PDB entry 1ol0

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HEADER    IMMUNOGLOBULIN                          02-AUG-03   1OL0              
TITLE     CRYSTAL STRUCTURE OF A CAMELISED HUMAN VH                             
COMPND    MOL_ID: 1;                                                            
COMPND   2 MOLECULE: IMMUNOGLOBULIN G;                                          
COMPND   3 CHAIN: A, B;                                                         
COMPND   4 FRAGMENT: VARIABLE HEAVY DOMAIN;                                     
COMPND   5 ENGINEERED: YES                                                      
SOURCE    MOL_ID: 1;                                                            
SOURCE   2 ORGANISM_SCIENTIFIC: HOMO SAPIENS;                                   
SOURCE   3 ORGANISM_COMMON: HUMAN;                                              
SOURCE   4 EXPRESSION_SYSTEM: ESCHERICHIA COLI;                                 
SOURCE   5 EXPRESSION_SYSTEM_STRAIN: B834                                       
KEYWDS    IMMUNOGLOBULIN, CAMELISED VARIABLE HEAVY DOMAIN                       
EXPDTA    X-RAY DIFFRACTION                                                     
AUTHOR    T.DOTTORINI,C.K.VAUGHAN,M.A.WALSH,P.LOSURDO,M.SOLLAZZO                
REVDAT   1   22-JAN-04 1OL0    0                                                
JRNL        AUTH   T.DOTTORINI,C.K.VAUGHAN,M.A.WALSH,P.LOSURDO,                 
JRNL        AUTH 2 M.SOLLAZZO                                                   
JRNL        TITL   CRYSTAL STRUCTURE OF A HUMAN VH: REQUIREMENTS FOR            
JRNL        TITL 2 MAINTAINING A MONOMERIC FRAGMENT.                            
JRNL        REF    BIOCHEMISTRY                  V.  43   622 2004              
JRNL        REFN   ASTM BICHAW  US ISSN 0006-2960                               
REMARK   1                                                                      
REMARK   1 REFERENCE 1                                                          
REMARK   1  AUTH   S.DIMARCO,M.RIZZI,C.VOLPARI,M.A.WALSH,F.NARJES,              
REMARK   1  AUTH 2 S.COLARUSSO,R.DEFRANCESCO,V.G.MATASSA,M.SOLLAZZO             
REMARK   1  TITL   INHIBITION OF THE HEPATITIS C VIRUS NS3/4A                   
REMARK   1  TITL 2 PROTEASE. THE CRYSTAL STRUCTURES OF TWO                      
REMARK   1  TITL 3 PROTEASE-INHIBITOR COMPLEXES                                 
REMARK   1  REF    J.BIOL.CHEM.                  V. 275  7152 2000              
REMARK   1  REFN   ASTM JBCHA3  US ISSN 0021-9258                               
REMARK   1 REFERENCE 2                                                          
REMARK   1  AUTH   F.MARTIN,C.VOLPARI,C.STEINKUHLER,M.DIMASIBRUNETTI,           
REMARK   1  AUTH 2 G.BIASIOL,S.ALTAMURA,R.CORTESE,R.DEFRANCESCO,                
REMARK   1  AUTH 3 M.SOLLAZZO                                                   
REMARK   1  TITL   AFFINITY SELECTION OF A CAMELIZED V(H) DOMAIN                
REMARK   1  TITL 2 ANTIBODY INHIBITOR OF HEPATITIS C VIRUS NS3                  
REMARK   1  TITL 3 PROTEASE                                                     
REMARK   1  REF    PROTEIN ENG.                  V.  10   607 1997              
REMARK   1  REFN   ASTM PRENE9  UK ISSN 0269-2139                               
REMARK   1 REFERENCE 3                                                          
REMARK   1  AUTH   L.RIECHMANN                                                  
REMARK   1  TITL   REARRANGEMENT OF THE FORMER VL INTERFACE IN THE              
REMARK   1  TITL 2 SOLUTION STRUCTURE OF A CAMELISED, SINGLE ANTIBODY           
REMARK   1  TITL 3 VH DOMAIN                                                    
REMARK   1  REF    J.MOL.BIOL.                   V. 259   957 1996              
REMARK   1  REFN   ASTM JMOBAK  UK ISSN 0022-2836                               
REMARK   2                                                                      
REMARK   2 RESOLUTION. 1.80 ANGSTROMS.                                          
REMARK   3                                                                      
REMARK   3 REFINEMENT.                                                          
REMARK   3   PROGRAM     : REFMAC                                               
REMARK   3   AUTHORS     : MURSHUDOV,VAGIN,DODSON                               
REMARK   3                                                                      
REMARK   3  DATA USED IN REFINEMENT.                                            
