            
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.60                           
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 49.58                          
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.000                          
REMARK   3   DATA CUTOFF HIGH         (ABS(F)) : 132767.560                     
REMARK   3   DATA CUTOFF LOW          (ABS(F)) : 0.0000                         
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : 93.9                           
REMARK   3   NUMBER OF REFLECTIONS             : 17150                          
REMARK   3                                                                      
REMARK   3  FIT TO DATA USED IN REFINEMENT.                                     
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT                      
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM                          
REMARK   3   R VALUE            (WORKING SET) : 0.190                           
REMARK   3   FREE R VALUE                     : 0.239                           
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 9.800                           
REMARK   3   FREE R VALUE TEST SET COUNT      : 1685                            
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : 0.006                           
REMARK   3                                                                      
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.                                  
REMARK   3   TOTAL NUMBER OF BINS USED           : 6                            
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 2.60                         
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 2.76                         
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 84.00                        
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : 2261                         
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2810                       
REMARK   3   BIN FREE R VALUE                    : 0.3220                       
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : 9.90                         
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : 248                          
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : 0.020                        
REMARK   3                                                                      
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.                    
REMARK   3   PROTEIN ATOMS            : 3666                                    
REMARK   3   NUCLEIC ACID ATOMS       : 0                                       
REMARK   3   HETEROGEN ATOMS          : 0                                       
REMARK   3   SOLVENT ATOMS            : 318                                     
REMARK   3                                                                      
REMARK   3  B VALUES.                                                           
REMARK   3   FROM WILSON PLOT           (A**2) : 28.50                          
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 40.60                          
REMARK   3   OVERALL ANISOTROPIC B VALUE.                                       
REMARK   3    B11 (A**2) : -2.42000                                             
REMARK   3    B22 (A**2) : -10.55000                                            
REMARK   3    B33 (A**2) : 12.98000                                             
REMARK   3    B12 (A**2) : 0.00000                                              
REMARK   3    B13 (A**2) : 0.00000                                              
REMARK   3    B23 (A**2) : 0.00000                                              
REMARK   3                                                                      
REMARK   3  ESTIMATED COORDINATE ERROR.                                         
REMARK   3   ESD FROM LUZZATI PLOT        (A) : 0.28                            
REMARK   3   ESD FROM SIGMAA              (A) : 0.36                            
REMARK   3   LOW RESOLUTION CUTOFF        (A) : 5.00                            
REMARK   3                                                                      
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.                         
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : 0.37                            
REMARK   3   ESD FROM C-V SIGMAA          (A) : 0.42                            
REMARK   3                                                                      
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.                                   
REMARK   3   BOND LENGTHS                 (A) : 0.006                           
REMARK   3   BOND ANGLES            (DEGREES) : 1.30                            
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : 26.00                           
REMARK   3   IMPROPER ANGLES        (DEGREES) : 0.87                            
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL MODEL : RESTRAINED                                
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA                
REMARK   3   MAIN-CHAIN BOND              (A**2) : 1.560 ; 1.500                
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : 2.620 ; 2.000                
REMARK   3   SIDE-CHAIN BOND              (A**2) : 2.340 ; 2.000                
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : 3.580 ; 2.500                
REMARK   3                                                                      
REMARK   3  BULK SOLVENT MODELING.                                              
REMARK   3   METHOD USED : FLAT MODEL                                           
REMARK   3   KSOL        : 0.31                                                 
REMARK   3   BSOL        : 34.43                                                
REMARK   3                                                                      
REMARK   3  NCS MODEL : NULL                                                    
REMARK   3                                                                      
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT          
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL                 
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL                 
REMARK   3                                                                      
REMARK   3  PARAMETER FILE  1  : PROTEIN_REP.PARAM                              
REMARK   3  PARAMETER FILE  2  : WATER_REP.PARAM                                
REMARK   3  PARAMETER FILE  3  : ION.PARAM                                      
REMARK   3  PARAMETER FILE  4  : NULL                                           
REMARK   3  TOPOLOGY FILE  1   : PROTEIN.TOP                                    
REMARK   3  TOPOLOGY FILE  2   : WATER.TOP                                      
REMARK   3  TOPOLOGY FILE  3   : ION.TOP                                        
REMARK   3  TOPOLOGY FILE  4   : NULL                                           
REMARK   3                                                                      
REMARK   3  OTHER REFINEMENT REMARKS: DISORDERED RESIDUES IN CHAIN A            
REMARK   3  INCLUDE N-TERMINAL RESIDUES 496-5 AND C-TERMINAL RESIDUES 704-      
REMARK   3  709. IN CHAIN A, SIDE CHAIN FOR LYS IS DISORDERED. DISORDERED       
REMARK   3  RESIDUES IN CHAIN B INCLUDE C-TERMINAL RESIDUES 266-288.            
