PDB entry 1nsd

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HEADER    HYDROLASE(O-GLYCOSYL)                   24-MAY-93   1NSD              
TITLE     INFLUENZA B VIRUS NEURAMINIDASE CAN SYNTHESIZE ITS OWN                
TITLE    2 INHIBITOR                                                            
COMPND    MOL_ID: 1;                                                            
COMPND   2 MOLECULE: NEURAMINIDASE;                                             
COMPND   3 CHAIN: A, B;                                                         
COMPND   4 EC: 3.2.1.18;                                                        
COMPND   5 ENGINEERED: YES                                                      
SOURCE    MOL_ID: 1;                                                            
SOURCE   2 ORGANISM_SCIENTIFIC: INFLUENZA B VIRUS;                              
SOURCE   3 STRAIN: B/BEIJING/1/87                                               
KEYWDS    HYDROLASE(O-GLYCOSYL)                                                 
EXPDTA    X-RAY DIFFRACTION                                                     
AUTHOR    W.P.BURMEISTER,R.W.H.RUIGROK,S.CUSACK                                 
REVDAT   2   01-APR-03 1NSD    1       JRNL                                     
REVDAT   1   31-OCT-93 1NSD    0                                                
JRNL        AUTH   W.P.BURMEISTER,B.HENRISSAT,C.BOSSO,S.CUSACK,                 
JRNL        AUTH 2 R.W.RUIGROK                                                  
JRNL        TITL   INFLUENZA B VIRUS NEURAMINIDASE CAN SYNTHESIZE ITS           
JRNL        TITL 2 OWN INHIBITOR.                                               
JRNL        REF    STRUCTURE                     V.   1    19 1993              
JRNL        REFN   ASTM STRUE6  UK ISSN 0969-2126                               
REMARK   1                                                                      
REMARK   1 REFERENCE 1                                                          
REMARK   1  AUTH   W.P.BURMEISTER,R.W.H.RUIGROK,S.CUSACK                        
REMARK   1  TITL   THE 2.2 ANGSTROMS RESOLUTION CRYSTAL STRUCTURE OF            
REMARK   1  TITL 2 INFLUENZA B NEURAMINIDASE AND ITS COMPLEX WITH               
REMARK   1  TITL 3 SIALIC ACID                                                  
REMARK   1  REF    EMBO J.                       V.  11    49 1992              
REMARK   1  REFN   ASTM EMJODG  UK ISSN 0261-4189                               
REMARK   1 REFERENCE 2                                                          
REMARK   1  AUTH   W.P.BURMEISTER,R.S.DANIELS,S.DAYAN,J.GAGNON,                 
REMARK   1  AUTH 2 S.CUSACK,R.W.H.RUIGROK                                       
REMARK   1  TITL   SEQUENCE AND CRYSTALLIZATION OF INFLUENZA VIRUS              
REMARK   1  TITL 2 B(SLASH)BEIJING(SLASH)1(SLASH)87 NEURAMINIDASE               
REMARK   1  REF    VIROLOGY                      V. 180   266 1991              
REMARK   1  REFN   ASTM VIRLAX  US ISSN 0042-6822                               
REMARK   2                                                                      
REMARK   2 RESOLUTION. 1.80 ANGSTROMS.                                          
REMARK   3                                                                      
REMARK   3 REFINEMENT.                                                          
REMARK   3   PROGRAM     : X-PLOR                                               
REMARK   3   AUTHORS     : BRUNGER                                              
REMARK   3                                                                      
REMARK   3  DATA USED IN REFINEMENT.                                            
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.80                           
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : NULL                           
REMARK   3   DATA CUTOFF            (SIGMA(F)) : NULL                           
REMARK   3   DATA CUTOFF HIGH         (ABS(F)) : NULL                           
REMARK   3   DATA CUTOFF LOW          (ABS(F)) : NULL                           
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : NULL                           
REMARK   3   NUMBER OF REFLECTIONS             : NULL                           
REMARK   3                                                                      
REMARK   3  FIT TO DATA USED IN REFINEMENT.                                     
REMARK   3   CROSS-VALIDATION METHOD          : NULL                            
REMARK   3   FREE R VALUE TEST SET SELECTION  : NULL                            
REMARK   3   R VALUE            (WORKING SET) : 0.169                           
REMARK   3   FREE R VALUE                     : NULL                            
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : NULL                            
REMARK   3   FREE R VALUE TEST SET COUNT      : NULL                            
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : NULL                            
REMARK   3                                                                      
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.                                  
