PDB entry 1n4x

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HEADER    IMMUNE SYSTEM                           02-NOV-02   1N4X              
TITLE     STRUCTURE OF SCFV 1696 AT ACIDIC PH                                   
COMPND    MOL_ID: 1;                                                            
COMPND   2 MOLECULE: IMMUNOGLOBULIN KAPPA CHAIN VARIABLE REGION;                
COMPND   3 CHAIN: L, M;                                                         
COMPND   4 SYNONYM: SCFV 1696 MOLECULE 1, MONOCLONAL ANTIBODY 1696;             
COMPND   5 ENGINEERED: YES;                                                     
COMPND   6 MOL_ID: 2;                                                           
COMPND   7 MOLECULE: IMMUNOGLOBULIN HEAVY CHAIN VARIABLE REGION;                
COMPND   8 CHAIN: H, I;                                                         
COMPND   9 SYNONYM: SCFV 1696 MOLECULE 2, MONOCLONAL ANTIBODY 1696;             
COMPND  10 ENGINEERED: YES                                                      
SOURCE    MOL_ID: 1;                                                            
SOURCE   2 ORGANISM_SCIENTIFIC: MUS MUSCULUS;                                   
SOURCE   3 ORGANISM_COMMON: MOUSE;                                              
SOURCE   4 EXPRESSION_SYSTEM: ESCHERICHIA COLI;                                 
SOURCE   5 EXPRESSION_SYSTEM_COMMON: BACTERIA;                                  
SOURCE   6 MOL_ID: 2;                                                           
SOURCE   7 ORGANISM_SCIENTIFIC: MUS MUSCULUS;                                   
SOURCE   8 ORGANISM_COMMON: MOUSE;                                              
SOURCE   9 EXPRESSION_SYSTEM: ESCHERICHIA COLI;                                 
SOURCE  10 EXPRESSION_SYSTEM_COMMON: BACTERIA                                   
KEYWDS    IMMUNOGLOBULIN                                                        
EXPDTA    X-RAY DIFFRACTION                                                     
AUTHOR    J.LESCAR,J.BRYNDA,M.FABRY,M.HOREJSI,P.REZACOVA,J.SEDLACEK,            
AUTHOR   2 G.A.BENTLEY                                                          
REVDAT   1   10-JUN-03 1N4X    0                                                
JRNL        AUTH   J.LESCAR,J.BRYNDA,M.FABRY,M.HOREJSI,P.REZACOVA,              
JRNL        AUTH 2 J.SEDLACEK,G.A.BENTLEY                                       
JRNL        TITL   STRUCTURE OF A SINGLE-CHAIN FV FRAGMENT OF AN                
JRNL        TITL 2 ANTIBODY THAT INHIBITS THE HIV-1 AND HIV-2                   
JRNL        TITL 3 PROTEASES.                                                   
JRNL        REF    ACTA CRYSTALLOGR.,SECT.D      V.  59   955 2003              
JRNL        REFN   ASTM ABCRE6  DK ISSN 0907-4449                               
REMARK   1                                                                      
REMARK   2                                                                      
REMARK   2 RESOLUTION. 1.70 ANGSTROMS.                                          
REMARK   3                                                                      
REMARK   3 REFINEMENT.                                                          
REMARK   3   PROGRAM     : CNS                                                  
REMARK   3   AUTHORS     : BRUNGER,ADAMS,CLORE,DELANO,GROS,GROSSE-              
REMARK   3               : KUNSTLEVE,JIANG,KUSZEWSKI,NILGES, PANNU,             
REMARK   3               : READ,RICE,SIMONSON,WARREN                            
REMARK   3                                                                      
REMARK   3  REFINEMENT TARGET : ENGH & HUBER                                    
REMARK   3                                                                      
REMARK   3  DATA USED IN REFINEMENT.                                            
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.70                           
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 20.00                          
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.000                          
REMARK   3   DATA CUTOFF HIGH         (ABS(F)) : NULL                           
REMARK   3   DATA CUTOFF LOW          (ABS(F)) : NULL                           
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : 91.0                           
REMARK   3   NUMBER OF REFLECTIONS             : 45422                          
REMARK   3                                                                      
REMARK   3  FIT TO DATA USED IN REFINEMENT.                                     
REMARK   3   CROSS-VALIDATION METHOD          : NULL                            
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM                          
REMARK   3   R VALUE            (WORKING SET) : 0.227                           
REMARK   3   FREE R VALUE                     : 0.256                           
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : NULL                            
REMARK   3   FREE R VALUE TEST SET COUNT      : 2280                            
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : NULL                            
REMARK   3                                                                      
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.                                  
