PDB entry 1rup

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HEADER    IMMUNE SYSTEM                           11-DEC-03   1RUP              
TITLE     CRYSTAL STRUCTURE (G) OF NATIVE CATIONIC CYCLIZATION                  
TITLE    2 ANTIBODY 4C6 FAB AT PH 8.5 WITH A DATA SET COLLECTED AT              
TITLE    3 APS BEAMLINE 19-ID                                                   
COMPND    MOL_ID: 1;                                                            
COMPND   2 MOLECULE: IMMUNOGLOBULIN IGG2A, HEAVY CHAIN;                         
COMPND   3 CHAIN: L;                                                            
COMPND   4 FRAGMENT: FAB;                                                       
COMPND   5 MOL_ID: 2;                                                           
COMPND   6 MOLECULE: IMMUNOGLOBULIN IGG2A, LIGHT CHAIN;                         
COMPND   7 CHAIN: H;                                                            
COMPND   8 FRAGMENT: FAB                                                        
SOURCE    MOL_ID: 1;                                                            
SOURCE   2 ORGANISM_SCIENTIFIC: MUS MUSCULUS;                                   
SOURCE   3 ORGANISM_COMMON: MOUSE;                                              
SOURCE   4 STRAIN: 129G1X+;                                                     
SOURCE   5 MOL_ID: 2;                                                           
SOURCE   6 ORGANISM_SCIENTIFIC: MUS MUSCULUS;                                   
SOURCE   7 ORGANISM_COMMON: MOUSE;                                              
SOURCE   8 STRAIN: 129G1X+                                                      
KEYWDS    IMMUNOGLOBULIN, CATALYTIC ANTIBODY, WATER OXIDATION, AMINO            
KEYWDS   2 ACID MODIFICATION                                                    
EXPDTA    X-RAY DIFFRACTION                                                     
AUTHOR    X.ZHU,P.WENTWORTH JR.,A.D.WENTWORTH,A.ESCHENMOSER,                    
AUTHOR   2 R.A.LERNER,I.A.WILSON                                                
REVDAT   1   02-MAR-04 1RUP    0                                                
JRNL        AUTH   X.ZHU,P.WENTWORTH JR.,A.D.WENTWORTH,A.ESCHENMOSER,           
JRNL        AUTH 2 R.A.LERNER,I.A.WILSON                                        
JRNL        TITL   PROBING THE ANTIBODY-CATALYZED WATER-OXIDATION               
JRNL        TITL 2 PATHWAY AT ATOMIC RESOLUTION.                                
JRNL        REF    PROC.NATL.ACAD.SCI.USA        V. 110  2247 2004              
JRNL        REFN   ASTM PNASA6  US ISSN 0027-8424                               
REMARK   1                                                                      
REMARK   1 REFERENCE 1                                                          
REMARK   1  AUTH   X.ZHU,A.HEINE,F.MONNAT,K.N.HOUK,K.D.JANDA,                   
REMARK   1  AUTH 2 I.A.WILSON                                                   
REMARK   1  TITL   STRUCTURAL BASIS FOR ANTIBODY CATALYSIS OF A                 
REMARK   1  TITL 2 CATIONIC CYCLIZATION REACTION                                
REMARK   1  REF    J.MOL.BIOL.                   V. 329    69 2003              
REMARK   1  REFN   ASTM JMOBAK  UK ISSN 0022-2836                               
REMARK   2                                                                      
REMARK   2 RESOLUTION. 1.40 ANGSTROMS.                                          
REMARK   3                                                                      
REMARK   3 REFINEMENT.                                                          
REMARK   3   PROGRAM     : SHELXL-97                                            
REMARK   3   AUTHORS     : G.M.SHELDRICK                                        
REMARK   3                                                                      
REMARK   3  DATA USED IN REFINEMENT.                                            
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.40                           
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 50.00                          
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.000                          
REMARK   3   COMPLETENESS FOR RANGE        (%) : 86.3                           
REMARK   3   CROSS-VALIDATION METHOD           : THROUGHOUT                     
REMARK   3   FREE R VALUE TEST SET SELECTION   : RANDOM                         
REMARK   3                                                                      
REMARK   3  FIT TO DATA USED IN REFINEMENT (NO CUTOFF).                         
