PDB entry 1mju

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HEADER    IMMUNE SYSTEM                           28-AUG-02   1MJU              
TITLE     1.22 ANGSTROM RESOLUTION CRYSTAL STRUCTURE OF THE FAB                 
TITLE    2 FRAGMENT OF ESTEROLYTIC ANTIBODY MS6-12                              
COMPND    MOL_ID: 1;                                                            
COMPND   2 MOLECULE: IMMUNOGLOBULIN MS6-12;                                     
COMPND   3 CHAIN: L;                                                            
COMPND   4 FRAGMENT: FAB FRAGMENT, LIGHT CHAIN;                                 
COMPND   5 ENGINEERED: YES;                                                     
COMPND   6 MOL_ID: 2;                                                           
COMPND   7 MOLECULE: IMMUNOGLOBULIN MS6-12;                                     
COMPND   8 CHAIN: H;                                                            
COMPND   9 FRAGMENT: FAB FRAGMENT, HEAVY CHAIN;                                 
COMPND  10 ENGINEERED: YES                                                      
SOURCE    MOL_ID: 1;                                                            
SOURCE   2 ORGANISM_SCIENTIFIC: MUS MUSCULUS;                                   
SOURCE   3 ORGANISM_COMMON: MOUSE;                                              
SOURCE   4 CELL_LINE: HYBRIDOMA;                                                
SOURCE   5 EXPRESSION_SYSTEM: MUS MUSCULUS;                                     
SOURCE   6 EXPRESSION_SYSTEM_COMMON: MOUSE;                                     
SOURCE   7 OTHER_DETAILS: FUSING OF MOUSE SPLENOCYTES WITH MYELOMA              
SOURCE   8 CELL;                                                                
SOURCE   9 MOL_ID: 2;                                                           
SOURCE  10 ORGANISM_SCIENTIFIC: MUS MUSCULUS;                                   
SOURCE  11 ORGANISM_COMMON: MOUSE;                                              
SOURCE  12 CELL_LINE: HYBRIDOMA;                                                
SOURCE  13 EXPRESSION_SYSTEM: MUS MUSCULUS;                                     
SOURCE  14 EXPRESSION_SYSTEM_COMMON: MOUSE;                                     
SOURCE  15 OTHER_DETAILS: FUSING OF MOUSE SPLENOCYTES WITH MYELOMA              
SOURCE  16 CELL                                                                 
KEYWDS    CATALYTIC ANTIBODY, ESTER HYDROLYSIS, ESTEROLYTIC, FAB,               
KEYWDS   2 IMMUNOGLOBULIN                                                       
EXPDTA    X-RAY DIFFRACTION                                                     
AUTHOR    S.N.RUZHEINIKOV,T.A.MURANOVA,S.E.SEDELNIKOVA,L.J.PARTRIDGE,           
AUTHOR   2 G.M.BLACKBURN,I.A.MURRAY,H.KAKINUMA,N.TAKASHI,K.SHIMAZAKI,           
AUTHOR   3 J.SUN,Y.NISHI,D.W.RICE                                               
REVDAT   1   23-SEP-03 1MJU    0                                                
JRNL        AUTH   S.N.RUZHEINIKOV,T.A.MURANOVA,S.E.SEDELNIKOVA,                
JRNL        AUTH 2 L.J.PARTRIDGE,G.M.BLACKBURN,I.A.MURRAY,H.KAKINUMA,           
JRNL        AUTH 3 N.TAKASHI,K.SHIMAZAKI,J.SUN,Y.NISHI,D.W.RICE                 
JRNL        TITL   HIGH-RESOLUTION CRYSTAL STRUCTURE OF THE                     
JRNL        TITL 2 FAB-FRAGMENTS OF A FAMILY OF MOUSE CATALYTIC                 
JRNL        TITL 3 ANTIBODIES WITH ESTERASE ACTIVITY                            
JRNL        REF    J.MOL.BIOL.                   V. 332   423 2003              
JRNL        REFN   ASTM JMOBAK  UK ISSN 0022-2836                               
REMARK   1                                                                      
REMARK   1 REFERENCE 1                                                          
REMARK   1  AUTH   T.A.MURANOVA,S.N.RUZHEINIKOV,S.E.SEDELNIKOVA,                
REMARK   1  AUTH 2 A.MOIR,L.J.PARTRIDGE,H.KAKINUMA,N.TAKASHI,                   