REMARK   3   TOTAL NUMBER OF BINS USED           : NULL                         
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : NULL                         
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : NULL                         
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : NULL                         
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : NULL                         
REMARK   3   BIN R VALUE           (WORKING SET) : NULL                         
REMARK   3   BIN FREE R VALUE                    : NULL                         
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : NULL                         
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : NULL                         
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : NULL                         
REMARK   3                                                                      
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.                    
REMARK   3   PROTEIN ATOMS            : 3656                                    
REMARK   3   NUCLEIC ACID ATOMS       : 0                                       
REMARK   3   HETEROGEN ATOMS          : 2                                       
REMARK   3   SOLVENT ATOMS            : 530                                     
REMARK   3                                                                      
REMARK   3  B VALUES.                                                           
REMARK   3   FROM WILSON PLOT           (A**2) : NULL                           
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : NULL                           
REMARK   3   OVERALL ANISOTROPIC B VALUE.                                       
REMARK   3    B11 (A**2) : NULL                                                 
REMARK   3    B22 (A**2) : NULL                                                 
REMARK   3    B33 (A**2) : NULL                                                 
REMARK   3    B12 (A**2) : NULL                                                 
REMARK   3    B13 (A**2) : NULL                                                 
REMARK   3    B23 (A**2) : NULL                                                 
REMARK   3                                                                      
REMARK   3  ESTIMATED COORDINATE ERROR.                                         
REMARK   3   ESD FROM LUZZATI PLOT        (A) : NULL                            
REMARK   3   ESD FROM SIGMAA              (A) : NULL                            
REMARK   3   LOW RESOLUTION CUTOFF        (A) : NULL                            
REMARK   3                                                                      
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.                         
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : NULL                            
REMARK   3   ESD FROM C-V SIGMAA          (A) : NULL                            
REMARK   3                                                                      
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.                                   
REMARK   3   BOND LENGTHS                 (A) : 0.005                           
REMARK   3   BOND ANGLES            (DEGREES) : NULL                            
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : NULL                            
REMARK   3   IMPROPER ANGLES        (DEGREES) : NULL                            
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL MODEL : NULL                                      
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA                
REMARK   3   MAIN-CHAIN BOND              (A**2) : NULL  ; NULL                 
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : NULL  ; NULL                 
REMARK   3   SIDE-CHAIN BOND              (A**2) : NULL  ; NULL                 
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : NULL  ; NULL                 
REMARK   3                                                                      
REMARK   3  BULK SOLVENT MODELING.                                              
REMARK   3   METHOD USED : NULL                                                 
REMARK   3   KSOL        : NULL                                                 
REMARK   3   BSOL        : NULL                                                 
REMARK   3                                                                      
REMARK   3  NCS MODEL : NULL                                                    
REMARK   3                                                                      
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT          
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL                 
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL                 
REMARK   3                                                                      
REMARK   3  PARAMETER FILE  1  : NULL                                           
REMARK   3  TOPOLOGY FILE  1   : NULL                                           
REMARK   3                                                                      
REMARK   3  OTHER REFINEMENT REMARKS: NULL                                      
REMARK   4                                                                      
REMARK   4 1N4X COMPLIES WITH FORMAT V. 3.0, 1-DEC-2006                         
REMARK   4                                                                      
REMARK   4 THIS IS THE REMEDIATED VERSION OF THIS PDB ENTRY.                    
REMARK   4 REMEDIATED DATA FILE REVISION 3.101 (2007-05-29)                     
REMARK 100                                                                      
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBJ .                              
REMARK 100 THE RCSB ID CODE IS RCSB017523.                                      