REMARK   3   R VALUE   (WORKING + TEST SET, NO CUTOFF) : NULL                   
REMARK   3   R VALUE          (WORKING SET, NO CUTOFF) : 0.161                  
REMARK   3   FREE R VALUE                  (NO CUTOFF) : 0.212                  
REMARK   3   FREE R VALUE TEST SET SIZE (%, NO CUTOFF) : NULL                   
REMARK   3   FREE R VALUE TEST SET COUNT   (NO CUTOFF) : 5069                   
REMARK   3   TOTAL NUMBER OF REFLECTIONS   (NO CUTOFF) : 100805                 
REMARK   3                                                                      
REMARK   3  FIT/AGREEMENT OF MODEL FOR DATA WITH F>4SIG(F).                     
REMARK   3   R VALUE   (WORKING + TEST SET, F>4SIG(F)) : NULL                   
REMARK   3   R VALUE          (WORKING SET, F>4SIG(F)) : 0.155                  
REMARK   3   FREE R VALUE                  (F>4SIG(F)) : 0.206                  
REMARK   3   FREE R VALUE TEST SET SIZE (%, F>4SIG(F)) : NULL                   
REMARK   3   FREE R VALUE TEST SET COUNT   (F>4SIG(F)) : 4363                   
REMARK   3   TOTAL NUMBER OF REFLECTIONS   (F>4SIG(F)) : 86423                  
REMARK   3                                                                      
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.                    
REMARK   3   PROTEIN ATOMS      : 3398                                          
REMARK   3   NUCLEIC ACID ATOMS : 0                                             
REMARK   3   HETEROGEN ATOMS    : 9                                             
REMARK   3   SOLVENT ATOMS      : 495                                           
REMARK   3                                                                      
REMARK   3  MODEL REFINEMENT.                                                   
REMARK   3   OCCUPANCY SUM OF NON-HYDROGEN ATOMS      : NULL                    
REMARK   3   OCCUPANCY SUM OF HYDROGEN ATOMS          : NULL                    
REMARK   3   NUMBER OF DISCRETELY DISORDERED RESIDUES : NULL                    
REMARK   3   NUMBER OF LEAST-SQUARES PARAMETERS       : NULL                    
REMARK   3   NUMBER OF RESTRAINTS                     : NULL                    
REMARK   3                                                                      
REMARK   3  RMS DEVIATIONS FROM RESTRAINT TARGET VALUES.                        
REMARK   3   BOND LENGTHS                         (A) : 0.013                   
REMARK   3   ANGLE DISTANCES                      (A) : 0.030                   
REMARK   3   SIMILAR DISTANCES (NO TARGET VALUES) (A) : 0.000                   
REMARK   3   DISTANCES FROM RESTRAINT PLANES      (A) : 0.032                   
REMARK   3   ZERO CHIRAL VOLUMES               (A**3) : 0.058                   
REMARK   3   NON-ZERO CHIRAL VOLUMES           (A**3) : 0.073                   
REMARK   3   ANTI-BUMPING DISTANCE RESTRAINTS     (A) : 0.023                   
REMARK   3   RIGID-BOND ADP COMPONENTS         (A**2) : 0.004                   
REMARK   3   SIMILAR ADP COMPONENTS            (A**2) : 0.054                   
REMARK   3   APPROXIMATELY ISOTROPIC ADPS      (A**2) : 0.082                   
REMARK   3                                                                      
REMARK   3  BULK SOLVENT MODELING.                                              
REMARK   3   METHOD USED: MOEWS & KRETSINGER, J.MOL.BIOL.91(1973)201-228        
REMARK   3                                                                      
REMARK   3  STEREOCHEMISTRY TARGET VALUES : ENGH & HUBER                        
REMARK   3   SPECIAL CASE: NULL                                                 
REMARK   3                                                                      
REMARK   3  OTHER REFINEMENT REMARKS: ANISOTROPIC SCALING APPLIED BY THE        
REMARK   3  METHOD OF PARKIN, MOEZZI & HOPE, J.APPL.CRYST.28(1995)53-56         
REMARK   4                                                                      
REMARK   4 1RUP COMPLIES WITH FORMAT V. 3.0, 1-DEC-2006                         
REMARK   4                                                                      
REMARK   4 THIS IS THE REMEDIATED VERSION OF THIS PDB ENTRY.                    