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.80                           
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 25.24                          
REMARK   3   DATA CUTOFF            (SIGMA(F)) : NULL                           
REMARK   3   COMPLETENESS FOR RANGE        (%) : 95.0                           
REMARK   3   NUMBER OF REFLECTIONS             : 29892                          
REMARK   3                                                                      
REMARK   3  FIT TO DATA USED IN REFINEMENT.                                     
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT EXCEPT IN LAST       
REMARK   3                                      RO                              
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM                          
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.156                           
REMARK   3   R VALUE            (WORKING SET) : 0.154                           
REMARK   3   FREE R VALUE                     : 0.180                           
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.000                           
REMARK   3   FREE R VALUE TEST SET COUNT      : 1506                            
REMARK   3                                                                      
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.                    
REMARK   3   PROTEIN ATOMS            : 1818                                    
REMARK   3   NUCLEIC ACID ATOMS       : 0                                       
REMARK   3   HETEROGEN ATOMS          : 87                                      
REMARK   3   SOLVENT ATOMS            : NULL                                    
REMARK   3                                                                      
REMARK   3  B VALUES.                                                           
REMARK   3   FROM WILSON PLOT           (A**2) : NULL                           
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 17.34                          
REMARK   3   OVERALL ANISOTROPIC B VALUE.                                       
REMARK   3    B11 (A**2) : 0.00000                                              
REMARK   3    B22 (A**2) : 0.00000                                              
REMARK   3    B33 (A**2) : 0.00000                                              
REMARK   3    B12 (A**2) : 0.00000                                              
REMARK   3    B13 (A**2) : 0.00000                                              
REMARK   3    B23 (A**2) : 0.00000                                              
REMARK   3                                                                      
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.                                 
REMARK   3   ESU BASED ON R VALUE                            (A): NULL          
REMARK   3   ESU BASED ON FREE R VALUE                       (A): NULL          
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): NULL          
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): NULL          
REMARK   3                                                                      
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.                                   
REMARK   3   DISTANCE RESTRAINTS.                    RMS    SIGMA               
REMARK   3    BOND LENGTH                     (A) : NULL  ; NULL                
REMARK   3    ANGLE DISTANCE                  (A) : NULL  ; NULL                
REMARK   3    INTRAPLANAR 1-4 DISTANCE        (A) : NULL  ; NULL                
REMARK   3    H-BOND OR METAL COORDINATION    (A) : NULL  ; NULL                
REMARK   3                                                                      
REMARK   3   PLANE RESTRAINT                  (A) : NULL  ; NULL                
REMARK   3   CHIRAL-CENTER RESTRAINT       (A**3) : NULL  ; NULL                
REMARK   3                                                                      
REMARK   3   NON-BONDED CONTACT RESTRAINTS.                                     
REMARK   3    SINGLE TORSION                  (A) : NULL  ; NULL                
REMARK   3    MULTIPLE TORSION                (A) : NULL  ; NULL                
REMARK   3    H-BOND (X...Y)                  (A) : NULL  ; NULL                
REMARK   3    H-BOND (X-H...Y)                (A) : NULL  ; NULL                
REMARK   3                                                                      
REMARK   3   CONFORMATIONAL TORSION ANGLE RESTRAINTS.                           