REMARK 200                                                                      
REMARK 200 EXPERIMENTAL DETAILS                                                 
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION                  
REMARK 200  DATE OF DATA COLLECTION        : 06-JAN-1998                        
REMARK 200  TEMPERATURE           (KELVIN) : NULL                               
REMARK 200  PH                             : NULL                               
REMARK 200  NUMBER OF CRYSTALS USED        : 3                                  
REMARK 200                                                                      
REMARK 200  SYNCHROTRON              (Y/N) : Y                                  
REMARK 200  RADIATION SOURCE               : ESRF                               
REMARK 200  BEAMLINE                       : ID2                                
REMARK 200  X-RAY GENERATOR MODEL          : NULL                               
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M                                  
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.99                               
REMARK 200  MONOCHROMATOR                  : NULL                               
REMARK 200  OPTICS                         : MIRRORS                            
REMARK 200                                                                      
REMARK 200  DETECTOR TYPE                  : AREA DETECTOR                      
REMARK 200  DETECTOR MANUFACTURER          : MARRESEARCH                        
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : MARXDS                             
REMARK 200  DATA SCALING SOFTWARE          : XDS                                
REMARK 200                                                                      
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : NULL                               
REMARK 200  RESOLUTION RANGE HIGH      (A) : NULL                               
REMARK 200  RESOLUTION RANGE LOW       (A) : NULL                               
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL                               
REMARK 200                                                                      
REMARK 200 OVERALL.                                                             
REMARK 200  COMPLETENESS FOR RANGE     (%) : NULL                               
REMARK 200  DATA REDUNDANCY                : NULL                               
REMARK 200  R MERGE                    (I) : NULL                               
REMARK 200  R SYM                      (I) : NULL                               
REMARK 200   FOR THE DATA SET  : NULL                               
REMARK 200                                                                      
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.                                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : NULL                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : NULL                     
REMARK 200  COMPLETENESS FOR SHELL     (%) : NULL                               
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL                               
REMARK 200  R MERGE FOR SHELL          (I) : NULL                               
REMARK 200  R SYM FOR SHELL            (I) : NULL                               
REMARK 200   FOR SHELL         : NULL                               
REMARK 200                                                                      
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH                              
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT        
REMARK 200 SOFTWARE USED: AMORE                                                 
REMARK 200 STARTING MODEL: NULL                                                 
REMARK 200                                                                      
REMARK 200 REMARK: NULL                                                         
REMARK 280                                                                      
REMARK 280 CRYSTAL                                                              
REMARK 280 SOLVENT CONTENT, VS   (%): 34.69                                     
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 1.90                     
REMARK 280                                                                      
REMARK 280 CRYSTALLIZATION CONDITIONS: NULL                                     
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY                                            
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 21                       
REMARK 290                                                                      
REMARK 290      SYMOP   SYMMETRY                                                
REMARK 290     NNNMMM   OPERATOR                                                
REMARK 290       1555   X,Y,Z                                                   
REMARK 290       2555   1/2-X,-Y,1/2+Z                                          
REMARK 290       3555   -X,1/2+Y,1/2-Z                                          
REMARK 290       4555   1/2+X,1/2-Y,-Z                                          
REMARK 290                                                                      
REMARK 290     WHERE NNN -> OPERATOR NUMBER                                     
REMARK 290           MMM -> TRANSLATION VECTOR                                  
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS                            
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM             
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY                
REMARK 290 RELATED MOLECULES.                                                   
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000       63.46500            
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       28.65000            
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       30.60500            
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000       28.65000            
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       63.46500            
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       30.60500            
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000            
REMARK 290                                                                      
REMARK 290 REMARK: NULL                                                         
REMARK 300                                                                      
REMARK 300 BIOMOLECULE: 1, 2                                                    
REMARK 300 THIS ENTRY CONTAINS THE CRYSTALLOGRAPHIC ASYMMETRIC UNIT             
REMARK 300 WHICH CONSISTS OF 4 CHAIN(S). SEE REMARK 350 FOR                     
REMARK 300 INFORMATION ON GENERATING THE BIOLOGICAL MOLECULE(S).                
REMARK 350                                                                      
REMARK 350 GENERATING THE BIOMOLECULE                                           
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN           
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE                
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS          
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND                          
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.                               
REMARK 350                                                                      
REMARK 350 BIOMOLECULE: 1                                                       
REMARK 350 APPLY THE FOLLOWING TO CHAINS: L, H                                  
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 350 BIOMOLECULE: 2                                                       
REMARK 350 APPLY THE FOLLOWING TO CHAINS: M, I                                  
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 465                                                                      
REMARK 465 MISSING RESIDUES                                                     
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE                       
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)                
REMARK 465                                                                      
REMARK 465   M RES C SSSEQI                                                     
REMARK 465     ALA I   420                                                      
REMARK 470                                                                      
REMARK 470 MISSING ATOM                                                         
REMARK 470 THE FOLLOWING RESIDUES HAVE MISSING ATOMS(M=MODEL NUMBER;            
REMARK 470 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;          
REMARK 470 I=INSERTION CODE):                                                   
REMARK 470   M RES CSSEQI  ATOMS                                                
REMARK 470     MET L   0    CG    SD    CE                                      
REMARK 470     LEU M   3    CG    CD1   CD2                                     
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT                     
REMARK 500                                                                      
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.                            