REMARK   4 REMEDIATED DATA FILE REVISION 3.100 (2007-03-17)                     
REMARK 100                                                                      
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB .                              
REMARK 100 THE RCSB ID CODE IS RCSB021049.                                      
REMARK 200                                                                      
REMARK 200 EXPERIMENTAL DETAILS                                                 
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION                  
REMARK 200  DATE OF DATA COLLECTION        : 06-JUN-2001                        
REMARK 200  TEMPERATURE           (KELVIN) : 100.0                              
REMARK 200  PH                             : 8.50                               
REMARK 200  NUMBER OF CRYSTALS USED        : 1                                  
REMARK 200                                                                      
REMARK 200  SYNCHROTRON              (Y/N) : Y                                  
REMARK 200  RADIATION SOURCE               : APS                                
REMARK 200  BEAMLINE                       : 19-ID                              
REMARK 200  X-RAY GENERATOR MODEL          : NULL                               
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M                                  
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.0332                             
REMARK 200  MONOCHROMATOR                  : NULL                               
REMARK 200  OPTICS                         : NULL                               
REMARK 200                                                                      
REMARK 200  DETECTOR TYPE                  : CCD                                
REMARK 200  DETECTOR MANUFACTURER          : SBC2                               
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : HKL-2000                           
REMARK 200  DATA SCALING SOFTWARE          : SCALEPACK                          
REMARK 200                                                                      
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 104925                             
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.400                              
REMARK 200  RESOLUTION RANGE LOW       (A) : 50.000                             
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : -3.000                             
REMARK 200                                                                      
REMARK 200 OVERALL.                                                             
REMARK 200  COMPLETENESS FOR RANGE     (%) : 93.4                               
REMARK 200  DATA REDUNDANCY                : 7.200                              
REMARK 200  R MERGE                    (I) : NULL                               
REMARK 200  R SYM                      (I) : 0.07900                            
REMARK 200   FOR THE DATA SET  : 31.4000                            
REMARK 200                                                                      
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.                                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.40                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.43                     
REMARK 200  COMPLETENESS FOR SHELL     (%) : 90.7                               
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL                               
REMARK 200  R MERGE FOR SHELL          (I) : NULL                               
REMARK 200  R SYM FOR SHELL            (I) : 0.61600                            
REMARK 200   FOR SHELL         : 2.400                              
REMARK 200                                                                      
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH                              
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT        
REMARK 200 SOFTWARE USED: CNS                                                   
REMARK 200 STARTING MODEL: PDB ENTRY 1NCW                                       
REMARK 200                                                                      
REMARK 200 REMARK: NULL                                                         