REMARK   3   TOTAL NUMBER OF BINS USED           : NULL                         
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : NULL                         
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : NULL                         
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : NULL                         
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : NULL                         
REMARK   3   BIN R VALUE           (WORKING SET) : NULL                         
REMARK   3   BIN FREE R VALUE                    : NULL                         
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : NULL                         
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : NULL                         
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : NULL                         
REMARK   3                                                                      
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.                    
REMARK   3   PROTEIN ATOMS            : 7474                                    
REMARK   3   NUCLEIC ACID ATOMS       : 0                                       
REMARK   3   HETEROGEN ATOMS          : 131                                     
REMARK   3   SOLVENT ATOMS            : 1510                                    
REMARK   3                                                                      
REMARK   3  B VALUES.                                                           
REMARK   3   FROM WILSON PLOT           (A**2) : NULL                           
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : NULL                           
REMARK   3   OVERALL ANISOTROPIC B VALUE.                                       
REMARK   3    B11 (A**2) : NULL                                                 
REMARK   3    B22 (A**2) : NULL                                                 
REMARK   3    B33 (A**2) : NULL                                                 
REMARK   3    B12 (A**2) : NULL                                                 
REMARK   3    B13 (A**2) : NULL                                                 
REMARK   3    B23 (A**2) : NULL                                                 
REMARK   3                                                                      
REMARK   3  ESTIMATED COORDINATE ERROR.                                         
REMARK   3   ESD FROM LUZZATI PLOT        (A) : NULL                            
REMARK   3   ESD FROM SIGMAA              (A) : NULL                            
REMARK   3   LOW RESOLUTION CUTOFF        (A) : NULL                            
REMARK   3                                                                      
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.                         
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : NULL                            
REMARK   3   ESD FROM C-V SIGMAA          (A) : NULL                            
REMARK   3                                                                      
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.                                   
REMARK   3   BOND LENGTHS                 (A) : 0.013                           
REMARK   3   BOND ANGLES            (DEGREES) : 2.86                            
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : NULL                            
REMARK   3   IMPROPER ANGLES        (DEGREES) : NULL                            
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL MODEL : NULL                                      
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA                
REMARK   3   MAIN-CHAIN BOND              (A**2) : NULL  ; NULL                 
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : NULL  ; NULL                 
REMARK   3   SIDE-CHAIN BOND              (A**2) : NULL  ; NULL                 
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : NULL  ; NULL                 
REMARK   3                                                                      
REMARK   3  NCS MODEL : NULL                                                    
REMARK   3                                                                      
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT          
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL                 
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL                 
REMARK   3                                                                      
REMARK   3  PARAMETER FILE  1  : NULL                                           
REMARK   3  TOPOLOGY FILE  1   : NULL                                           
REMARK   3                                                                      
REMARK   3  OTHER REFINEMENT REMARKS: NULL                                      
REMARK   4                                                                      
REMARK   4 1NSD COMPLIES WITH FORMAT V. 3.0, 1-DEC-2006                         
REMARK   4                                                                      
REMARK   4 THIS IS THE REMEDIATED VERSION OF THIS PDB ENTRY.                    
REMARK   4 REMEDIATED DATA FILE REVISION 3.100 (2007-03-17)                     
REMARK 200                                                                      
REMARK 200 EXPERIMENTAL DETAILS                                                 
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION                  
REMARK 200  DATE OF DATA COLLECTION        : NULL                               
REMARK 200  TEMPERATURE           (KELVIN) : NULL                               
REMARK 200  PH                             : NULL                               
REMARK 200  NUMBER OF CRYSTALS USED        : NULL                               
REMARK 200                                                                      
REMARK 200  SYNCHROTRON              (Y/N) : NULL                               
REMARK 200  RADIATION SOURCE               : NULL                               
REMARK 200  BEAMLINE                       : NULL                               
REMARK 200  X-RAY GENERATOR MODEL          : NULL                               
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : NULL                               
REMARK 200  WAVELENGTH OR RANGE        (A) : NULL                               
REMARK 200  MONOCHROMATOR                  : NULL                               
REMARK 200  OPTICS                         : NULL                               
REMARK 200                                                                      
REMARK 200  DETECTOR TYPE                  : NULL                               
REMARK 200  DETECTOR MANUFACTURER          : NULL                               
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : NULL                               
REMARK 200  DATA SCALING SOFTWARE          : NULL                               
REMARK 200                                                                      
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : NULL                               
REMARK 200  RESOLUTION RANGE HIGH      (A) : NULL                               
REMARK 200  RESOLUTION RANGE LOW       (A) : NULL                               
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL                               