REMARK   3    SPECIFIED                 (DEGREES) : NULL  ; NULL                
REMARK   3    PLANAR                    (DEGREES) : NULL  ; NULL                
REMARK   3    STAGGERED                 (DEGREES) : NULL  ; NULL                
REMARK   3    TRANSVERSE                (DEGREES) : NULL  ; NULL                
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA                
REMARK   3   MAIN-CHAIN BOND              (A**2) : NULL  ; NULL                 
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : NULL  ; NULL                 
REMARK   3   SIDE-CHAIN BOND              (A**2) : NULL  ; NULL                 
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : NULL  ; NULL                 
REMARK   3                                                                      
REMARK   3  OTHER REFINEMENT REMARKS: NULL                                      
REMARK   4                                                                      
REMARK   4 1OL0 COMPLIES WITH FORMAT V. 3.0, 1-DEC-2006                         
REMARK   4                                                                      
REMARK   4 THIS IS THE REMEDIATED VERSION OF THIS PDB ENTRY.                    
REMARK   4 REMEDIATED DATA FILE REVISION 3.101 (2007-05-31)                     
REMARK 100                                                                      
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY EBI  .                              
REMARK 100 THE EBI ID CODE IS  EBI-13218 .                                      
REMARK 200                                                                      
REMARK 200 EXPERIMENTAL DETAILS                                                 
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION                  
REMARK 200  DATE OF DATA COLLECTION        : 15-OCT-2000                        
REMARK 200  TEMPERATURE           (KELVIN) : 100.0                              
REMARK 200  PH                             : 7.50                               
REMARK 200  NUMBER OF CRYSTALS USED        : 1                                  
REMARK 200                                                                      
REMARK 200  SYNCHROTRON              (Y/N) : Y                                  
REMARK 200  RADIATION SOURCE               : ESRF                               
REMARK 200  BEAMLINE                       : ID14-2                             
REMARK 200  X-RAY GENERATOR MODEL          : NULL                               
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M                                  
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.933                              
REMARK 200  MONOCHROMATOR                  : DIAMON 100/GE 220                  
REMARK 200  OPTICS                         : TOROIDAL                           
REMARK 200                                                                      
REMARK 200  DETECTOR TYPE                  : CCD                                
REMARK 200  DETECTOR MANUFACTURER          : ADSC                               
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : DENZO                              
REMARK 200  DATA SCALING SOFTWARE          : SCALEPACK                          
REMARK 200                                                                      
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 29910                              
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.800                              
REMARK 200  RESOLUTION RANGE LOW       (A) : 30.000                             
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : -3.000                             
REMARK 200                                                                      
REMARK 200 OVERALL.                                                             
REMARK 200  COMPLETENESS FOR RANGE     (%) : 94.9                               
REMARK 200  DATA REDUNDANCY                : 2.300                              
REMARK 200  R MERGE                    (I) : 0.05100                            
REMARK 200  R SYM                      (I) : NULL                               
REMARK 200   FOR THE DATA SET  : 17.2000                            
REMARK 200                                                                      
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.                                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.80                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.83                     
REMARK 200  COMPLETENESS FOR SHELL     (%) : 71.5                               
REMARK 200  DATA REDUNDANCY IN SHELL       : 2.00                               
REMARK 200  R MERGE FOR SHELL          (I) : 0.22700                            
REMARK 200  R SYM FOR SHELL            (I) : NULL                               
REMARK 200   FOR SHELL         : 1.500                              
REMARK 200                                                                      
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH                              
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT        
REMARK 200 SOFTWARE USED: AMORE                                                 
REMARK 200 STARTING MODEL: PDB ENTRY 1HCV                                       
REMARK 200                                                                      
REMARK 200 REMARK: NULL                                                         
REMARK 280                                                                      