REMARK 280                                                                      
REMARK 280 CRYSTAL                                                              
REMARK 280 SOLVENT CONTENT, VS   (%): 56.60                                     
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.86                     
REMARK 280                                                                      
REMARK 280 CRYSTALLIZATION CONDITIONS: 0.1 M TRIS HYDROCHLORIDE, 0.2 M          
REMARK 280  SODIUM ACETATE, 15% (W/V) PEG 4000 , PH 8.5, VAPOR DIFFUSION,       
REMARK 280  SITTING DROP, TEMPERATURE 295K                                      
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY                                            
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 41 21 2                        
REMARK 290                                                                      
REMARK 290      SYMOP   SYMMETRY                                                
REMARK 290     NNNMMM   OPERATOR                                                
REMARK 290       1555   X,Y,Z                                                   
REMARK 290       2555   -X,-Y,1/2+Z                                             
REMARK 290       3555   1/2-Y,1/2+X,1/4+Z                                       
REMARK 290       4555   1/2+Y,1/2-X,3/4+Z                                       
REMARK 290       5555   1/2-X,1/2+Y,1/4-Z                                       
REMARK 290       6555   1/2+X,1/2-Y,3/4-Z                                       
REMARK 290       7555   Y,X,-Z                                                  
REMARK 290       8555   -Y,-X,1/2-Z                                             
REMARK 290                                                                      
REMARK 290     WHERE NNN -> OPERATOR NUMBER                                     
REMARK 290           MMM -> TRANSLATION VECTOR                                  
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS                            
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM             
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY                
REMARK 290 RELATED MOLECULES.                                                   
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000      133.23750            
REMARK 290   SMTRY1   3  0.000000 -1.000000  0.000000       32.09850            
REMARK 290   SMTRY2   3  1.000000  0.000000  0.000000       32.09850            
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000       66.61875            
REMARK 290   SMTRY1   4  0.000000  1.000000  0.000000       32.09850            
REMARK 290   SMTRY2   4 -1.000000  0.000000  0.000000       32.09850            
REMARK 290   SMTRY3   4  0.000000  0.000000  1.000000      199.85625            
REMARK 290   SMTRY1   5 -1.000000  0.000000  0.000000       32.09850            
REMARK 290   SMTRY2   5  0.000000  1.000000  0.000000       32.09850            
REMARK 290   SMTRY3   5  0.000000  0.000000 -1.000000       66.61875            
REMARK 290   SMTRY1   6  1.000000  0.000000  0.000000       32.09850            
REMARK 290   SMTRY2   6  0.000000 -1.000000  0.000000       32.09850            
REMARK 290   SMTRY3   6  0.000000  0.000000 -1.000000      199.85625            
REMARK 290   SMTRY1   7  0.000000  1.000000  0.000000        0.00000            
REMARK 290   SMTRY2   7  1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY3   7  0.000000  0.000000 -1.000000        0.00000            
REMARK 290   SMTRY1   8  0.000000 -1.000000  0.000000        0.00000            
REMARK 290   SMTRY2   8 -1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY3   8  0.000000  0.000000 -1.000000      133.23750            
REMARK 290                                                                      
REMARK 290 REMARK: NULL                                                         
REMARK 300                                                                      
REMARK 300 BIOMOLECULE: 1                                                       
REMARK 300 THIS ENTRY CONTAINS THE CRYSTALLOGRAPHIC ASYMMETRIC UNIT             
REMARK 300 WHICH CONSISTS OF 2 CHAIN(S). SEE REMARK 350 FOR                     
REMARK 300 INFORMATION ON GENERATING THE BIOLOGICAL MOLECULE(S).                