REMARK 200                                                                      
REMARK 200 OVERALL.                                                             
REMARK 200  COMPLETENESS FOR RANGE     (%) : NULL                               
REMARK 200  DATA REDUNDANCY                : NULL                               
REMARK 200  R MERGE                    (I) : NULL                               
REMARK 200  R SYM                      (I) : NULL                               
REMARK 200   FOR THE DATA SET  : NULL                               
REMARK 200                                                                      
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.                                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : NULL                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : NULL                     
REMARK 200  COMPLETENESS FOR SHELL     (%) : NULL                               
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL                               
REMARK 200  R MERGE FOR SHELL          (I) : NULL                               
REMARK 200  R SYM FOR SHELL            (I) : NULL                               
REMARK 200   FOR SHELL         : NULL                               
REMARK 200                                                                      
REMARK 200 DIFFRACTION PROTOCOL: NULL                                           
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: NULL                         
REMARK 200 SOFTWARE USED: X-PLOR                                                
REMARK 200 STARTING MODEL: NULL                                                 
REMARK 200                                                                      
REMARK 200 REMARK: NULL                                                         
REMARK 280                                                                      
REMARK 280 CRYSTAL                                                              
REMARK 280 SOLVENT CONTENT, VS   (%): 57.95                                     
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.92                     
REMARK 280                                                                      
REMARK 280 CRYSTALLIZATION CONDITIONS: NULL                                     
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY                                            
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 31 2 1                         
REMARK 290                                                                      
REMARK 290      SYMOP   SYMMETRY                                                
REMARK 290     NNNMMM   OPERATOR                                                
REMARK 290       1555   X,Y,Z                                                   
REMARK 290       2555   -Y,X-Y,1/3+Z                                            
REMARK 290       3555   -X+Y,-X,2/3+Z                                           
REMARK 290       4555   Y,X,-Z                                                  
REMARK 290       5555   X-Y,-Y,2/3-Z                                            
REMARK 290       6555   -X,-X+Y,1/3-Z                                           
REMARK 290                                                                      
REMARK 290     WHERE NNN -> OPERATOR NUMBER                                     
REMARK 290           MMM -> TRANSLATION VECTOR                                  
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS                            
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM             
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY                
REMARK 290 RELATED MOLECULES.                                                   
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   2 -0.500000 -0.866025  0.000000        0.00000            
REMARK 290   SMTRY2   2  0.866025 -0.500000  0.000000        0.00000            
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       74.26667            
REMARK 290   SMTRY1   3 -0.500000  0.866025  0.000000        0.00000            
REMARK 290   SMTRY2   3 -0.866025 -0.500000  0.000000        0.00000            
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000      148.53333            
REMARK 290   SMTRY1   4 -0.500000  0.866025  0.000000        0.00000            
REMARK 290   SMTRY2   4  0.866025  0.500000  0.000000        0.00000            
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000            
REMARK 290   SMTRY1   5  1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   5  0.000000 -1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   5  0.000000  0.000000 -1.000000      148.53333            
REMARK 290   SMTRY1   6 -0.500000 -0.866025  0.000000        0.00000            
REMARK 290   SMTRY2   6 -0.866025  0.500000  0.000000        0.00000            
REMARK 290   SMTRY3   6  0.000000  0.000000 -1.000000       74.26667            
REMARK 290                                                                      
REMARK 290 REMARK: NULL                                                         
REMARK 300                                                                      
REMARK 300 BIOMOLECULE: 1                                                       
REMARK 300 THIS ENTRY CONTAINS THE CRYSTALLOGRAPHIC ASYMMETRIC UNIT             
REMARK 300 WHICH CONSISTS OF 2 CHAIN(S). SEE REMARK 350 FOR                     
REMARK 300 INFORMATION ON GENERATING THE BIOLOGICAL MOLECULE(S).                
REMARK 350                                                                      
REMARK 350 GENERATING THE BIOMOLECULE                                           
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN           
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE                
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS          
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND                          
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.                               
REMARK 350                                                                      
REMARK 350 BIOMOLECULE: 1                                                       
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B                                  
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 350   BIOMT1   2 -0.500000  0.866025  0.000000        0.00000            
REMARK 350   BIOMT2   2  0.866025  0.500000  0.000000        0.00000            
REMARK 350   BIOMT3   2  0.000000  0.000000 -1.000000        0.00000            
REMARK 375                                                                      
REMARK 375 SPECIAL POSITION                                                     
REMARK 375 THE FOLLOWING ATOMS ARE FOUND TO BE WITHIN 0.15 ANGSTROMS            
REMARK 375 OF A SYMMETRY RELATED ATOM AND ARE ASSUMED TO BE ON SPECIAL          
REMARK 375 POSITIONS.                                                           
REMARK 375                                                                      
REMARK 375 ATOM RES CSSEQI                                                      
REMARK 375 CA    CA   470   LIES ON A SPECIAL POSITION.                         