REMARK   4                                                                      
REMARK   4 1SQ0 COMPLIES WITH FORMAT V. 3.0, 1-DEC-2006                         
REMARK   4                                                                      
REMARK   4 THIS IS THE REMEDIATED VERSION OF THIS PDB ENTRY.                    
REMARK   4 REMEDIATED DATA FILE REVISION 3.101 (2007-05-29)                     
REMARK 100                                                                      
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB .                              
REMARK 100 THE RCSB ID CODE IS RCSB021902.                                      
REMARK 200                                                                      
REMARK 200 EXPERIMENTAL DETAILS                                                 
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION                  
REMARK 200  DATE OF DATA COLLECTION        : 31-AUG-2002; 18-MAY-2002           
REMARK 200  TEMPERATURE           (KELVIN) : 100.0                              
REMARK 200  PH                             : 8.00                               
REMARK 200  NUMBER OF CRYSTALS USED        : 2                                  
REMARK 200                                                                      
REMARK 200  SYNCHROTRON              (Y/N) : Y; Y                               
REMARK 200  RADIATION SOURCE               : ALS; ALS                           
REMARK 200  BEAMLINE                       : 5.0.2; 5.0.2                       
REMARK 200  X-RAY GENERATOR MODEL          : NULL                               
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M; M                               
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.97942, 0.97959, 0.9686; 1        
REMARK 200  MONOCHROMATOR                  : DOUBLE-CRYSTAL, SI(111)            
REMARK 200  OPTICS                         : NULL                               
REMARK 200                                                                      
REMARK 200  DETECTOR TYPE                  : CCD; CCD                           
REMARK 200  DETECTOR MANUFACTURER          : ADSC QUANTUM 4; ADSC QUANTUM 4     
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : DENZO                              
REMARK 200  DATA SCALING SOFTWARE          : CCP4 (SCALA)                       
REMARK 200                                                                      
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 17150                              
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.600                              
REMARK 200  RESOLUTION RANGE LOW       (A) : 49.580                             
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 0.000                              
REMARK 200                                                                      
REMARK 200 OVERALL.                                                             
REMARK 200  COMPLETENESS FOR RANGE     (%) : NULL                               
REMARK 200  DATA REDUNDANCY                : NULL                               
REMARK 200  R MERGE                    (I) : NULL                               
REMARK 200  R SYM                      (I) : NULL                               
REMARK 200   FOR THE DATA SET  : NULL                               
REMARK 200                                                                      
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.                                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.60                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.75                     
REMARK 200  COMPLETENESS FOR SHELL     (%) : 84.0                               
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL                               
REMARK 200  R MERGE FOR SHELL          (I) : NULL                               
REMARK 200  R SYM FOR SHELL            (I) : NULL                               
REMARK 200   FOR SHELL         : NULL                               
REMARK 200                                                                      
REMARK 200 DIFFRACTION PROTOCOL: MAD                                            
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MAD                          
REMARK 200 SOFTWARE USED: SHELXS                                                
REMARK 200 STARTING MODEL: NULL                                                 
REMARK 200                                                                      
REMARK 200 REMARK: NULL                                                         
REMARK 280                                                                      
REMARK 280 CRYSTAL                                                              
REMARK 280 SOLVENT CONTENT, VS   (%): 56.50                                     
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.85                     
REMARK 280                                                                      
REMARK 280 CRYSTALLIZATION CONDITIONS: PEG-3350; POTASSIUM THIOCYANATE, PH      
REMARK 280  8.0, VAPOR DIFFUSION, HANGING DROP, TEMPERATURE 291K                
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY                                            
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 2                        
REMARK 290                                                                      
REMARK 290      SYMOP   SYMMETRY                                                
REMARK 290     NNNMMM   OPERATOR                                                
REMARK 290       1555   X,Y,Z                                                   
REMARK 290       2555   -X,-Y,Z                                                 
REMARK 290       3555   1/2-X,1/2+Y,-Z                                          
REMARK 290       4555   1/2+X,1/2-Y,-Z                                          
REMARK 290                                                                      
REMARK 290     WHERE NNN -> OPERATOR NUMBER                                     