REMARK 350                                                                      
REMARK 350 GENERATING THE BIOMOLECULE                                           
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN           
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE                
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS          
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND                          
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.                               
REMARK 350                                                                      
REMARK 350 BIOMOLECULE: 1                                                       
REMARK 350 APPLY THE FOLLOWING TO CHAINS: L, H                                  
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES                                       
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)              
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3                                     
REMARK 500    LEU H  18   CA  -  CB  -  CG  ANGL. DEV. = 22.4 DEGREES           
REMARK 500    SER H  65   C   -  N   -  CA  ANGL. DEV. = 15.6 DEGREES           
REMARK 900                                                                      
REMARK 900 RELATED ENTRIES                                                      
REMARK 900 RELATED ID: 1NCW   RELATED DB: PDB                                   
REMARK 900 SAME NATIVE PROTEIN                                                  
REMARK 900 RELATED ID: 1ND0   RELATED DB: PDB                                   
REMARK 900 SAME NATIVE PROTEIN COMPLEX WITH TRANSITION STATE ANALOG             
REMARK 900 RELATED ID: 1RU9   RELATED DB: PDB                                   
REMARK 900 CRYSTAL STRUCTURE OF U.V.-IRRADIATED CATIONIC CYCLIZATION            
REMARK 900 ANTIBODY 4C6 FAB AT PH 4.6 WITH A DATA SET COLLECTED IN-             
REMARK 900 HOUSE.                                                               
REMARK 900 RELATED ID: 1RUA   RELATED DB: PDB                                   
REMARK 900 CRYSTAL STRUCTURE OF U.V.-IRRADIATED CATIONIC CYCLIZATION            
REMARK 900 ANTIBODY 4C6 FAB AT PH 4.6 WITH A DATA SET COLLECTED AT              
REMARK 900 SSRL BEAMLINE 11-1.                                                  
REMARK 900 RELATED ID: 1RUK   RELATED DB: PDB                                   
REMARK 900 CRYSTAL STRUCTURE OF NATIVE CATIONIC CYCLIZATION ANTIBODY            
REMARK 900 4C6 FAB AT PH 4.6 WITH A DATA SET COLLECTED AT SSRL                  
REMARK 900 BEAMLINE 9-1                                                         
REMARK 900 RELATED ID: 1RUL   RELATED DB: PDB                                   
REMARK 900 CRYSTAL STRUCTURE OF U.V.-IRRADIATED CATIONIC CYCLIZATION            
REMARK 900 ANTIBODY 4C6 FAB AT PH 5.6 WITH A DATA SET COLLECTED AT              
REMARK 900 SSRL BEAMLINE 11-1.                                                  
REMARK 900 RELATED ID: 1RUM   RELATED DB: PDB                                   
REMARK 900 CRYSTAL STRUCTURE OF H2O2-SOAKED CATIONIC CYCLIZATION                
REMARK 900 ANTIBODY 4C6 FAB AT PH 8.5 WITH A DATA SET COLLECTED AT              
REMARK 900 SSRL BEAMLINE 9-1.                                                   
DBREF  1RUP H  115   231  UNP    P01865   GCAM_MOUSE       1    102             
SEQRES   1 L  219  ASP VAL VAL MET THR GLN SER PRO LYS THR ILE SER VAL          
SEQRES   2 L  219  THR ILE GLY GLN PRO ALA SER ILE SER CYS LYS SER SER          
SEQRES   3 L  219  GLN ARG LEU LEU ASN SER ASN GLY LYS THR PHE LEU ASN          
SEQRES   4 L  219  TRP LEU LEU GLN ARG PRO GLY GLN SER PRO LYS ARG LEU          
SEQRES   5 L  219  ILE TYR LEU GLY THR LYS LEU ASP SER GLY VAL PRO ASP          
SEQRES   6 L  219  ARG PHE THR GLY SER GLY SER GLY THR ASP PHE THR LEU          
SEQRES   7 L  219  LYS ILE SER ARG VAL GLU ALA GLU ASP LEU GLY VAL TYR          
SEQRES   8 L  219  TYR CYS TRP GLN GLY THR HIS PHE PRO TYR THR PHE GLY          
SEQRES   9 L  219  GLY GLY THR LYS LEU GLU ILE LYS ARG ALA ASP ALA ALA          
SEQRES  10 L  219  PRO THR VAL SER ILE PHE PRO PRO SER SER GLU GLN LEU          
SEQRES  11 L  219  THR SER GLY GLY ALA SER VAL VAL CYS PHE LEU ASN ASN          
SEQRES  12 L  219  PHE TYR PRO LYS ASP ILE ASN VAL LYS TRP LYS ILE ASP          