REMARK 375      HOH   179   LIES ON A SPECIAL POSITION.                         
REMARK 375      HOH   279   LIES ON A SPECIAL POSITION.                         
REMARK 375      HOH   384   LIES ON A SPECIAL POSITION.                         
REMARK 375      HOH   452   LIES ON A SPECIAL POSITION.                         
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES                                       
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)              
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3                                     
REMARK 500    ILE A 163   N   -  CA  -  C   ANGL. DEV. =-17.2 DEGREES           
REMARK 525                                                                      
REMARK 525 SOLVENT                                                              
REMARK 525 THE FOLLOWING SOLVENT MOLECULES LIE FARTHER THAN EXPECTED            
REMARK 525 FROM THE PROTEIN OR NUCLEIC ACID MOLECULE AND MAY BE                 
REMARK 525 ASSOCIATED WITH A SYMMETRY RELATED MOLECULE (M=MODEL                 
REMARK 525 NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE          
REMARK 525 NUMBER; I=INSERTION CODE):                                           
REMARK 525                                                                      
REMARK 525  M RES CSSEQI                                                        
REMARK 525    HOH   272        DISTANCE =  6.32 ANGSTROMS                       
REMARK 525    HOH   488        DISTANCE =  5.63 ANGSTROMS                       
DBREF  1NSD A   76   465  UNP    P27907   NRAM_INBBE      76    465             
DBREF  1NSD B   76   465  UNP    P27907   NRAM_INBBE      76    465             
SEQRES   1 A  390  GLU PRO GLU TRP THR TYR PRO ARG LEU SER CYS GLN GLY          
SEQRES   2 A  390  SER THR PHE GLN LYS ALA LEU LEU ILE SER PRO HIS ARG          
SEQRES   3 A  390  PHE GLY GLU ALA ARG GLY ASN SER ALA PRO LEU ILE ILE          
SEQRES   4 A  390  ARG GLU PRO PHE ILE ALA CYS GLY PRO LYS GLU CYS LYS          
SEQRES   5 A  390  HIS PHE ALA LEU THR HIS TYR ALA ALA GLN PRO GLY GLY          
SEQRES   6 A  390  TYR TYR ASN GLY THR ARG GLU ASP ARG ASN LYS LEU ARG          
SEQRES   7 A  390  HIS LEU ILE SER VAL LYS LEU GLY LYS ILE PRO THR VAL          
SEQRES   8 A  390  GLU ASN SER ILE PHE HIS MET ALA ALA TRP SER GLY SER          
SEQRES   9 A  390  ALA CYS HIS ASP GLY ARG GLU TRP THR TYR ILE GLY VAL          
SEQRES  10 A  390  ASP GLY PRO ASP SER ASN ALA LEU ILE LYS ILE LYS TYR          
SEQRES  11 A  390  GLY GLU ALA TYR THR ASP THR TYR HIS SER TYR ALA ASN          
SEQRES  12 A  390  ASN ILE LEU ARG THR GLN GLU SER ALA CYS ASN CYS ILE          
SEQRES  13 A  390  GLY GLY ASP CYS TYR LEU MET ILE THR ASP GLY SER ALA          
SEQRES  14 A  390  SER GLY ILE SER LYS CYS ARG PHE LEU LYS ILE ARG GLU          
SEQRES  15 A  390  GLY ARG ILE ILE LYS GLU ILE PHE PRO THR GLY ARG VAL          
SEQRES  16 A  390  GLU HIS THR GLU GLU CYS THR CYS GLY PHE ALA SER ASN          
SEQRES  17 A  390  LYS THR ILE GLU CYS ALA CYS ARG ASP ASN SER TYR THR          
SEQRES  18 A  390  ALA LYS ARG PRO PHE VAL LYS LEU ASN VAL GLU THR ASP          
SEQRES  19 A  390  THR ALA GLU ILE ARG LEU MET CYS THR GLU THR TYR LEU          
SEQRES  20 A  390  ASP THR PRO ARG PRO ASP ASP GLY SER ILE THR GLY PRO          
SEQRES  21 A  390  CYS GLU SER ASN GLY ASP LYS GLY ARG GLY GLY ILE LYS          