REMARK 290           MMM -> TRANSLATION VECTOR                                  
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS                            
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM             
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY                
REMARK 290 RELATED MOLECULES.                                                   
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000       36.24850            
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       58.05200            
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000        0.00000            
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       36.24850            
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       58.05200            
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000            
REMARK 290                                                                      
REMARK 290 REMARK: NULL                                                         
REMARK 300                                                                      
REMARK 300 BIOMOLECULE: 1                                                       
REMARK 300 THIS ENTRY CONTAINS THE CRYSTALLOGRAPHIC ASYMMETRIC UNIT             
REMARK 300 WHICH CONSISTS OF 2 CHAIN(S). SEE REMARK 350 FOR                     
REMARK 300 INFORMATION ON GENERATING THE BIOLOGICAL MOLECULE(S).                
REMARK 350                                                                      
REMARK 350 GENERATING THE BIOMOLECULE                                           
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN           
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE                
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS          
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND                          
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.                               
REMARK 350                                                                      
REMARK 350 BIOMOLECULE: 1                                                       
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B                                  
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 465                                                                      
REMARK 465 MISSING RESIDUES                                                     
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE                       
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)                
REMARK 465                                                                      
REMARK 465   M RES C SSSEQI                                                     
REMARK 465     MET A   496                                                      
REMARK 465     GLU A   497                                                      
REMARK 465     ASP A   498                                                      
REMARK 465     ILE A   499                                                      
REMARK 465     SER A   500                                                      
REMARK 465     GLU A   501                                                      
REMARK 465     PRO A   502                                                      
REMARK 465     PRO A   503                                                      
REMARK 465     LEU A   504                                                      
REMARK 465     HIS A   505                                                      
REMARK 465     PRO A   704                                                      
REMARK 465     THR A   705                                                      
REMARK 465     LEU A   706                                                      
REMARK 465     PRO A   707                                                      
REMARK 465     PRO A   708                                                      
REMARK 465     HIS A   709                                                      
REMARK 465     THR B   266                                                      
REMARK 465     LEU B   267                                                      
REMARK 465     GLY B   268                                                      
REMARK 465     ASP B   269                                                      
REMARK 465     GLU B   270                                                      
REMARK 465     GLY B   271                                                      
REMARK 465     ASP B   272                                                      
REMARK 465     THR B   273                                                      
REMARK 465     ASP B   274                                                      
REMARK 465     LEU B   275                                                      
REMARK 465     TYR B   276                                                      
REMARK 465     ASP B   277                                                      
REMARK 465     TYR B   278                                                      
REMARK 465     TYR B   279                                                      
REMARK 465     PRO B   280                                                      
REMARK 465     GLU B   281                                                      
REMARK 465     GLU B   282                                                      
REMARK 465     ASP B   283                                                      
REMARK 465     THR B   284                                                      
REMARK 465     GLU B   285                                                      
REMARK 465     GLY B   286                                                      
REMARK 465     ASP B   287                                                      
REMARK 465     LYS B   288                                                      
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT                     
REMARK 500                                                                      
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.                            