SEQRES  13 L  219  GLY SER GLU ARG GLN ASN GLY VAL LEU ASN SER TRP THR          
SEQRES  14 L  219  ASP GLN ASP SER LYS ASP SER THR TYR SER MET SER SER          
SEQRES  15 L  219  THR LEU THR LEU THR LYS ASP GLU TYR GLU ARG HIS ASN          
SEQRES  16 L  219  SER TYR THR CYS GLU ALA THR HIS LYS THR SER THR SER          
SEQRES  17 L  219  PRO ILE VAL LYS SER PHE ASN ARG ASN GLU CYS                  
SEQRES   1 H  222  ARG VAL GLN LEU GLN GLN SER GLY PRO GLY LEU VAL LYS          
SEQRES   2 H  222  PRO SER GLN SER LEU SER LEU THR CYS THR VAL THR GLY          
SEQRES   3 H  222  TYR SER ILE THR SER ASP PHE ALA TRP ASN TRP ILE ARG          
SEQRES   4 H  222  GLN PHE PRO GLY ASN LYS LEU GLU TRP MET GLY TYR ILE          
SEQRES   5 H  222  ASN TYR SER GLY PHE THR SER HIS ASN PRO SER LEU LYS          
SEQRES   6 H  222  SER ARG ILE SER ILE THR ARG ASP THR SER LYS ASN GLN          
SEQRES   7 H  222  PHE PHE LEU GLN LEU ASN SER VAL THR THR GLU ASP THR          
SEQRES   8 H  222  ALA THR TYR TYR CYS ALA GLY LEU LEU TRP TYR ASP GLY          
SEQRES   9 H  222  GLY ALA GLY SER TRP GLY GLN GLY THR LEU VAL THR VAL          
SEQRES  10 H  222  SER ALA ALA LYS THR THR ALA PRO SER VAL TYR PRO LEU          
SEQRES  11 H  222  ALA PRO VAL CYS GLY ASP THR THR GLY SER SER VAL THR          
SEQRES  12 H  222  LEU GLY CYS LEU VAL LYS GLY TYR PHE PRO GLU PRO VAL          
SEQRES  13 H  222  THR LEU THR TRP ASN SER GLY SER LEU SER SER GLY VAL          
SEQRES  14 H  222  HIS THR PHE PRO ALA VAL LEU GLN SER ASP LEU TYR THR          
SEQRES  15 H  222  LEU SER SER SER VAL THR VAL THR SER SER THR TRP PRO          
SEQRES  16 H  222  SER GLN SER ILE THR CYS ASN VAL ALA HIS PRO ALA SER          
SEQRES  17 H  222  SER THR LYS VAL ASP LYS LYS ILE GLU PRO ARG GLY PRO          
SEQRES  18 H  222  THR                                                          
HET    BEZ    601       9                                                       
HET    GOL    611       6                                                       
HET    GOL    612       6                                                       
HET    GOL    613       6                                                       
HET    GOL    614       6                                                       
HETNAM     BEZ BENZOIC ACID                                                     
HETNAM     GOL GLYCEROL                                                         
FORMUL   3  BEZ    C7 H6 O2                                                     
FORMUL   4  GOL    4(C3 H8 O3)                                                  
FORMUL   8  HOH   *471(H2 O)                                                    
HELIX    1   1 GLU L   79  LEU L   83  5                                   5    
HELIX    2   2 SER L  121  SER L  127  1                                   7    
HELIX    3   3 LYS L  183  HIS L  189  1                                   7    
HELIX    4   4 PRO H   61  LYS H   64  5                                   4    
HELIX    5   5 THR H   83  THR H   87  5                                   5    
HELIX    6   6 VAL H  127  THR H  133  5                                   5    
HELIX    7   7 SER H  163  SER H  165  5                                   3    
HELIX    8   8 SER H  196  TRP H  199  5                                   3    
HELIX    9   9 PRO H  213  SER H  216  5                                   4    
SHEET    1   A 4 MET L   4  SER L   7  0                                        
SHEET    2   A 4 ALA L  19  SER L  25 -1  O  SER L  22   N  SER L   7           
SHEET    3   A 4 ASP L  70  ILE L  75 -1  O  PHE L  71   N  CYS L  23           
SHEET    4   A 4 PHE L  62  SER L  67 -1  N  THR L  63   O  LYS L  74           
SHEET    1   B 6 THR L  10  THR L  14  0                                        
SHEET    2   B 6 THR L 102  LYS L 107  1  O  GLU L 105   N  ILE L  11           
SHEET    3   B 6 GLY L  84  GLN L  90 -1  N  GLY L  84   O  LEU L 104           
SHEET    4   B 6 LEU L  33  GLN L  38 -1  N  GLN L  38   O  VAL L  85           
SHEET    5   B 6 PRO L  44  TYR L  49 -1  O  LYS L  45   N  LEU L  37           
SHEET    6   B 6 LYS L  53  LEU L  54 -1  O  LYS L  53   N  TYR L  49           
SHEET    1   C 4 THR L  10  THR L  14  0                                        
