PDB entry 1pbx

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HEADER    OXYGEN TRANSPORT                        04-NOV-91   1PBX              
TITLE     HAEMOGLOBIN OF THE ANTARCTIC FISH PAGOTHENIA BERNACCHII:              
TITLE    2 AMINO ACID SEQUENCE, OXYGEN EQUILIBRIA AND CRYSTAL                   
TITLE    3 STRUCTURE OF ITS CARBONMONOXY DERIVATIVE                             
COMPND    MOL_ID: 1;                                                            
COMPND   2 MOLECULE: HEMOGLOBIN (CARBONMONOXY) (ALPHA CHAIN);                   
COMPND   3 CHAIN: A;                                                            
COMPND   4 ENGINEERED: YES;                                                     
COMPND   5 MOL_ID: 2;                                                           
COMPND   6 MOLECULE: HEMOGLOBIN (CARBONMONOXY) (BETA CHAIN);                    
COMPND   7 CHAIN: B;                                                            
COMPND   8 ENGINEERED: YES                                                      
SOURCE    MOL_ID: 1;                                                            
SOURCE   2 ORGANISM_SCIENTIFIC: PAGOTHENIA BERNACCHII;                          
SOURCE   3 MOL_ID: 2;                                                           
SOURCE   4 ORGANISM_SCIENTIFIC: PAGOTHENIA BERNACCHII                           
KEYWDS    OXYGEN TRANSPORT                                                      
EXPDTA    X-RAY DIFFRACTION                                                     
AUTHOR    G.FERMI                                                               
REVDAT   2   01-APR-03 1PBX    1       JRNL                                     
REVDAT   1   15-OCT-92 1PBX    0                                                
JRNL        AUTH   L.CAMARDELLA,C.CARUSO,R.D'AVINO,G.DI PRISCO,                 
JRNL        AUTH 2 B.RUTIGLIANO,M.TAMBURRINI,G.FERMI,M.F.PERUTZ                 
JRNL        TITL   HAEMOGLOBIN OF THE ANTARCTIC FISH PAGOTHENIA                 
JRNL        TITL 2 BERNACCHII. AMINO ACID SEQUENCE, OXYGEN EQUILIBRIA           
JRNL        TITL 3 AND CRYSTAL STRUCTURE OF ITS CARBONMONOXY                    
JRNL        TITL 4 DERIVATIVE.                                                  
JRNL        REF    J.MOL.BIOL.                   V. 224   449 1992              
JRNL        REFN   ASTM JMOBAK  UK ISSN 0022-2836                               
REMARK   1                                                                      
REMARK   2                                                                      
REMARK   2 RESOLUTION. 2.50 ANGSTROMS.                                          
REMARK   3                                                                      
REMARK   3 REFINEMENT.                                                          
REMARK   3   PROGRAM     : X-PLOR                                               
REMARK   3   AUTHORS     : BRUNGER                                              
REMARK   3                                                                      
REMARK   3  DATA USED IN REFINEMENT.                                            
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.50                           
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 10.00                          
REMARK   3   DATA CUTOFF            (SIGMA(F)) : NULL                           
REMARK   3   DATA CUTOFF HIGH         (ABS(F)) : NULL                           
REMARK   3   DATA CUTOFF LOW          (ABS(F)) : NULL                           
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : NULL                           
REMARK   3   NUMBER OF REFLECTIONS             : 13785                          
REMARK   3                                                                      
REMARK   3  FIT TO DATA USED IN REFINEMENT.                                     
REMARK   3   CROSS-VALIDATION METHOD          : NULL                            
REMARK   3   FREE R VALUE TEST SET SELECTION  : NULL                            
REMARK   3   R VALUE            (WORKING SET) : 0.178                           
REMARK   3   FREE R VALUE                     : NULL                            
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : NULL                            
REMARK   3   FREE R VALUE TEST SET COUNT      : NULL                            
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : NULL                            
REMARK   3                                                                      
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.                                  
REMARK   3   TOTAL NUMBER OF BINS USED           : NULL                         
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : NULL                         
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : NULL                         
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : NULL                         
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : NULL                         
REMARK   3   BIN R VALUE           (WORKING SET) : NULL                         
REMARK   3   BIN FREE R VALUE                    : NULL                         
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : NULL                         
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : NULL                         
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : NULL                         
REMARK   3                                                                      
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.                    