REMARK   1  AUTH 3 K.SHIMAZAKI,J.SUN,Y.NISHI,D.W.RICE                           
REMARK   1  TITL   THE PREPARATION AND CRYSTALLIZATION OF FAB                   
REMARK   1  TITL 2 FRAGMENTS OF A FAMILY OF MOUSE ESTEROLYTIC                   
REMARK   1  TITL 3 CATALYTIC ANTIBODIES AND THEIR COMPLEXES WITH A              
REMARK   1  TITL 4 TRANSITION-STATE ANALOGUE                                    
REMARK   1  REF    ACTA CRYSTALLOGR., SECT.D     V.  57  1192 2001              
REMARK   1  REFN   ASTM ABCRE6  DK ISSN 0907-4449                               
REMARK   2                                                                      
REMARK   2 RESOLUTION. 1.22 ANGSTROMS.                                          
REMARK   3                                                                      
REMARK   3 REFINEMENT.                                                          
REMARK   3   PROGRAM     : REFMAC 5, CNS 1.1                                    
REMARK   3   AUTHORS     : BRUNGER,ADAMS,CLORE,DELANO,GROS,GROSSE-              
REMARK   3               : KUNSTLEVE,JIANG,KUSZEWSKI,NILGES, PANNU,             
REMARK   3               : READ,RICE,SIMONSON,WARREN                            
REMARK   3                                                                      
REMARK   3  REFINEMENT TARGET : MAXIMUM LIKELIHOOD                              
REMARK   3                                                                      
REMARK   3  DATA USED IN REFINEMENT.                                            
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.22                           
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 10.00                          
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.000                          
REMARK   3   DATA CUTOFF HIGH         (ABS(F)) : NULL                           
REMARK   3   DATA CUTOFF LOW          (ABS(F)) : NULL                           
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : 93.4                           
REMARK   3   NUMBER OF REFLECTIONS             : 141368                         
REMARK   3                                                                      
REMARK   3  FIT TO DATA USED IN REFINEMENT.                                     
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT                      
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM                          
REMARK   3   R VALUE            (WORKING SET) : 0.123                           
REMARK   3   FREE R VALUE                     : 0.154                           
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.000                           
REMARK   3   FREE R VALUE TEST SET COUNT      : 7075                            
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : NULL                            
REMARK   3                                                                      
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.                                  
REMARK   3   TOTAL NUMBER OF BINS USED           : 20                           
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 1.22                         
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 1.25                         
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : NULL                         
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : 6913                         
REMARK   3   BIN R VALUE           (WORKING SET) : 0.1700                       
REMARK   3   BIN FREE R VALUE                    : 0.2120                       
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : NULL                         
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : 352                          
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : NULL                         
REMARK   3                                                                      
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.                    