SEQRES  22 A  390  GLY GLY PHE VAL HIS GLN ARG MET ALA SER LYS ILE GLY          
SEQRES  23 A  390  ARG TRP TYR SER ARG THR MET SER LYS THR GLU ARG MET          
SEQRES  24 A  390  GLY MET GLU LEU TYR VAL ARG TYR ASP GLY ASP PRO TRP          
SEQRES  25 A  390  THR ASP SER ASP ALA LEU ALA HIS SER GLY VAL MET VAL          
SEQRES  26 A  390  SER MET LYS GLU PRO GLY TRP TYR SER PHE GLY PHE GLU          
SEQRES  27 A  390  ILE LYS ASP LYS LYS CYS ASP VAL PRO CYS ILE GLY ILE          
SEQRES  28 A  390  GLU MET VAL HIS ASP GLY GLY LYS LYS THR TRP HIS SER          
SEQRES  29 A  390  ALA ALA THR ALA ILE TYR CYS LEU MET GLY SER GLY GLN          
SEQRES  30 A  390  LEU LEU TRP ASP THR VAL THR GLY VAL ASP MET ALA LEU          
SEQRES   1 B  390  GLU PRO GLU TRP THR TYR PRO ARG LEU SER CYS GLN GLY          
SEQRES   2 B  390  SER THR PHE GLN LYS ALA LEU LEU ILE SER PRO HIS ARG          
SEQRES   3 B  390  PHE GLY GLU ALA ARG GLY ASN SER ALA PRO LEU ILE ILE          
SEQRES   4 B  390  ARG GLU PRO PHE ILE ALA CYS GLY PRO LYS GLU CYS LYS          
SEQRES   5 B  390  HIS PHE ALA LEU THR HIS TYR ALA ALA GLN PRO GLY GLY          
SEQRES   6 B  390  TYR TYR ASN GLY THR ARG GLU ASP ARG ASN LYS LEU ARG          
SEQRES   7 B  390  HIS LEU ILE SER VAL LYS LEU GLY LYS ILE PRO THR VAL          
SEQRES   8 B  390  GLU ASN SER ILE PHE HIS MET ALA ALA TRP SER GLY SER          
SEQRES   9 B  390  ALA CYS HIS ASP GLY ARG GLU TRP THR TYR ILE GLY VAL          
SEQRES  10 B  390  ASP GLY PRO ASP SER ASN ALA LEU ILE LYS ILE LYS TYR          
SEQRES  11 B  390  GLY GLU ALA TYR THR ASP THR TYR HIS SER TYR ALA ASN          
SEQRES  12 B  390  ASN ILE LEU ARG THR GLN GLU SER ALA CYS ASN CYS ILE          
SEQRES  13 B  390  GLY GLY ASP CYS TYR LEU MET ILE THR ASP GLY SER ALA          
SEQRES  14 B  390  SER GLY ILE SER LYS CYS ARG PHE LEU LYS ILE ARG GLU          
SEQRES  15 B  390  GLY ARG ILE ILE LYS GLU ILE PHE PRO THR GLY ARG VAL          
SEQRES  16 B  390  GLU HIS THR GLU GLU CYS THR CYS GLY PHE ALA SER ASN          
SEQRES  17 B  390  LYS THR ILE GLU CYS ALA CYS ARG ASP ASN SER TYR THR          
SEQRES  18 B  390  ALA LYS ARG PRO PHE VAL LYS LEU ASN VAL GLU THR ASP          
SEQRES  19 B  390  THR ALA GLU ILE ARG LEU MET CYS THR GLU THR TYR LEU          
SEQRES  20 B  390  ASP THR PRO ARG PRO ASP ASP GLY SER ILE THR GLY PRO          
SEQRES  21 B  390  CYS GLU SER ASN GLY ASP LYS GLY ARG GLY GLY ILE LYS          
SEQRES  22 B  390  GLY GLY PHE VAL HIS GLN ARG MET ALA SER LYS ILE GLY          
SEQRES  23 B  390  ARG TRP TYR SER ARG THR MET SER LYS THR GLU ARG MET          
SEQRES  24 B  390  GLY MET GLU LEU TYR VAL ARG TYR ASP GLY ASP PRO TRP          
SEQRES  25 B  390  THR ASP SER ASP ALA LEU ALA HIS SER GLY VAL MET VAL          
SEQRES  26 B  390  SER MET LYS GLU PRO GLY TRP TYR SER PHE GLY PHE GLU          
SEQRES  27 B  390  ILE LYS ASP LYS LYS CYS ASP VAL PRO CYS ILE GLY ILE          
SEQRES  28 B  390  GLU MET VAL HIS ASP GLY GLY LYS LYS THR TRP HIS SER          
SEQRES  29 B  390  ALA ALA THR ALA ILE TYR CYS LEU MET GLY SER GLY GLN          
SEQRES  30 B  390  LEU LEU TRP ASP THR VAL THR GLY VAL ASP MET ALA LEU          
MODRES 1NSD ASN A  283  ASN  GLYCOSYLATION SITE                                 
MODRES 1NSD ASN B  283  ASN  GLYCOSYLATION SITE                                 
FTNOTE   1 CA 468 BELONGS TO THE CAH SITE OF MONOMER 1.                         