REMARK 500                                                                      
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI                             
REMARK 500   NH2  ARG M    24     O    HOH     442              1.96            
REMARK 500   O    HOH     198     O    HOH     319              2.00            
REMARK 500   O    HOH      20     O    HOH     413              2.02            
REMARK 500   O    HOH      95     O    HOH     319              2.02            
REMARK 500   O    SER L    48     O    HOH     460              2.08            
REMARK 500   O    HOH      64     O    HOH     308              2.08            
REMARK 500   O    HOH      45     O    HOH     414              2.14            
REMARK 500   O    HOH     224     O    HOH     382              2.14            
REMARK 500   O    HOH     245     O    HOH     409              2.14            
REMARK 500   O    HOH     163     O    HOH     373              2.15            
REMARK 500   O    HOH     238     O    HOH     410              2.15            
REMARK 500   O    HOH     144     O    HOH     436              2.16            
REMARK 500   N    MET M     0     O    HOH     308              2.17            
REMARK 500   O    HOH      20     O    HOH     290              2.17            
REMARK 500   O    HOH      44     O    HOH     301              2.17            
REMARK 500   O    HOH      45     O    HOH     316              2.17            
REMARK 500   OG   SER H   377     O    HOH     521              2.18            
REMARK 500   O    HOH      45     O    HOH     405              2.18            
REMARK 500   O    HOH      50     O    HOH     309              2.18            
REMARK 500   O    HOH     117     O    HOH     370              2.18            
REMARK 500   O    HOH     141     O    HOH     426              2.18            
REMARK 500   O    HOH     177     O    HOH     460              2.18            
REMARK 500   O    HOH     229     O    HOH     437              2.18            
REMARK 500   OD1  ASP L     1     O    HOH     480              2.19            
REMARK 500   OE1  GLN L    43     O    HOH     491              2.19            
REMARK 500   O    GLY H   366     O    HOH     329              2.19            
REMARK 500   O    HOH     152     O    HOH     330              2.19            
REMARK 500   O    HOH     181     O    HOH     355              2.19            
REMARK 500   O    HOH     222     O    HOH     423              2.19            
REMARK 500   O    HOH     237     O    HOH     386              2.19            
REMARK 500   O    HOH     409     O    HOH     442              2.19            
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: CLOSE CONTACTS                                             
REMARK 500                                                                      
REMARK 500 THE FOLLOWING ATOMS THAT ARE RELATED BY CRYSTALLOGRAPHIC             
REMARK 500 SYMMETRY ARE IN CLOSE CONTACT.  AN ATOM LOCATED WITHIN 0.15          
REMARK 500 ANGSTROMS OF A SYMMETRY RELATED ATOM IS ASSUMED TO BE ON A           
REMARK 500 SPECIAL POSITION AND IS, THEREFORE, LISTED IN REMARK 375             
REMARK 500 INSTEAD OF REMARK 500.  ATOMS WITH NON-BLANK ALTERNATE               
REMARK 500 LOCATION INDICATORS ARE NOT INCLUDED IN THE CALCULATIONS.            
REMARK 500                                                                      
REMARK 500 DISTANCE CUTOFF:                                                     
REMARK 500 2.2 ANGSTROMS FOR CONTACTS NOT INVOLVING HYDROGEN ATOMS              
REMARK 500 1.6 ANGSTROMS FOR CONTACTS INVOLVING HYDROGEN ATOMS                  
REMARK 500                                                                      
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI  SSYMOP   DISTANCE          
REMARK 500   O    HOH     406     O    HOH     413     2565     2.17            
REMARK 500   O    HOH      63     O    HOH     426     2564     2.18            
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND LENGTHS                                      