REMARK 500                                                                      
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI                             
REMARK 500   O    HOH      21     O    HOH     243              2.17            
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND LENGTHS                                      
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,2(A3,1X,A1,I4,A1,1X,A4,3X),F6.3)                  
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   RES CSSEQI ATM2   DEVIATION                     
REMARK 500    MET A 541   SD    MET A 541   CE    -0.057                        
REMARK 500    PRO A 703   CB    PRO A 703   CG     0.050                        
REMARK 500    ARG B 218   CG    ARG B 218   CD     0.038                        
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES                                       
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)              
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3                                     
REMARK 500    ASP A 514   N   -  CA  -  C   ANGL. DEV. = -8.4 DEGREES           
REMARK 500    ARG A 552   N   -  CA  -  C   ANGL. DEV. = -8.2 DEGREES           
REMARK 500    GLU A 700   N   -  CA  -  C   ANGL. DEV. =  9.4 DEGREES           
REMARK 500    LEU B  76   N   -  CA  -  C   ANGL. DEV. = -8.3 DEGREES           
REMARK 500    ASN B 157   N   -  CA  -  C   ANGL. DEV. =  8.0 DEGREES           
REMARK 500    ASN B 158   N   -  CA  -  C   ANGL. DEV. = -8.4 DEGREES           
REMARK 500    ALA B 238   N   -  CA  -  C   ANGL. DEV. =  7.6 DEGREES           
REMARK 500    GLN B 247   N   -  CA  -  C   ANGL. DEV. =  8.6 DEGREES           
REMARK 500    GLY B 263   N   -  CA  -  C   ANGL. DEV. = 10.6 DEGREES           
REMARK 900                                                                      
REMARK 900 RELATED ENTRIES                                                      
REMARK 900 RELATED ID: 1M10   RELATED DB: PDB                                   
REMARK 900 CRYSTAL STRUCTURE OF THE COMPLEX OF GLYCOPROTEIN IB ALPHA            
REMARK 900 AND THE VON WILLEBRAND FACTOR A1 DOMAIN WITH GAIN-OF-                
REMARK 900 FUNCTION MUTATIONS                                                   
REMARK 900 RELATED ID: 1M0Z   RELATED DB: PDB                                   
REMARK 900 CRYSTAL STRUCTURE OF THE VON WILLEBRAND FACTOR BINDING               
REMARK 900 DOMAIN OF GLYCOPROTEIN IB ALPHA                                      
REMARK 900 RELATED ID: 1AUQ   RELATED DB: PDB                                   
REMARK 900 CRYSTAL STRUCTURE OF THE A1 DOMAIN OF VON WILLEBRAND FACTOR          
REMARK 900 RELATED ID: 1P8V   RELATED DB: PDB                                   
REMARK 900 CRYSTAL STRUCTURE OF THE COMPLEX OF GLYCOPROTEIN IB-ALPHA            
REMARK 900 AND ALPHA-THROMBIN                                                   
REMARK 900 RELATED ID: 1OOK   RELATED DB: PDB                                   
REMARK 900 CRYSTAL STRUCTURE OF THE COMPLEX OF GLYCOPROTEIN IB-ALPHA            
REMARK 900 AND ALPHA-THROMBIN                                                   
REMARK 999                                                                      
REMARK 999 SEQUENCE                                                             
REMARK 999 ASN RESIDUES AT POSITIONS 21 AND 159 OF CHAIN B, WHICH ARE           
REMARK 999 SITES FOR N-LINKED GLYCOSYLATION, WERE MUTATED TO ASP.               