REMARK   3   PROTEIN ATOMS            : 2242                                    
REMARK   3   NUCLEIC ACID ATOMS       : 0                                       
REMARK   3   HETEROGEN ATOMS          : 90                                      
REMARK   3   SOLVENT ATOMS            : 28                                      
REMARK   3                                                                      
REMARK   3  B VALUES.                                                           
REMARK   3   FROM WILSON PLOT           (A**2) : NULL                           
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : NULL                           
REMARK   3   OVERALL ANISOTROPIC B VALUE.                                       
REMARK   3    B11 (A**2) : NULL                                                 
REMARK   3    B22 (A**2) : NULL                                                 
REMARK   3    B33 (A**2) : NULL                                                 
REMARK   3    B12 (A**2) : NULL                                                 
REMARK   3    B13 (A**2) : NULL                                                 
REMARK   3    B23 (A**2) : NULL                                                 
REMARK   3                                                                      
REMARK   3  ESTIMATED COORDINATE ERROR.                                         
REMARK   3   ESD FROM LUZZATI PLOT        (A) : NULL                            
REMARK   3   ESD FROM SIGMAA              (A) : NULL                            
REMARK   3   LOW RESOLUTION CUTOFF        (A) : NULL                            
REMARK   3                                                                      
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.                         
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : NULL                            
REMARK   3   ESD FROM C-V SIGMAA          (A) : NULL                            
REMARK   3                                                                      
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.                                   
REMARK   3   BOND LENGTHS                 (A) : 0.019                           
REMARK   3   BOND ANGLES            (DEGREES) : 2.00                            
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : NULL                            
REMARK   3   IMPROPER ANGLES        (DEGREES) : NULL                            
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL MODEL : NULL                                      
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA                
REMARK   3   MAIN-CHAIN BOND              (A**2) : NULL  ; NULL                 
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : NULL  ; NULL                 
REMARK   3   SIDE-CHAIN BOND              (A**2) : NULL  ; NULL                 
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : NULL  ; NULL                 
REMARK   3                                                                      
REMARK   3  NCS MODEL : NULL                                                    
REMARK   3                                                                      
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT          
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL                 
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL                 
REMARK   3                                                                      
REMARK   3  PARAMETER FILE  1  : NULL                                           
REMARK   3  TOPOLOGY FILE  1   : NULL                                           
REMARK   3                                                                      
REMARK   3  OTHER REFINEMENT REMARKS: NULL                                      
REMARK   4                                                                      
REMARK   4 1PBX COMPLIES WITH FORMAT V. 3.0, 1-DEC-2006                         
REMARK   4                                                                      
REMARK   4 THIS IS THE REMEDIATED VERSION OF THIS PDB ENTRY.                    
REMARK   4 REMEDIATED DATA FILE REVISION 3.100 (2007-03-23)                     
REMARK 200                                                                      
REMARK 200 EXPERIMENTAL DETAILS                                                 
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION                  
REMARK 200  DATE OF DATA COLLECTION        : NULL                               
REMARK 200  TEMPERATURE           (KELVIN) : NULL                               
REMARK 200  PH                             : NULL                               
REMARK 200  NUMBER OF CRYSTALS USED        : NULL                               
REMARK 200                                                                      
REMARK 200  SYNCHROTRON              (Y/N) : NULL                               
REMARK 200  RADIATION SOURCE               : NULL                               
REMARK 200  BEAMLINE                       : NULL                               
REMARK 200  X-RAY GENERATOR MODEL          : NULL                               
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : NULL                               
REMARK 200  WAVELENGTH OR RANGE        (A) : NULL                               
REMARK 200  MONOCHROMATOR                  : NULL                               
REMARK 200  OPTICS                         : NULL                               
REMARK 200                                                                      
REMARK 200  DETECTOR TYPE                  : NULL                               
REMARK 200  DETECTOR MANUFACTURER          : NULL                               
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : NULL                               
REMARK 200  DATA SCALING SOFTWARE          : NULL                               
REMARK 200                                                                      
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : NULL                               
REMARK 200  RESOLUTION RANGE HIGH      (A) : NULL                               