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,2(A3,1X,A1,I4,A1,1X,A4,3X),F6.3)                  
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   RES CSSEQI ATM2   DEVIATION                     
REMARK 500    PRO H 358   CB    PRO H 358   CG     0.033                        
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES                                       
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)              
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3                                     
REMARK 500    CYS L  93   N   -  CA  -  C   ANGL. DEV. = -9.0 DEGREES           
REMARK 500    GLY L 104   N   -  CA  -  C   ANGL. DEV. = -8.8 DEGREES           
REMARK 500    LEU H 311   N   -  CA  -  C   ANGL. DEV. = -8.7 DEGREES           
REMARK 500    ASN H 352   N   -  CA  -  C   ANGL. DEV. = -8.7 DEGREES           
REMARK 500    ASP H 362   N   -  CA  -  C   ANGL. DEV. =  8.3 DEGREES           
REMARK 500    ASP H 363   N   -  CA  -  C   ANGL. DEV. =  8.6 DEGREES           
REMARK 500    ASP H 373   N   -  CA  -  C   ANGL. DEV. =-11.5 DEGREES           
REMARK 500    CYS H 396   N   -  CA  -  C   ANGL. DEV. =-11.3 DEGREES           
REMARK 500    VAL M  56   N   -  CA  -  C   ANGL. DEV. =  9.3 DEGREES           
REMARK 500    GLY M 104   N   -  CA  -  C   ANGL. DEV. = -8.4 DEGREES           
REMARK 500    ASN I 326   N   -  CA  -  C   ANGL. DEV. = 11.1 DEGREES           
REMARK 500    GLY I 343   N   -  CA  -  C   ANGL. DEV. =  9.7 DEGREES           
REMARK 500    ASN I 352   N   -  CA  -  C   ANGL. DEV. = -9.5 DEGREES           
REMARK 500    ALA I 361   N   -  CA  -  C   ANGL. DEV. =-10.5 DEGREES           
REMARK 500    ASP I 362   N   -  CA  -  C   ANGL. DEV. =  8.8 DEGREES           
REMARK 500    GLY I 366   N   -  CA  -  C   ANGL. DEV. = 14.7 DEGREES           
REMARK 500    LEU I 381   N   -  CA  -  C   ANGL. DEV. = -9.3 DEGREES           
REMARK 500    CYS I 396   N   -  CA  -  C   ANGL. DEV. =-10.5 DEGREES           
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: TORSION ANGLES                                             
REMARK 500                                                                      
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:            
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;               
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                             
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)                    
REMARK 500                                                                      
REMARK 500  M RES CSSEQI        PSI       PHI                                   
REMARK 500    VAL L  56      -46.94     71.43                                   
REMARK 500    VAL M  56      -53.93     69.75                                   
REMARK 525                                                                      
REMARK 525 SOLVENT                                                              
REMARK 525 THE FOLLOWING SOLVENT MOLECULES LIE FARTHER THAN EXPECTED            
REMARK 525 FROM THE PROTEIN OR NUCLEIC ACID MOLECULE AND MAY BE                 
REMARK 525 ASSOCIATED WITH A SYMMETRY RELATED MOLECULE (M=MODEL                 
REMARK 525 NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE          
REMARK 525 NUMBER; I=INSERTION CODE):                                           
REMARK 525                                                                      
REMARK 525  M RES CSSEQI                                                        
REMARK 525    HOH    95        DISTANCE =  5.62 ANGSTROMS                       
REMARK 525    HOH   135        DISTANCE =  5.07 ANGSTROMS                       
REMARK 525    HOH   150        DISTANCE =  5.96 ANGSTROMS                       
REMARK 525    HOH   396        DISTANCE =  6.19 ANGSTROMS                       
REMARK 525    HOH   450        DISTANCE =  6.39 ANGSTROMS                       
REMARK 525    HOH   513        DISTANCE =  5.32 ANGSTROMS                       
REMARK 900                                                                      
REMARK 900 RELATED ENTRIES                                                      