DBREF  1SQ0 A  496   709  UNP    P04275   VWF_HUMAN     1259   1472             
DBREF  1SQ0 B    1   288  UNP    P07359   GP1BA_HUMAN     17    304             
SEQADV 1SQ0 MET A  496  UNP  P04275    VAL  1259 ENGINEERED                     
SEQADV 1SQ0 ASP B   21  UNP  P07359    ASN    37 SEE REMARK 999                 
SEQADV 1SQ0 ASP B  159  UNP  P07359    ASN   175 SEE REMARK 999                 
SEQRES   1 A  214  MET GLU ASP ILE SER GLU PRO PRO LEU HIS ASP PHE TYR          
SEQRES   2 A  214  CYS SER ARG LEU LEU ASP LEU VAL PHE LEU LEU ASP GLY          
SEQRES   3 A  214  SER SER ARG LEU SER GLU ALA GLU PHE GLU VAL LEU LYS          
SEQRES   4 A  214  ALA PHE VAL VAL ASP MET MET GLU ARG LEU ARG ILE SER          
SEQRES   5 A  214  GLN LYS TRP VAL ARG VAL ALA VAL VAL GLU TYR HIS ASP          
SEQRES   6 A  214  GLY SER HIS ALA TYR ILE GLY LEU LYS ASP ARG LYS ARG          
SEQRES   7 A  214  PRO SER GLU LEU ARG ARG ILE ALA SER GLN VAL LYS TYR          
SEQRES   8 A  214  ALA GLY SER GLN VAL ALA SER THR SER GLU VAL LEU LYS          
SEQRES   9 A  214  TYR THR LEU PHE GLN ILE PHE SER LYS ILE ASP ARG PRO          
SEQRES  10 A  214  GLU ALA SER ARG ILE ALA LEU LEU LEU MET ALA SER GLN          
SEQRES  11 A  214  GLU PRO GLN ARG MET SER ARG ASN PHE VAL ARG TYR VAL          
SEQRES  12 A  214  GLN GLY LEU LYS LYS LYS LYS VAL ILE VAL ILE PRO VAL          
SEQRES  13 A  214  GLY ILE GLY PRO HIS ALA ASN LEU LYS GLN ILE ARG LEU          
SEQRES  14 A  214  ILE GLU LYS GLN ALA PRO GLU ASN LYS ALA PHE VAL LEU          
SEQRES  15 A  214  SER SER VAL ASP GLU LEU GLU GLN GLN ARG ASP GLU ILE          
SEQRES  16 A  214  VAL SER TYR LEU CYS ASP LEU ALA PRO GLU ALA PRO PRO          
SEQRES  17 A  214  PRO THR LEU PRO PRO HIS                                      
SEQRES   1 B  288  HIS PRO ILE CYS GLU VAL SER LYS VAL ALA SER HIS LEU          
SEQRES   2 B  288  GLU VAL ASN CYS ASP LYS ARG ASP LEU THR ALA LEU PRO          
SEQRES   3 B  288  PRO ASP LEU PRO LYS ASP THR THR ILE LEU HIS LEU SER          
SEQRES   4 B  288  GLU ASN LEU LEU TYR THR PHE SER LEU ALA THR LEU MET          
SEQRES   5 B  288  PRO TYR THR ARG LEU THR GLN LEU ASN LEU ASP ARG CYS          
SEQRES   6 B  288  GLU LEU THR LYS LEU GLN VAL ASP GLY THR LEU PRO VAL          
SEQRES   7 B  288  LEU GLY THR LEU ASP LEU SER HIS ASN GLN LEU GLN SER          
SEQRES   8 B  288  LEU PRO LEU LEU GLY GLN THR LEU PRO ALA LEU THR VAL          
SEQRES   9 B  288  LEU ASP VAL SER PHE ASN ARG LEU THR SER LEU PRO LEU          
SEQRES  10 B  288  GLY ALA LEU ARG GLY LEU GLY GLU LEU GLN GLU LEU TYR          
SEQRES  11 B  288  LEU LYS GLY ASN GLU LEU LYS THR LEU PRO PRO GLY LEU          
SEQRES  12 B  288  LEU THR PRO THR PRO LYS LEU GLU LYS LEU SER LEU ALA          
SEQRES  13 B  288  ASN ASN ASP LEU THR GLU LEU PRO ALA GLY LEU LEU ASN          
SEQRES  14 B  288  GLY LEU GLU ASN LEU ASP THR LEU LEU LEU GLN GLU ASN          
SEQRES  15 B  288  SER LEU TYR THR ILE PRO LYS GLY PHE PHE GLY SER HIS          
SEQRES  16 B  288  LEU LEU PRO PHE ALA PHE LEU HIS GLY ASN PRO TRP LEU          
SEQRES  17 B  288  CYS ASN CYS GLU ILE LEU TYR PHE ARG ARG TRP LEU GLN          
SEQRES  18 B  288  ASP ASN ALA GLU ASN VAL TYR VAL TRP LYS GLN GLY VAL          
SEQRES  19 B  288  ASP VAL LYS ALA MET THR SER ASN VAL ALA SER VAL GLN          
SEQRES  20 B  288  CYS ASP ASN SER ASP LYS PHE PRO VAL TYR LYS TYR PRO          
SEQRES  21 B  288  GLY LYS GLY CYS PRO THR LEU GLY ASP GLU GLY ASP THR          
SEQRES  22 B  288  ASP LEU TYR ASP TYR TYR PRO GLU GLU ASP THR GLU GLY          
SEQRES  23 B  288  ASP LYS                                                      