REMARK 200  RESOLUTION RANGE LOW       (A) : NULL                               
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL                               
REMARK 200                                                                      
REMARK 200 OVERALL.                                                             
REMARK 200  COMPLETENESS FOR RANGE     (%) : NULL                               
REMARK 200  DATA REDUNDANCY                : NULL                               
REMARK 200  R MERGE                    (I) : NULL                               
REMARK 200  R SYM                      (I) : NULL                               
REMARK 200   FOR THE DATA SET  : NULL                               
REMARK 200                                                                      
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.                                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : NULL                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : NULL                     
REMARK 200  COMPLETENESS FOR SHELL     (%) : NULL                               
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL                               
REMARK 200  R MERGE FOR SHELL          (I) : NULL                               
REMARK 200  R SYM FOR SHELL            (I) : NULL                               
REMARK 200   FOR SHELL         : NULL                               
REMARK 200                                                                      
REMARK 200 DIFFRACTION PROTOCOL: NULL                                           
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: NULL                         
REMARK 200 SOFTWARE USED: NULL                                                  
REMARK 200 STARTING MODEL: NULL                                                 
REMARK 200                                                                      
REMARK 200 REMARK: NULL                                                         
REMARK 280                                                                      
REMARK 280 CRYSTAL                                                              
REMARK 280 SOLVENT CONTENT, VS   (%): 64.73                                     
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 3.49                     
REMARK 280                                                                      
REMARK 280 CRYSTALLIZATION CONDITIONS: NULL                                     
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY                                            
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: C 1 2 1                          
REMARK 290                                                                      
REMARK 290      SYMOP   SYMMETRY                                                
REMARK 290     NNNMMM   OPERATOR                                                
REMARK 290       1555   X,Y,Z                                                   
REMARK 290       2555   -X,Y,-Z                                                 
REMARK 290       3555   1/2+X,1/2+Y,Z                                           
REMARK 290       4555   1/2-X,1/2+Y,-Z                                          
REMARK 290                                                                      
REMARK 290     WHERE NNN -> OPERATOR NUMBER                                     
REMARK 290           MMM -> TRANSLATION VECTOR                                  
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS                            
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM             
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY                
REMARK 290 RELATED MOLECULES.                                                   
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   2  0.000000  1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   2  0.000000  0.000000 -1.000000        0.00000            
REMARK 290   SMTRY1   3  1.000000  0.000000  0.000000       45.69000            
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       44.27000            
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   4 -1.000000  0.000000  0.000000       45.69000            
REMARK 290   SMTRY2   4  0.000000  1.000000  0.000000       44.27000            
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000            
REMARK 290                                                                      
REMARK 290 REMARK: NULL                                                         
REMARK 300                                                                      
REMARK 300 BIOMOLECULE: 1                                                       
REMARK 300 THIS ENTRY CONTAINS THE CRYSTALLOGRAPHIC ASYMMETRIC UNIT             
REMARK 300 WHICH CONSISTS OF 2 CHAIN(S). SEE REMARK 350 FOR                     
REMARK 300 INFORMATION ON GENERATING THE BIOLOGICAL MOLECULE(S).                
REMARK 350                                                                      
REMARK 350 GENERATING THE BIOMOLECULE                                           
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN           
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE                
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS          
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND                          
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.                               
REMARK 350                                                                      
REMARK 350 BIOMOLECULE: 1                                                       
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B                                  
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 350   BIOMT1   2 -1.000000  0.000000  0.000000        0.00000            
REMARK 350   BIOMT2   2  0.000000  1.000000  0.000000        0.00000            
REMARK 350   BIOMT3   2  0.000000  0.000000 -1.000000        0.00000            
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT                     
REMARK 500                                                                      
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.                            