FTNOTE   2 CA 469 BELONGS TO THE CAH SITE OF MONOMER 2.                         
FTNOTE   3 CA 470 BELONGS TO THE CAL SITE ON THE FOUR-FOLD AXIS.                
FTNOTE   4 CIS PROLINE - PRO A 138                                              
FTNOTE   5 CIS PROLINE - PRO A 325                                              
FTNOTE   6 CIS PROLINE - PRO A 138                                              
FTNOTE   7 CIS PROLINE - PRO B 325                                              
HET    NAG  A 466      28                                                       
HET    NAG  B 466      28                                                       
HET     CA    468       1                                                       
HET     CA    469       1                                                       
HET     CA    470       1                                                       
HET    DAN  A 467      36                                                       
HET    DAN  B 467      36                                                       
HETNAM     NAG N-ACETYL-D-GLUCOSAMINE                                           
HETNAM      CA CALCIUM ION                                                      
HETNAM     DAN 2-DEOXY-2,3-DEHYDRO-N-ACETYL-NEURAMINIC ACID                     
HETSYN     NAG NAG                                                              
FORMUL   3  NAG    2(C8 H15 N O6)                                               
FORMUL   5   CA    3(CA 2+)                                                     
FORMUL   8  DAN    2(C11 H17 N O8)                                              
FORMUL  10  HOH   *506(H2 O)                                                    
HELIX    1   1 SER A   98  GLY A  103  5                                   6    
HELIX    2   2 PRO A  195  ASN A  198  5                                   4    
HELIX    3   3 SER B   98  GLY B  103  5                                   6    
HELIX    4   4 PRO B  195  ASN B  198  5                                   4    
SHEET    1  1A 4 ILE A 113  CYS A 121  0                                        
SHEET    2  1A 4 CYS A 126  THR A 132 -1  N  LEU A 131   O  ARG A 115           
SHEET    3  1A 4 HIS A 154  LYS A 159  1  O  VAL A 158   N  HIS A 128           
SHEET    4  1A 4 ILE A 170  ALA A 174 -1  O  ILE A 170   N  SER A 157           
SHEET    1  2A 4 SER A 177  HIS A 182  0                                        
SHEET    2  2A 4 TRP A 187  ASP A 193 -1  N  VAL A 192   O  SER A 177           
SHEET    3  2A 4 LEU A 200  TYR A 205  1  N  LYS A 202   O  GLY A 191           
SHEET    4  2A 4 ALA A 208  HIS A 214 -1  N  TYR A 213   O  ILE A 201           
SHEET    1 3AA 4 ARG A 222  THR A 223  0                                        
SHEET    2 3AA 4 ASP A 234  GLY A 242 -1  O  THR A 240   N  ARG A 222           
SHEET    3 3AA 4 SER A 248  ARG A 256  1  N  LEU A 253   O  LEU A 237           
SHEET    4 3AA 4 ARG A 259  ILE A 264 -1  N  ARG A 259   O  ARG A 256           
SHEET    1 3BA 4 ASN A 229  ILE A 231  0                                        
SHEET    2 3BA 4 ASP A 234  GLY A 242 -1  O  TYR A 236   N  ASN A 229           
SHEET    3 3BA 4 SER A 248  ARG A 256  1  N  LEU A 253   O  LEU A 237           
SHEET    4 3BA 4 ARG A 259  ILE A 264 -1  N  ARG A 259   O  ARG A 256           
SHEET    1  4A 4 GLU A 274  PHE A 280  0                                        
SHEET    2  4A 4 THR A 285  ARG A 291 -1  O  ARG A 291   N  GLU A 274           
SHEET    3  4A 4 PRO A 300  ASN A 305  1  O  PRO A 300   N  CYS A 290           
SHEET    4  4A 4 THR A 310  LEU A 315 -1  O  THR A 310   N  ASN A 305           
SHEET    1  5A 4 PHE A 351  ARG A 355  0                                        
SHEET    2  5A 4 ILE A 360  ARG A 366 -1  O  TRP A 363   N  VAL A 352           
SHEET    3  5A 4 MET A 376  TYR A 382  1  N  GLU A 377   O  ARG A 366           
SHEET    4  5A 4 ALA A 394  VAL A 400 -1  N  ALA A 394   O  VAL A 380           
SHEET    1  6A 4 SER A 409  LYS A 415  0                                        
SHEET    2  6A 4 ASP A 420  HIS A 430 -1  O  GLY A 425   N  PHE A 410           