REMARK 280 CRYSTAL                                                              
REMARK 280 SOLVENT CONTENT, VS   (%): 59.50                                     
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 3.10                     
REMARK 280                                                                      
REMARK 280 CRYSTALLIZATION CONDITIONS: PH 7.5                                   
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY                                            
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 32 2 1                         
REMARK 290                                                                      
REMARK 290      SYMOP   SYMMETRY                                                
REMARK 290     NNNMMM   OPERATOR                                                
REMARK 290       1555   X,Y,Z                                                   
REMARK 290       2555   -Y,X-Y,2/3+Z                                            
REMARK 290       3555   -X+Y,-X,1/3+Z                                           
REMARK 290       4555   Y,X,-Z                                                  
REMARK 290       5555   X-Y,-Y,1/3-Z                                            
REMARK 290       6555   -X,-X+Y,2/3-Z                                           
REMARK 290                                                                      
REMARK 290     WHERE NNN -> OPERATOR NUMBER                                     
REMARK 290           MMM -> TRANSLATION VECTOR                                  
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS                            
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM             
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY                
REMARK 290 RELATED MOLECULES.                                                   
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   2 -0.500000 -0.866025  0.000000        0.00000            
REMARK 290   SMTRY2   2  0.866025 -0.500000  0.000000        0.00000            
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       67.85333            
REMARK 290   SMTRY1   3 -0.500000  0.866025  0.000000        0.00000            
REMARK 290   SMTRY2   3 -0.866025 -0.500000  0.000000        0.00000            
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000       33.92667            
REMARK 290   SMTRY1   4 -0.500000  0.866025  0.000000        0.00000            
REMARK 290   SMTRY2   4  0.866025  0.500000  0.000000        0.00000            
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000            
REMARK 290   SMTRY1   5  1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   5  0.000000 -1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   5  0.000000  0.000000 -1.000000       33.92667            
REMARK 290   SMTRY1   6 -0.500000 -0.866025  0.000000        0.00000            
REMARK 290   SMTRY2   6 -0.866025  0.500000  0.000000        0.00000            
REMARK 290   SMTRY3   6  0.000000  0.000000 -1.000000       67.85333            
REMARK 290                                                                      
REMARK 290 REMARK: NULL                                                         
REMARK 300                                                                      
REMARK 300 BIOMOLECULE: 1                                                       
REMARK 300 THIS ENTRY CONTAINS THE CRYSTALLOGRAPHIC ASYMMETRIC UNIT             
REMARK 300 WHICH CONSISTS OF 2 CHAIN(S). SEE REMARK 350 FOR                     
REMARK 300 INFORMATION ON GENERATING THE BIOLOGICAL MOLECULE(S).                
REMARK 350                                                                      
REMARK 350 GENERATING THE BIOMOLECULE                                           
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN           
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE                
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS          
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND                          
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.                               
REMARK 350                                                                      
REMARK 350 BIOMOLECULE: 1                                                       
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B, Y, Z                            
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT                     
REMARK 500                                                                      
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.                            
REMARK 500                                                                      
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI                             
REMARK 500   O    HOH Y    57     O    HOH Y   147              2.08            
REMARK 500   O    HOH Y    92     O    HOH Y    99              2.10            
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: CLOSE CONTACTS                                             
REMARK 500                                                                      
REMARK 500 THE FOLLOWING ATOMS THAT ARE RELATED BY CRYSTALLOGRAPHIC             
REMARK 500 SYMMETRY ARE IN CLOSE CONTACT.  AN ATOM LOCATED WITHIN 0.15          
REMARK 500 ANGSTROMS OF A SYMMETRY RELATED ATOM IS ASSUMED TO BE ON A           
REMARK 500 SPECIAL POSITION AND IS, THEREFORE, LISTED IN REMARK 375             
REMARK 500 INSTEAD OF REMARK 500.  ATOMS WITH NON-BLANK ALTERNATE               
REMARK 500 LOCATION INDICATORS ARE NOT INCLUDED IN THE CALCULATIONS.            