REMARK 500                                                                      
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI                             
REMARK 500   NE2  HIS A    88    FE    HEM A   144              2.14            
REMARK 500   NE2  HIS B    92    FE    HEM B   148              2.15            
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: TORSION ANGLES                                             
REMARK 500                                                                      
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:            
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;               
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                             
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)                    
REMARK 500                                                                      
REMARK 500  M RES CSSEQI        PSI       PHI                                   
REMARK 500    SER A   1      113.48    102.22                                   
DBREF  1PBX A    1   142  UNP    P80043   HBA_PAGBE        1    142             
DBREF  1PBX B    1   146  UNP    P80044   HBB_PAGBE        1    146             
SEQRES   1 A  143  ACE SER LEU SER ASP LYS ASP LYS ALA ALA VAL ARG ALA          
SEQRES   2 A  143  LEU TRP SER LYS ILE GLY LYS SER ALA ASP ALA ILE GLY          
SEQRES   3 A  143  ASN ASP ALA LEU SER ARG MET ILE VAL VAL TYR PRO GLN          
SEQRES   4 A  143  THR LYS THR TYR PHE SER HIS TRP PRO ASP VAL THR PRO          
SEQRES   5 A  143  GLY SER PRO HIS ILE LYS ALA HIS GLY LYS LYS VAL MET          
SEQRES   6 A  143  GLY GLY ILE ALA LEU ALA VAL SER LYS ILE ASP ASP LEU          
SEQRES   7 A  143  LYS THR GLY LEU MET GLU LEU SER GLU GLN HIS ALA TYR          
SEQRES   8 A  143  LYS LEU ARG VAL ASP PRO ALA ASN PHE LYS ILE LEU ASN          
SEQRES   9 A  143  HIS CYS ILE LEU VAL VAL ILE SER THR MET PHE PRO LYS          
SEQRES  10 A  143  GLU PHE THR PRO GLU ALA HIS VAL SER LEU ASP LYS PHE          
SEQRES  11 A  143  LEU SER GLY VAL ALA LEU ALA LEU ALA GLU ARG TYR ARG          
SEQRES   1 B  146  VAL GLU TRP THR ASP LYS GLU ARG SER ILE ILE SER ASP          
SEQRES   2 B  146  ILE PHE SER HIS MET ASP TYR ASP ASP ILE GLY PRO LYS          
SEQRES   3 B  146  ALA LEU SER ARG CYS LEU ILE VAL TYR PRO TRP THR GLN          
SEQRES   4 B  146  ARG HIS PHE SER GLY PHE GLY ASN LEU TYR ASN ALA GLU          
SEQRES   5 B  146  ALA ILE ILE GLY ASN ALA ASN VAL ALA ALA HIS GLY ILE          
SEQRES   6 B  146  LYS VAL LEU HIS GLY LEU ASP ARG GLY VAL LYS ASN MET          
SEQRES   7 B  146  ASP ASN ILE ALA ALA THR TYR ALA ASP LEU SER THR LEU          
SEQRES   8 B  146  HIS SER GLU LYS LEU HIS VAL ASP PRO ASP ASN PHE LYS          
SEQRES   9 B  146  LEU LEU SER ASP CYS ILE THR ILE VAL LEU ALA ALA LYS          
SEQRES  10 B  146  MET GLY HIS ALA PHE THR ALA GLU THR GLN GLY ALA PHE          
SEQRES  11 B  146  GLN LYS PHE LEU ALA VAL VAL VAL SER ALA LEU GLY LYS          
SEQRES  12 B  146  GLN TYR HIS                                                  
HET    ACE  A   0       3                                                       
HET    HEM  A 144      43                                                       
HET    CMO  A 145       2                                                       
HET    HEM  B 148      43                                                       
HET    CMO  B 149       2                                                       
HETNAM     ACE ACETYL GROUP                                                     
HETNAM     HEM PROTOPORPHYRIN IX CONTAINING FE                                  
HETNAM     CMO CARBON MONOXIDE                                                  
HETSYN     HEM HEME                                                             
FORMUL   1  ACE    C2 H4 O                                                      
FORMUL   3  HEM    2(C34 H32 FE N4 O4)                                          
FORMUL   4  CMO    2(C O)                                                       
FORMUL   7  HOH   *28(H2 O)                                                     
HELIX    1  AA SER A    3  GLY A   18  1                                  16    
HELIX    2  BA SER A   20  VAL A   35  1                                  16    
HELIX    3  CA TYR A   36  TYR A   42  1                                   7    
HELIX    4  EA SER A   53  SER A   72  1                                  20    
HELIX    5  FA LEU A   81  ALA A   89  1                                   9    
HELIX    6  GA ASP A   95  MET A  113  1                                  19    
HELIX    7  HA THR A  119  GLU A  139  1                                  21    
HELIX    8  AB THR B    4  MET B   18  1                                  15    
HELIX    9  BB ASP B   19  VAL B   34  1                                  16    
HELIX   10  CB TYR B   35  HIS B   41  1                                   7    
HELIX   11  DB ASN B   50  GLY B   56  1                                   7    
HELIX   12  EB ASN B   57  LYS B   76  1                                  20    
HELIX   13  FB TYR B   85  SER B   93  1                                   9    
HELIX   14  GB ASP B   99  LYS B  117  1                                  19    
HELIX   15  HB THR B  123  LYS B  143  1                                  21    
LINK         C   CMO A 145                FE   HEM A 144                        
LINK         C   CMO B 149                FE   HEM B 148                        
SITE     1 STA  2 HIS A  88  HEM A 144                                          
SITE     1 STB  2 HIS B  92  HEM B 148                                          
CRYST1   91.380   88.540   55.340  90.00  97.16  90.00 C 1 2 1       4          
ORIGX1      1.000000  0.000000  0.000000        0.00000                         
ORIGX2      0.000000  1.000000  0.000000        0.00000                         
ORIGX3      0.000000  0.000000  1.000000        0.00000                         
SCALE1      0.010943  0.000000  0.001375        0.00000                         
SCALE2      0.000000  0.011294  0.000000        0.00000                         
SCALE3      0.000000  0.000000  0.018212        0.00000