REMARK   3   PROTEIN ATOMS            : 3362                                    
REMARK   3   NUCLEIC ACID ATOMS       : 0                                       
REMARK   3   HETEROGEN ATOMS          : 0                                       
REMARK   3   SOLVENT ATOMS            : 971                                     
REMARK   3                                                                      
REMARK   3  B VALUES.                                                           
REMARK   3   FROM WILSON PLOT           (A**2) : NULL                           
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 13.54                          
REMARK   3   OVERALL ANISOTROPIC B VALUE.                                       
REMARK   3    B11 (A**2) : -0.21000                                             
REMARK   3    B22 (A**2) : 0.56000                                              
REMARK   3    B33 (A**2) : -0.35000                                             
REMARK   3    B12 (A**2) : 0.00000                                              
REMARK   3    B13 (A**2) : 0.00000                                              
REMARK   3    B23 (A**2) : 0.00000                                              
REMARK   3                                                                      
REMARK   3  ESTIMATED COORDINATE ERROR.                                         
REMARK   3   ESD FROM LUZZATI PLOT        (A) : NULL                            
REMARK   3   ESD FROM SIGMAA              (A) : NULL                            
REMARK   3   LOW RESOLUTION CUTOFF        (A) : NULL                            
REMARK   3                                                                      
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.                         
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : NULL                            
REMARK   3   ESD FROM C-V SIGMAA          (A) : NULL                            
REMARK   3                                                                      
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.                                   
REMARK   3   BOND LENGTHS                 (A) : NULL                            
REMARK   3   BOND ANGLES            (DEGREES) : NULL                            
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : NULL                            
REMARK   3   IMPROPER ANGLES        (DEGREES) : NULL                            
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL MODEL : ANISOTROPIC                               
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA                
REMARK   3   MAIN-CHAIN BOND              (A**2) : NULL  ; NULL                 
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : NULL  ; NULL                 
REMARK   3   SIDE-CHAIN BOND              (A**2) : NULL  ; NULL                 
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : NULL  ; NULL                 
REMARK   3                                                                      
REMARK   3  BULK SOLVENT MODELING.                                              
REMARK   3   METHOD USED : BABINET MODEL WITH MASK                              
REMARK   3   KSOL        : NULL                                                 
REMARK   3   BSOL        : NULL                                                 
REMARK   3                                                                      
REMARK   3  NCS MODEL : NULL                                                    
REMARK   3                                                                      
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT          
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL                 
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL                 
REMARK   3                                                                      
REMARK   3  PARAMETER FILE  1  : NULL                                           
REMARK   3  TOPOLOGY FILE  1   : NULL                                           
REMARK   3                                                                      
REMARK   3  OTHER REFINEMENT REMARKS: HYDROGENS HAVE BEEN ADDED IN THE          
REMARK   3  RIDING POSITIONS                                                    
REMARK   4                                                                      
REMARK   4 1MJU COMPLIES WITH FORMAT V. 3.0, 1-DEC-2006                         
REMARK   4                                                                      
REMARK   4 THIS IS THE REMEDIATED VERSION OF THIS PDB ENTRY.                    
REMARK   4 REMEDIATED DATA FILE REVISION 3.101 (2007-05-29)                     
REMARK 100                                                                      
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB .                              
REMARK 100 THE RCSB ID CODE IS RCSB016966.                                      