REMARK 500                                                                      
REMARK 500 DISTANCE CUTOFF:                                                     
REMARK 500 2.2 ANGSTROMS FOR CONTACTS NOT INVOLVING HYDROGEN ATOMS              
REMARK 500 1.6 ANGSTROMS FOR CONTACTS INVOLVING HYDROGEN ATOMS                  
REMARK 500                                                                      
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI  SSYMOP   DISTANCE          
REMARK 500   O    GLY B     9     NZ   LYS A    43     2654     2.13            
REMARK 800                                                                      
REMARK 800 SITE                                                                 
REMARK 800 SITE_IDENTIFIER: AC1                                                 
REMARK 800 SITE_DESCRIPTION: GOL BINDING SITE FOR CHAIN A                       
DBREF  1OL0 A    1   121  PDB    1OL0     1OL0             1    121             
DBREF  1OL0 B    1   121  PDB    1OL0     1OL0             1    121             
SEQRES   1 A  121  GLN VAL GLN LEU VAL GLU SER GLY GLY GLY LEU VAL GLN          
SEQRES   2 A  121  PRO GLY GLY SER LEU ARG LEU SER CYS ALA ALA SER GLY          
SEQRES   3 A  121  PHE THR PHE SER SER TYR ALA MSE SER TRP PHE ARG GLN          
SEQRES   4 A  121  ALA PRO GLY LYS GLU ARG GLU ILE VAL SER ALA VAL SER          
SEQRES   5 A  121  GLY SER GLY GLY SER THR TYR TYR ALA ASP SER VAL ARG          
SEQRES   6 A  121  GLY ARG PHE THR ILE SER ARG ASP ASN SER LYS ASN THR          
SEQRES   7 A  121  LEU TYR LEU GLN MSE ASN SER LEU ARG ALA GLU ASP THR          
SEQRES   8 A  121  ALA VAL TYR TYR CYS ALA ARG GLU PRO ARG ILE PRO ARG          
SEQRES   9 A  121  PRO PRO SER PHE ASP TYR TRP GLY GLN GLY THR LEU VAL          
SEQRES  10 A  121  THR VAL SER SER                                              
SEQRES   1 B  121  GLN VAL GLN LEU VAL GLU SER GLY GLY GLY LEU VAL GLN          
SEQRES   2 B  121  PRO GLY GLY SER LEU ARG LEU SER CYS ALA ALA SER GLY          
SEQRES   3 B  121  PHE THR PHE SER SER TYR ALA MSE SER TRP PHE ARG GLN          
SEQRES   4 B  121  ALA PRO GLY LYS GLU ARG GLU ILE VAL SER ALA VAL SER          
SEQRES   5 B  121  GLY SER GLY GLY SER THR TYR TYR ALA ASP SER VAL ARG          
SEQRES   6 B  121  GLY ARG PHE THR ILE SER ARG ASP ASN SER LYS ASN THR          
SEQRES   7 B  121  LEU TYR LEU GLN MSE ASN SER LEU ARG ALA GLU ASP THR          
SEQRES   8 B  121  ALA VAL TYR TYR CYS ALA ARG GLU PRO ARG ILE PRO ARG          
SEQRES   9 B  121  PRO PRO SER PHE ASP TYR TRP GLY GLN GLY THR LEU VAL          
SEQRES  10 B  121  THR VAL SER SER                                              
MODRES 1OL0 MSE A   34  MET  SELENOMETHIONINE                                   
MODRES 1OL0 MSE A   83  MET  SELENOMETHIONINE                                   
MODRES 1OL0 MSE B   34  MET  SELENOMETHIONINE                                   
MODRES 1OL0 MSE B   83  MET  SELENOMETHIONINE                                   
HET    MSE  A  34       8                                                       
HET    MSE  A  83       8                                                       
HET    MSE  B  34       8                                                       
HET    MSE  B  83       8                                                       
HET    SO4  A1122       5                                                       
HET    SO4  B1122       5                                                       
HET    SO4  B1123       5                                                       
HET    SO4  B1124       5                                                       