SHEET    2   C 4 THR L 102  LYS L 107  1  O  GLU L 105   N  ILE L  11           
SHEET    3   C 4 GLY L  84  GLN L  90 -1  N  GLY L  84   O  LEU L 104           
SHEET    4   C 4 THR L  97  PHE L  98 -1  O  THR L  97   N  GLN L  90           
SHEET    1   D 4 THR L 114  PHE L 118  0                                        
SHEET    2   D 4 GLY L 129  PHE L 139 -1  O  ASN L 137   N  THR L 114           
SHEET    3   D 4 TYR L 173  THR L 182 -1  O  SER L 177   N  CYS L 134           
SHEET    4   D 4 VAL L 159  TRP L 163 -1  N  SER L 162   O  SER L 176           
SHEET    1   E 4 SER L 153  GLU L 154  0                                        
SHEET    2   E 4 ASN L 145  ILE L 150 -1  N  ILE L 150   O  SER L 153           
SHEET    3   E 4 SER L 191  HIS L 198 -1  O  THR L 197   N  ASN L 145           
SHEET    4   E 4 SER L 201  ASN L 210 -1  O  ILE L 205   N  ALA L 196           
SHEET    1   F 4 GLN H   3  SER H   7  0                                        
SHEET    2   F 4 LEU H  18  THR H  25 -1  O  THR H  21   N  SER H   7           
SHEET    3   F 4 GLN H  77  LEU H  82 -1  O  PHE H  78   N  CYS H  22           
SHEET    4   F 4 ILE H  67  ASP H  72 -1  N  THR H  70   O  PHE H  79           
SHEET    1   G 6 LEU H  11  VAL H  12  0                                        
SHEET    2   G 6 THR H 107  VAL H 111  1  O  THR H 110   N  VAL H  12           
SHEET    3   G 6 ALA H  88  LEU H  96 -1  N  TYR H  90   O  THR H 107           
SHEET    4   G 6 PHE H  33  PHE H  40 -1  N  ILE H  37   O  TYR H  91           
SHEET    5   G 6 LYS H  44  ASN H  52 -1  O  MET H  48   N  TRP H  36           
SHEET    6   G 6 THR H  57  HIS H  59 -1  O  SER H  58   N  TYR H  50           
SHEET    1   H 4 LEU H  11  VAL H  12  0                                        
SHEET    2   H 4 THR H 107  VAL H 111  1  O  THR H 110   N  VAL H  12           
SHEET    3   H 4 ALA H  88  LEU H  96 -1  N  TYR H  90   O  THR H 107           
SHEET    4   H 4 SER H 102  TRP H 103 -1  O  SER H 102   N  GLY H  94           
SHEET    1   I 4 SER H 120  LEU H 124  0                                        
SHEET    2   I 4 SER H 137  TYR H 147 -1  O  LEU H 143   N  TYR H 122           
SHEET    3   I 4 LEU H 184  THR H 194 -1  O  LEU H 187   N  VAL H 144           
SHEET    4   I 4 VAL H 171  THR H 173 -1  N  HIS H 172   O  SER H 190           
SHEET    1   J 4 SER H 120  LEU H 124  0                                        
SHEET    2   J 4 SER H 137  TYR H 147 -1  O  LEU H 143   N  TYR H 122           
SHEET    3   J 4 LEU H 184  THR H 194 -1  O  LEU H 187   N  VAL H 144           
SHEET    4   J 4 VAL H 177  GLN H 179 -1  N  GLN H 179   O  LEU H 184           
SHEET    1   K 3 THR H 153  TRP H 157  0                                        
SHEET    2   K 3 THR H 206  HIS H 212 -1  O  ASN H 209   N  THR H 156           
SHEET    3   K 3 THR H 217  LYS H 222 -1  O  VAL H 219   N  VAL H 210           
SSBOND   1 CYS L   23    CYS L   88                                             
SSBOND   2 CYS L  134    CYS L  194                                             
SSBOND   3 CYS L  214    CYS H  128                                             
SSBOND   4 CYS H   22    CYS H   92                                             
SSBOND   5 CYS H  142    CYS H  208                                             
CISPEP   1 SER L    7    PRO L    8          0        -3.07                     
CISPEP   2 PHE L   94    PRO L   95          0        -8.42                     
CISPEP   3 TYR L  140    PRO L  141          0        -2.36                     
CISPEP   4 PHE H  148    PRO H  149          0        -6.23                     
CISPEP   5 GLU H  150    PRO H  151          0        -4.19                     
CISPEP   6 TRP H  199    PRO H  200          0         9.67                     
CRYST1   64.197   64.197  266.475  90.00  90.00  90.00 P 41 21 2     8          
ORIGX1      1.000000  0.000000  0.000000        0.00000                         
ORIGX2      0.000000  1.000000  0.000000        0.00000                         
ORIGX3      0.000000  0.000000  1.000000        0.00000                         
SCALE1      0.015577  0.000000  0.000000        0.00000                         
SCALE2      0.000000  0.015577  0.000000        0.00000                         
SCALE3      0.000000  0.000000  0.003753        0.00000