REMARK 900 RELATED ID: 1CL7   RELATED DB: PDB                                   
REMARK 900 FREE FAB 1696                                                        
REMARK 900 RELATED ID: 1JP5   RELATED DB: PDB                                   
REMARK 900 COMPLEXED SCFV1696                                                   
DBREF  1N4X L    1   110  UNP    Q99M37   Q99M37          23    130             
DBREF  1N4X H  304   417  UNP    Q921A6   Q921A6_MOUSE     2    117             
DBREF  1N4X M    1   110  UNP    Q99M37   Q99M37          23    130             
DBREF  1N4X I  304   417  UNP    Q921A6   Q921A6_MOUSE     2    117             
SEQADV 1N4X MET L    0  UNP  Q99M37              INITIATING MET                 
SEQADV 1N4X ILE L  111  UNP  Q99M37              CLONING ARTIFACT               
SEQADV 1N4X ALA L  112  UNP  Q99M37              CLONING ARTIFACT               
SEQADV 1N4X MET M    0  UNP  Q99M37              INITIATING MET                 
SEQADV 1N4X ILE M  111  UNP  Q99M37              CLONING ARTIFACT               
SEQADV 1N4X ALA M  112  UNP  Q99M37              CLONING ARTIFACT               
SEQADV 1N4X GLU H  301  UNP  Q921A6              CLONING ARTIFACT               
SEQADV 1N4X VAL H  302  UNP  Q921A6              CLONING ARTIFACT               
SEQADV 1N4X GLN H  303  UNP  Q921A6              CLONING ARTIFACT               
SEQADV 1N4X VAL H  418  UNP  Q921A6              CLONING ARTIFACT               
SEQADV 1N4X SER H  419  UNP  Q921A6              CLONING ARTIFACT               
SEQADV 1N4X ALA H  420  UNP  Q921A6              CLONING ARTIFACT               
SEQADV 1N4X GLU I  301  UNP  Q921A6              CLONING ARTIFACT               
SEQADV 1N4X VAL I  302  UNP  Q921A6              CLONING ARTIFACT               
SEQADV 1N4X GLN I  303  UNP  Q921A6              CLONING ARTIFACT               
SEQADV 1N4X VAL I  418  UNP  Q921A6              CLONING ARTIFACT               
SEQADV 1N4X SER I  419  UNP  Q921A6              CLONING ARTIFACT               
SEQADV 1N4X ALA I  420  UNP  Q921A6              CLONING ARTIFACT               
SEQRES   1 L  113  MET ASP ILE LEU MET THR GLN THR PRO LEU TYR LEU PRO          
SEQRES   2 L  113  VAL SER LEU GLY ASP GLN ALA SER ILE SER CYS ARG SER          
SEQRES   3 L  113  SER GLN THR ILE VAL HIS ASN ASN GLY ASN THR TYR LEU          
SEQRES   4 L  113  GLU TRP TYR LEU GLN LYS PRO GLY GLN SER PRO GLN LEU          
SEQRES   5 L  113  LEU ILE TYR LYS VAL SER ASN ARG PHE SER GLY VAL PRO          
SEQRES   6 L  113  ASP ARG PHE SER GLY SER GLY SER GLY THR ASP PHE THR          
SEQRES   7 L  113  LEU LYS ILE SER ARG VAL GLU ALA GLU ASP LEU GLY ILE          
SEQRES   8 L  113  TYR TYR CYS PHE GLN GLY SER HIS PHE PRO PRO THR PHE          
SEQRES   9 L  113  GLY GLY GLY THR LYS LEU GLU ILE ALA                          
SEQRES   1 H  120  GLU VAL GLN LEU GLN GLN SER GLY PRO GLU LEU LYS LYS          
SEQRES   2 H  120  PRO GLY GLU THR VAL LYS ILE SER CYS LYS ALA THR ASN          
SEQRES   3 H  120  TYR ALA PHE THR ASP TYR SER MET HIS TRP VAL LYS GLN          
SEQRES   4 H  120  ALA PRO GLY GLY ASP LEU LYS TYR VAL GLY TRP ILE ASN          
SEQRES   5 H  120  THR GLU THR ASP GLU PRO THR PHE ALA ASP ASP PHE LYS          
SEQRES   6 H  120  GLY ARG PHE ALA PHE SER LEU ASP THR SER THR SER THR          
SEQRES   7 H  120  ALA PHE LEU GLN ILE ASN ASN LEU LYS ASN GLU ASP THR          
SEQRES   8 H  120  ALA THR TYR PHE CYS VAL ARG ASP ARG HIS ASP TYR GLY          
SEQRES   9 H  120  GLU ILE PHE THR TYR TRP GLY GLN GLY THR THR VAL THR          
SEQRES  10 H  120  VAL SER ALA                                                  
SEQRES   1 M  113  MET ASP ILE LEU MET THR GLN THR PRO LEU TYR LEU PRO          
SEQRES   2 M  113  VAL SER LEU GLY ASP GLN ALA SER ILE SER CYS ARG SER          
SEQRES   3 M  113  SER GLN THR ILE VAL HIS ASN ASN GLY ASN THR TYR LEU          
SEQRES   4 M  113  GLU TRP TYR LEU GLN LYS PRO GLY GLN SER PRO GLN LEU          
SEQRES   5 M  113  LEU ILE TYR LYS VAL SER ASN ARG PHE SER GLY VAL PRO          
SEQRES   6 M  113  ASP ARG PHE SER GLY SER GLY SER GLY THR ASP PHE THR          