HET    SO4  B1125       5                                                       
HET    GOL  A1123       6                                                       
HET    GOL  A1124       6                                                       
HET    GOL  A1125       6                                                       
HET    GOL  A1126       6                                                       
HET    GOL  B1126       6                                                       
HETNAM     MSE SELENOMETHIONINE                                                 
HETNAM     SO4 SULFATE ION                                                      
HETNAM     GOL GLYCEROL                                                         
FORMUL   1  MSE    4(C5 H11 N O2 SE)                                            
FORMUL   3  SO4    5(O4 S 2-)                                                   
FORMUL   8  GOL    5(C3 H8 O3)                                                  
FORMUL  13  HOH   *388(H2 O)                                                    
HELIX    1   1 THR A   28  TYR A   32  5                                   5    
HELIX    2   2 ASP A   62  ARG A   65  5                                   4    
HELIX    3   3 ARG A   87  THR A   91  5                                   5    
HELIX    4   4 PRO A  105  ASP A  109  5                                   5    
HELIX    5   5 THR B   28  TYR B   32  5                                   5    
HELIX    6   6 ASP B   62  ARG B   65  5                                   4    
HELIX    7   7 ARG B   87  THR B   91  5                                   5    
HELIX    8   8 PRO B  105  ASP B  109  5                                   5    
SHEET    1  AA 4 GLN A   3  GLY A   8  0                                        
SHEET    2  AA 4 LEU A  18  SER A  25 -1  O  SER A  21   N  SER A   7           
SHEET    3  AA 4 THR A  78  MSE A  83 -1  O  LEU A  79   N  CYS A  22           
SHEET    4  AA 4 PHE A  68  ASP A  73 -1  O  THR A  69   N  GLN A  82           
SHEET    1  AB 6 GLY A  10  VAL A  12  0                                        
SHEET    2  AB 6 THR A 115  VAL A 119  1  O  LEU A 116   N  GLY A  10           
SHEET    3  AB 6 ALA A  92  ARG A  98 -1  O  ALA A  92   N  VAL A 117           
SHEET    4  AB 6 MSE A  34  GLN A  39 -1  O  SER A  35   N  ALA A  97           
SHEET    5  AB 6 GLU A  46  VAL A  51 -1  O  GLU A  46   N  ARG A  38           
SHEET    6  AB 6 THR A  58  TYR A  60 -1  O  TYR A  59   N  ALA A  50           
SHEET    1  AC 4 GLY A  10  VAL A  12  0                                        
SHEET    2  AC 4 THR A 115  VAL A 119  1  O  LEU A 116   N  GLY A  10           
SHEET    3  AC 4 ALA A  92  ARG A  98 -1  O  ALA A  92   N  VAL A 117           
SHEET    4  AC 4 TYR A 110  TRP A 111 -1  O  TYR A 110   N  ARG A  98           
SHEET    1  BA 4 GLN B   3  GLY B   8  0                                        
SHEET    2  BA 4 LEU B  18  SER B  25 -1  O  SER B  21   N  SER B   7           
SHEET    3  BA 4 THR B  78  MSE B  83 -1  O  LEU B  79   N  CYS B  22           
SHEET    4  BA 4 PHE B  68  ASP B  73 -1  O  THR B  69   N  GLN B  82           
SHEET    1  BB 6 GLY B  10  VAL B  12  0                                        
SHEET    2  BB 6 THR B 115  VAL B 119  1  O  LEU B 116   N  GLY B  10           
SHEET    3  BB 6 ALA B  92  ARG B  98 -1  O  ALA B  92   N  VAL B 117           
SHEET    4  BB 6 MSE B  34  GLN B  39 -1  O  SER B  35   N  ALA B  97           
SHEET    5  BB 6 GLU B  46  VAL B  51 -1  O  GLU B  46   N  ARG B  38           
SHEET    6  BB 6 THR B  58  TYR B  60 -1  O  TYR B  59   N  ALA B  50           