SHEET    3  6A 4 SER A 439  LEU A 447  1  O  ALA A 443   N  ILE A 426           
SHEET    4  6A 4 PHE A  91  ILE A  97 -1  O  ILE A  97   N  THR A 442           
SHEET    1  1B 4 ILE B 113  CYS B 121  0                                        
SHEET    2  1B 4 CYS B 126  THR B 132 -1  N  LEU B 131   O  ARG B 115           
SHEET    3  1B 4 HIS B 154  LYS B 159  1  O  VAL B 158   N  HIS B 128           
SHEET    4  1B 4 ILE B 170  ALA B 174 -1  O  ILE B 170   N  SER B 157           
SHEET    1  2B 4 SER B 177  HIS B 182  0                                        
SHEET    2  2B 4 TRP B 187  ASP B 193 -1  N  VAL B 192   O  SER B 177           
SHEET    3  2B 4 LEU B 200  TYR B 205  1  N  LYS B 202   O  GLY B 191           
SHEET    4  2B 4 ALA B 208  HIS B 214 -1  N  TYR B 213   O  ILE B 201           
SHEET    1 3AB 4 ARG B 222  THR B 223  0                                        
SHEET    2 3AB 4 ASP B 234  GLY B 242 -1  O  THR B 240   N  ARG B 222           
SHEET    3 3AB 4 SER B 248  ARG B 256  1  N  LEU B 253   O  LEU B 237           
SHEET    4 3AB 4 ARG B 259  ILE B 264 -1  N  ARG B 259   O  ARG B 256           
SHEET    1 3BB 4 ASN B 229  ILE B 231  0                                        
SHEET    2 3BB 4 ASP B 234  GLY B 242 -1  O  TYR B 236   N  ASN B 229           
SHEET    3 3BB 4 SER B 248  ARG B 256  1  N  LEU B 253   O  LEU B 237           
SHEET    4 3BB 4 ARG B 259  ILE B 264 -1  N  ARG B 259   O  ARG B 256           
SHEET    1  4B 4 GLU B 274  PHE B 280  0                                        
SHEET    2  4B 4 THR B 285  ARG B 291 -1  O  ARG B 291   N  GLU B 274           
SHEET    3  4B 4 PRO B 300  ASN B 305  1  O  PRO B 300   N  CYS B 290           
SHEET    4  4B 4 THR B 310  LEU B 315 -1  O  THR B 310   N  ASN B 305           
SHEET    1  5B 4 PHE B 351  ARG B 355  0                                        
SHEET    2  5B 4 ILE B 360  ARG B 366 -1  O  TRP B 363   N  VAL B 352           
SHEET    3  5B 4 MET B 376  TYR B 382  1  N  GLU B 377   O  ARG B 366           
SHEET    4  5B 4 ALA B 394  VAL B 400 -1  N  ALA B 394   O  VAL B 380           
SHEET    1  6B 4 SER B 409  LYS B 415  0                                        
SHEET    2  6B 4 ASP B 420  HIS B 430 -1  O  GLY B 425   N  PHE B 410           
SHEET    3  6B 4 SER B 439  LEU B 447  1  O  ALA B 443   N  ILE B 426           
SHEET    4  6B 4 PHE B  91  ILE B  97 -1  O  ILE B  97   N  THR B 442           
TURN     1  A1 GLU A 104  GLY A 107                                             
TURN     2  A2 TYR A 142  THR A 145                                             
TURN     3  A3 LYS A 159  LYS A 162     BETA 1L34                               
TURN     4  A4 PRO A 195  ASN A 198     BETA 1L23                               
TURN     5  A5 TYR A 205  ALA A 208     BETA 2L23                               
TURN     6  A6 ILE A 231  ASP A 234     BETA 3L12                               
TURN     7  A7 SER A 243  GLY A 246     BETA 3L23                               
TURN     8  A8 ARG A 256  ARG A 259     BETA 3L34                               
TURN     9  A9 ASP A 328  SER A 331                                             
TURN    10 A10 PRO A 335  SER A 338                                             
TURN    11 A11 ASP A 385  THR A 388     BETA 5L34                               
TURN    12 A12 SER A 401  GLU A 404                                             
TURN    13 A13 GLY A 433  THR A 436     BETA 6L23                               
TURN    14 A14 ASP A 462  LEU A 465                                             
TURN    15  B1 GLU B 104  GLY B 107                                             
TURN    16  B2 TYR B 142  THR B 145                                             
TURN    17  B3 LYS B 159  LYS B 162     BETA 1L34                               
TURN    18  B4 PRO B 195  ASN B 198     BETA 1L23                               
TURN    19  B5 TYR B 205  ALA B 208     BETA 2L23                               
TURN    20  B6 ILE B 231  ASP B 234     BETA 3L12                               
TURN    21  B7 SER B 243  GLY B 246     BETA 3L23                               