SEQRES   7 M  113  LEU LYS ILE SER ARG VAL GLU ALA GLU ASP LEU GLY ILE          
SEQRES   8 M  113  TYR TYR CYS PHE GLN GLY SER HIS PHE PRO PRO THR PHE          
SEQRES   9 M  113  GLY GLY GLY THR LYS LEU GLU ILE ALA                          
SEQRES   1 I  120  GLU VAL GLN LEU GLN GLN SER GLY PRO GLU LEU LYS LYS          
SEQRES   2 I  120  PRO GLY GLU THR VAL LYS ILE SER CYS LYS ALA THR ASN          
SEQRES   3 I  120  TYR ALA PHE THR ASP TYR SER MET HIS TRP VAL LYS GLN          
SEQRES   4 I  120  ALA PRO GLY GLY ASP LEU LYS TYR VAL GLY TRP ILE ASN          
SEQRES   5 I  120  THR GLU THR ASP GLU PRO THR PHE ALA ASP ASP PHE LYS          
SEQRES   6 I  120  GLY ARG PHE ALA PHE SER LEU ASP THR SER THR SER THR          
SEQRES   7 I  120  ALA PHE LEU GLN ILE ASN ASN LEU LYS ASN GLU ASP THR          
SEQRES   8 I  120  ALA THR TYR PHE CYS VAL ARG ASP ARG HIS ASP TYR GLY          
SEQRES   9 I  120  GLU ILE PHE THR TYR TRP GLY GLN GLY THR THR VAL THR          
SEQRES  10 I  120  VAL SER ALA                                                  
HET     CL   1001       1                                                       
HET     CL   1002       1                                                       
HETNAM      CL CHLORIDE ION                                                     
FORMUL   5   CL    2(CL 1-)                                                     
FORMUL   7  HOH   *530(H2 O)                                                    
HELIX    1   1 GLU L   84  LEU L   88  5                                   5    
HELIX    2   2 ALA H  328  TYR H  332  5                                   5    
HELIX    3   3 THR H  374  THR H  376  5                                   3    
HELIX    4   4 LYS H  387  THR H  391  5                                   5    
HELIX    5   5 GLU M   84  LEU M   88  5                                   5    
HELIX    6   6 ALA I  328  TYR I  332  5                                   5    
HELIX    7   7 ASP I  362  LYS I  365  5                                   4    
HELIX    8   8 THR I  374  THR I  376  5                                   3    
HELIX    9   9 LYS I  387  THR I  391  5                                   5    
SHEET    1   A 4 MET L   4  THR L   7  0                                        
SHEET    2   A 4 ALA L  19  SER L  25 -1  O  SER L  22   N  THR L   7           
SHEET    3   A 4 ASP L  75  ILE L  80 -1  O  ILE L  80   N  ALA L  19           
SHEET    4   A 4 PHE L  67  GLY L  71 -1  N  SER L  70   O  THR L  77           
SHEET    1   B 5 ASN L  58  ARG L  59  0                                        
SHEET    2   B 5 GLN L  50  TYR L  54 -1  N  TYR L  54   O  ASN L  58           
SHEET    3   B 5 LEU L  38  GLN L  43 -1  N  TRP L  40   O  LEU L  52           
SHEET    4   B 5 GLY L  89  GLN L  95 -1  O  TYR L  92   N  TYR L  41           
SHEET    5   B 5 THR L 102  PHE L 103 -1  O  THR L 102   N  GLN L  95           
SHEET    1   C 6 ASN L  58  ARG L  59  0                                        
SHEET    2   C 6 GLN L  50  TYR L  54 -1  N  TYR L  54   O  ASN L  58           
SHEET    3   C 6 LEU L  38  GLN L  43 -1  N  TRP L  40   O  LEU L  52           
SHEET    4   C 6 GLY L  89  GLN L  95 -1  O  TYR L  92   N  TYR L  41           
SHEET    5   C 6 THR L 107  ILE L 111 -1  O  THR L 107   N  TYR L  91           
SHEET    6   C 6 TYR L  10  VAL L  13  1  N  LEU L  11   O  LYS L 108           
SHEET    1   D 4 GLN H 303  GLN H 306  0                                        
SHEET    2   D 4 VAL H 318  THR H 325 -1  O  LYS H 323   N  GLN H 305           
SHEET    3   D 4 THR H 378  ILE H 383 -1  O  LEU H 381   N  ILE H 320           
SHEET    4   D 4 PHE H 368  ASP H 373 -1  N  ALA H 369   O  GLN H 382           
SHEET    1   E 5 GLU H 357  PHE H 360  0                                        
SHEET    2   E 5 LEU H 345  ASN H 352 -1  N  ASN H 352   O  GLU H 357           
SHEET    3   E 5 MET H 334  GLN H 339 -1  N  LYS H 338   O  LYS H 346           
SHEET    4   E 5 ALA H 392  ASP H 399 -1  O  VAL H 397   N  HIS H 335           
SHEET    5   E 5 PHE H 407  TRP H 410 -1  O  TYR H 409   N  ARG H 398           