SHEET    1  BC 4 GLY B  10  VAL B  12  0                                        
SHEET    2  BC 4 THR B 115  VAL B 119  1  O  LEU B 116   N  GLY B  10           
SHEET    3  BC 4 ALA B  92  ARG B  98 -1  O  ALA B  92   N  VAL B 117           
SHEET    4  BC 4 TYR B 110  TRP B 111 -1  O  TYR B 110   N  ARG B  98           
SSBOND   1 CYS A   22    CYS A   96                                             
SSBOND   2 CYS B   22    CYS B   96                                             
LINK         C   ALA A  33                 N   MSE A  34                        
LINK         C   MSE A  34                 N   SER A  35                        
LINK         C   GLN A  82                 N   MSE A  83                        
LINK         C   MSE A  83                 N   ASN A  84                        
LINK         C   ALA B  33                 N   MSE B  34                        
LINK         C   MSE B  34                 N   SER B  35                        
LINK         C   GLN B  82                 N   MSE B  83                        
LINK         C   MSE B  83                 N   ASN B  84                        
CISPEP   1 ILE A  102    PRO A  103          0        -1.63                     
CISPEP   2 ILE B  102    PRO B  103          0        -1.83                     
SITE     1 AC1 10 GLY A   8  GLU B  44  ARG B  45  HOH Y  74                    
SITE     2 AC1 10 HOH Y 117  HOH Y 180  HOH Y 181  HOH Y 182                    
SITE     3 AC1 10 HOH Y 183  HOH Y 184                                          
SITE     1 AC2  4 ARG A  65  ARG B 104  HOH Y 186  HOH Z 105                    
SITE     1 AC3  5 PHE B  27  THR B  28  TYR B  32  ARG B 101                    
SITE     2 AC3  5 HOH Y  49                                                     
SITE     1 AC4  4 ARG A 104  ARG B  65  HOH Y  91  HOH Y 187                    
SITE     1 AC5  6 PHE A  27  THR A  28  TYR A  32  ARG A 101                    
SITE     2 AC5  6 HOH Z  47  HOH Z 197                                          
SITE     1 AC6  7 THR A  58  TYR A  60  THR B  58  TYR B  60                    
SITE     2 AC6  7 HOH Y 105  HOH Z 130  HOH Z 198                               
SITE     1 AC7  5 GLU B  46  ASP B  62  SER B  63  HOH Y 188                    
SITE     2 AC7  5 HOH Y 189                                                     
SITE     1 AC8  6 GLN A  13  SER A 120  SER A 121  PRO B  41                    
SITE     2 AC8  6 HOH Z  16  HOH Z 199                                          
SITE     1 AC9  5 ARG A  19  ARG B  19  HOH Y 118  HOH Z  34                    
SITE     2 AC9  5 HOH Z 147                                                     
SITE     1 BC1  5 PHE A  27  THR A  28  PHE A  29  ASN A  77                    
SITE     2 BC1  5 HOH Z  49                                                     
CRYST1   75.300   75.300  101.780  90.00  90.00 120.00 P 32 2 1     12          
ORIGX1      1.000000  0.000000  0.000000        0.00000                         
ORIGX2      0.000000  1.000000  0.000000        0.00000                         
ORIGX3      0.000000  0.000000  1.000000        0.00000                         
SCALE1      0.013280  0.007667  0.000000        0.00000                         
SCALE2      0.000000  0.015335  0.000000        0.00000                         
SCALE3      0.000000  0.000000  0.009825        0.00000                         
MTRIX1   1 -0.999943 -0.009568 -0.004770       95.64020    1                    
MTRIX2   1  0.001752 -0.586769  0.809753       37.70880    1                    
MTRIX3   1 -0.010547  0.809698  0.586752      -18.93450    1