FORMUL   3  HOH   *318(H2 O)                                                    
HELIX    1   1 SER A  526  LEU A  544  1                                  19    
HELIX    2   2 ARG A  573  VAL A  584  1                                  12    
HELIX    3   3 SER A  593  ILE A  605  1                                  13    
HELIX    4   4 PRO A  627  SER A  631  5                                   5    
HELIX    5   5 ASN A  633  LYS A  644  1                                  12    
HELIX    6   6 ASN A  658  ALA A  669  1                                  12    
HELIX    7   7 SER A  679  LEU A  683  5                                   5    
HELIX    8   8 GLN A  686  ASP A  696  1                                  11    
HELIX    9   9 ALA B   49  MET B   52  5                                   4    
HELIX   10  10 ASN B  210  GLU B  212  5                                   3    
HELIX   11  11 ILE B  213  ASN B  223  1                                  11    
HELIX   12  12 VAL B  243  VAL B  246  5                                   4    
HELIX   13  13 GLN B  247  SER B  251  5                                   5    
HELIX   14  14 PRO B  255  TYR B  259  5                                   5    
SHEET    1   A18 PHE A 675  LEU A 677  0                                        
SHEET    2   A18 VAL A 646  ILE A 653  1  N  GLY A 652   O  LEU A 677           
SHEET    3   A18 SER A 615  MET A 622  1  N  ARG A 616   O  ILE A 647           
SHEET    4   A18 LEU A 513  ASP A 520  1  N  VAL A 516   O  ILE A 617           
SHEET    5   A18 VAL A 551  TYR A 558  1  O  VAL A 556   N  PHE A 517           
SHEET    6   A18 SER A 562  ILE A 566 -1  O  HIS A 563   N  GLU A 557           
SHEET    7   A18 LYS B 237  SER B 241 -1  O  MET B 239   N  SER A 562           
SHEET    8   A18 VAL B 227  GLN B 232 -1  N  VAL B 229   O  THR B 240           
SHEET    9   A18 PHE B 199  PHE B 201  1  N  ALA B 200   O  TYR B 228           
SHEET   10   A18 THR B 176  LEU B 178  1  N  LEU B 177   O  PHE B 199           
SHEET   11   A18 LYS B 152  SER B 154  1  N  LEU B 153   O  THR B 176           
SHEET   12   A18 GLU B 128  TYR B 130  1  N  LEU B 129   O  LYS B 152           
SHEET   13   A18 VAL B 104  ASP B 106  1  N  LEU B 105   O  TYR B 130           
SHEET   14   A18 THR B  81  ASP B  83  1  N  LEU B  82   O  VAL B 104           
SHEET   15   A18 GLN B  59  ASN B  61  1  N  LEU B  60   O  THR B  81           
SHEET   16   A18 ILE B  35  HIS B  37  1  N  LEU B  36   O  ASN B  61           
SHEET   17   A18 HIS B  12  ASN B  16  1  N  VAL B  15   O  ILE B  35           
SHEET   18   A18 GLU B   5  VAL B   9 -1  N  VAL B   9   O  HIS B  12           
SHEET    1   B 2 THR B  45  SER B  47  0                                        
SHEET    2   B 2 LYS B  69  GLN B  71  1  O  GLN B  71   N  PHE B  46           
SSBOND   1 CYS A  509    CYS A  695                                             
SSBOND   2 CYS B    4    CYS B   17                                             
SSBOND   3 CYS B  209    CYS B  248                                             
SSBOND   4 CYS B  211    CYS B  264                                             
CRYST1   72.497  116.104   67.962  90.00  90.00  90.00 P 21 21 2     4          
ORIGX1      1.000000  0.000000  0.000000        0.00000                         
ORIGX2      0.000000  1.000000  0.000000        0.00000                         
ORIGX3      0.000000  0.000000  1.000000        0.00000                         
SCALE1      0.013794  0.000000  0.000000        0.00000                         
SCALE2      0.000000  0.008613  0.000000        0.00000                         
SCALE3      0.000000  0.000000  0.014714        0.00000