SHEET    1   F 6 GLU H 357  PHE H 360  0                                        
SHEET    2   F 6 LEU H 345  ASN H 352 -1  N  ASN H 352   O  GLU H 357           
SHEET    3   F 6 MET H 334  GLN H 339 -1  N  LYS H 338   O  LYS H 346           
SHEET    4   F 6 ALA H 392  ASP H 399 -1  O  VAL H 397   N  HIS H 335           
SHEET    5   F 6 THR H 414  VAL H 418 -1  O  VAL H 416   N  ALA H 392           
SHEET    6   F 6 GLU H 310  LYS H 312  1  N  GLU H 310   O  THR H 417           
SHEET    1   G 4 MET M   4  THR M   7  0                                        
SHEET    2   G 4 ALA M  19  SER M  25 -1  O  SER M  22   N  THR M   7           
SHEET    3   G 4 ASP M  75  ILE M  80 -1  O  ILE M  80   N  ALA M  19           
SHEET    4   G 4 PHE M  67  SER M  72 -1  N  SER M  68   O  LYS M  79           
SHEET    1   H 5 ASN M  58  ARG M  59  0                                        
SHEET    2   H 5 GLN M  50  TYR M  54 -1  N  TYR M  54   O  ASN M  58           
SHEET    3   H 5 LEU M  38  GLN M  43 -1  N  TRP M  40   O  LEU M  52           
SHEET    4   H 5 GLY M  89  GLN M  95 -1  O  ILE M  90   N  GLN M  43           
SHEET    5   H 5 THR M 102  PHE M 103 -1  O  THR M 102   N  GLN M  95           
SHEET    1   I 6 ASN M  58  ARG M  59  0                                        
SHEET    2   I 6 GLN M  50  TYR M  54 -1  N  TYR M  54   O  ASN M  58           
SHEET    3   I 6 LEU M  38  GLN M  43 -1  N  TRP M  40   O  LEU M  52           
SHEET    4   I 6 GLY M  89  GLN M  95 -1  O  ILE M  90   N  GLN M  43           
SHEET    5   I 6 THR M 107  ILE M 111 -1  O  THR M 107   N  TYR M  91           
SHEET    6   I 6 TYR M  10  VAL M  13  1  N  LEU M  11   O  LYS M 108           
SHEET    1   J 4 GLN I 303  GLN I 306  0                                        
SHEET    2   J 4 VAL I 318  THR I 325 -1  O  LYS I 323   N  GLN I 305           
SHEET    3   J 4 THR I 378  ILE I 383 -1  O  LEU I 381   N  ILE I 320           
SHEET    4   J 4 PHE I 368  ASP I 373 -1  N  SER I 371   O  PHE I 380           
SHEET    1   K 5 PRO I 358  PHE I 360  0                                        
SHEET    2   K 5 LEU I 345  ILE I 351 -1  N  TRP I 350   O  THR I 359           
SHEET    3   K 5 MET I 334  GLN I 339 -1  N  LYS I 338   O  LYS I 346           
SHEET    4   K 5 ALA I 392  ASP I 399 -1  O  PHE I 395   N  VAL I 337           
SHEET    5   K 5 PHE I 407  TRP I 410 -1  O  TYR I 409   N  ARG I 398           
SHEET    1   L 6 PRO I 358  PHE I 360  0                                        
SHEET    2   L 6 LEU I 345  ILE I 351 -1  N  TRP I 350   O  THR I 359           
SHEET    3   L 6 MET I 334  GLN I 339 -1  N  LYS I 338   O  LYS I 346           
SHEET    4   L 6 ALA I 392  ASP I 399 -1  O  PHE I 395   N  VAL I 337           
SHEET    5   L 6 THR I 414  VAL I 418 -1  O  THR I 414   N  TYR I 394           
SHEET    6   L 6 GLU I 310  LYS I 312  1  N  LYS I 312   O  THR I 417           
SSBOND   1 CYS L   23    CYS L   93                                             
SSBOND   2 CYS H  322    CYS H  396                                             
SSBOND   3 CYS M   23    CYS M   93                                             
SSBOND   4 CYS I  322    CYS I  396                                             
CISPEP   1 THR L    7    PRO L    8          0        -0.37                     
CISPEP   2 PHE L   99    PRO L  100          0        -0.29                     
CISPEP   3 THR M    7    PRO M    8          0        -0.22                     
CISPEP   4 PHE M   99    PRO M  100          0         0.17                     
CRYST1  126.930   61.210   57.300  90.00  90.00  90.00 P 21 21 21    8          
ORIGX1      1.000000  0.000000  0.000000        0.00000                         
ORIGX2      0.000000  1.000000  0.000000        0.00000                         
ORIGX3      0.000000  0.000000  1.000000        0.00000                         
SCALE1      0.007878  0.000000  0.000000        0.00000                         
SCALE2      0.000000  0.016337  0.000000        0.00000                         
SCALE3      0.000000  0.000000  0.017452        0.00000