TURN    22  B8 ARG B 256  ARG B 259     BETA 3L34                               
TURN    23  B9 ASP B 328  SER B 331                                             
TURN    24 B10 PRO B 335  SER B 338                                             
TURN    25 B11 ASP B 385  THR B 388     BETA 5L34                               
TURN    26 B12 SER B 401  GLU B 404                                             
TURN    27 B13 GLY B 433  THR B 436     BETA 6L23                               
TURN    28 B14 ASP B 462  LEU B 465                                             
SSBOND   1 CYS A   86    CYS A  419                                             
SSBOND   2 CYS A  121    CYS A  126                                             
SSBOND   3 CYS A  181    CYS A  228                                             
SSBOND   4 CYS A  230    CYS A  235                                             
SSBOND   5 CYS A  276    CYS A  290                                             
SSBOND   6 CYS A  278    CYS A  288                                             
SSBOND   7 CYS A  317    CYS A  336                                             
SSBOND   8 CYS A  423    CYS A  446                                             
SSBOND   9 CYS B   86    CYS B  419                                             
SSBOND  10 CYS B  121    CYS B  126                                             
SSBOND  11 CYS B  181    CYS B  228                                             
SSBOND  12 CYS B  230    CYS B  235                                             
SSBOND  13 CYS B  276    CYS B  290                                             
SSBOND  14 CYS B  278    CYS B  288                                             
SSBOND  15 CYS B  317    CYS B  336                                             
SSBOND  16 CYS B  423    CYS B  446                                             
LINK         ND2 ASN A 283                 C1  NAG A 466                        
LINK         ND2 ASN B 283                 C1  NAG B 466                        
CISPEP   1 GLN A  137    PRO A  138          0        -6.95                     
CISPEP   2 THR A  324    PRO A  325          0         0.31                     
CISPEP   3 GLN B  137    PRO B  138          0        -2.22                     
CISPEP   4 THR B  324    PRO B  325          0         2.99                     
SITE     1 CHA  5 ASP A 323  GLY A 343  GLY A 345  THR A 296                    
SITE     2 CHA  5 ASP A 292                                                     
SITE     1 CLA  1 GLU A 167                                                     
SITE     1 NEA 10 ASP A 323  ARG A 115  ARG A 291  ARG A 149                    
SITE     2 NEA 10 ILE A 220  ARG A 222  TRP A 176  GLU A 274                    
SITE     3 NEA 10 ASP A 148  TYR A 408                                          
SITE     1 CHB  5 ASP B 323  GLY B 343  GLY B 345  THR B 296                    
SITE     2 CHB  5 ASP B 292                                                     
SITE     1 CLB  1 GLU B 167                                                     
SITE     1 NEB 10 ASP B 323  ARG B 115  ARG B 291  ARG B 149                    
SITE     2 NEB 10 ILE B 220  ARG B 222  TRP B 176  GLU B 274                    
SITE     3 NEB 10 ASP B 148  TYR B 408                                          
CRYST1   88.900   88.900  222.800  90.00  90.00 120.00 P 31 2 1     12          
ORIGX1      1.000000  0.000000  0.000000        0.00000                         
ORIGX2      0.000000  1.000000  0.000000        0.00000                         
ORIGX3      0.000000  0.000000  1.000000        0.00000                         
SCALE1      0.011249  0.006494  0.000000        0.00000                         
SCALE2      0.000000  0.012989  0.000000        0.00000                         
SCALE3      0.000000  0.000000  0.004488        0.00000                         
MTRIX1   1  0.250000  0.433000 -0.866000        0.00000    1                    
MTRIX2   1  0.433000  0.750000  0.500000        0.00000    1                    
MTRIX3   1  0.866000 -0.500000  0.000000        0.00000    1