REMARK 200                                                                      
REMARK 200 EXPERIMENTAL DETAILS                                                 
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION                  
REMARK 200  DATE OF DATA COLLECTION        : 09-NOV-2000                        
REMARK 200  TEMPERATURE           (KELVIN) : 100.0                              
REMARK 200  PH                             : 5.00                               
REMARK 200  NUMBER OF CRYSTALS USED        : 1                                  
REMARK 200                                                                      
REMARK 200  SYNCHROTRON              (Y/N) : Y                                  
REMARK 200  RADIATION SOURCE               : SRS                                
REMARK 200  BEAMLINE                       : PX9.6                              
REMARK 200  X-RAY GENERATOR MODEL          : NULL                               
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M                                  
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.870                              
REMARK 200  MONOCHROMATOR                  : SI 111                             
REMARK 200  OPTICS                         : MIRRORS                            
REMARK 200                                                                      
REMARK 200  DETECTOR TYPE                  : CCD                                
REMARK 200  DETECTOR MANUFACTURER          : ADSC QUANTUM 4                     
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : DENZO                              
REMARK 200  DATA SCALING SOFTWARE          : SCALEPACK                          
REMARK 200                                                                      
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 141368                             
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.220                              
REMARK 200  RESOLUTION RANGE LOW       (A) : 10.000                             
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : -3.000                             
REMARK 200                                                                      
REMARK 200 OVERALL.                                                             
REMARK 200  COMPLETENESS FOR RANGE     (%) : 93.4                               
REMARK 200  DATA REDUNDANCY                : 8.300                              
REMARK 200  R MERGE                    (I) : 0.03500                            
REMARK 200  R SYM                      (I) : NULL                               
REMARK 200   FOR THE DATA SET  : 32.0000                            
REMARK 200                                                                      
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.                                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.22                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.24                     
REMARK 200  COMPLETENESS FOR SHELL     (%) : 63.3                               
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL                               
REMARK 200  R MERGE FOR SHELL          (I) : 0.21900                            
REMARK 200  R SYM FOR SHELL            (I) : NULL                               
REMARK 200   FOR SHELL         : 3.400                              
REMARK 200                                                                      
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH                              
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MIR                          
REMARK 200 SOFTWARE USED: CNS                                                   
REMARK 200 STARTING MODEL: PDB ENTRY 1MH5                                       
REMARK 200                                                                      
REMARK 200 REMARK: NULL                                                         
REMARK 280                                                                      
REMARK 280 CRYSTAL                                                              
REMARK 280 SOLVENT CONTENT, VS   (%): 52.89                                     
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.61                     
REMARK 280                                                                      
REMARK 280 CRYSTALLIZATION CONDITIONS: SODIUM ACETATE, NACL, PH 5.0, VAPOR      
REMARK 280  DIFFUSION, HANGING DROP, TEMPERATURE 290K                           
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY                                            
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 21                       
REMARK 290                                                                      
REMARK 290      SYMOP   SYMMETRY                                                
REMARK 290     NNNMMM   OPERATOR                                                
REMARK 290       1555   X,Y,Z                                                   
REMARK 290       2555   1/2-X,-Y,1/2+Z                                          
REMARK 290       3555   -X,1/2+Y,1/2-Z                                          
REMARK 290       4555   1/2+X,1/2-Y,-Z                                          
REMARK 290                                                                      
REMARK 290     WHERE NNN -> OPERATOR NUMBER                                     
REMARK 290           MMM -> TRANSLATION VECTOR                                  
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS                            
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM             
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY                
REMARK 290 RELATED MOLECULES.                                                   
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000       28.02300            
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       69.28150            
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       32.67450            
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000       69.28150            
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       28.02300            
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       32.67450            
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000            
REMARK 290                                                                      
REMARK 290 REMARK: NULL                                                         
REMARK 300                                                                      
REMARK 300 BIOMOLECULE: 1                                                       
REMARK 300 THIS ENTRY CONTAINS THE CRYSTALLOGRAPHIC ASYMMETRIC UNIT             
REMARK 300 WHICH CONSISTS OF 2 CHAIN(S). SEE REMARK 350 FOR                     
REMARK 300 INFORMATION ON GENERATING THE BIOLOGICAL MOLECULE(S).                
REMARK 350                                                                      
REMARK 350 GENERATING THE BIOMOLECULE                                           
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN           
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE                
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS          
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND                          
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.                               
REMARK 350                                                                      
REMARK 350 BIOMOLECULE: 1                                                       
REMARK 350 APPLY THE FOLLOWING TO CHAINS: L, H                                  
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 465                                                                      
REMARK 465 MISSING RESIDUES                                                     
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE                       
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)                
REMARK 465                                                                      
REMARK 465   M RES C SSSEQI                                                     
REMARK 465     GLY H   131                                                      
REMARK 465     ASP H   132                                                      
REMARK 465     THR H   133                                                      
REMARK 465     SER H   134                                                      
REMARK 465     GLY H   135                                                      
REMARK 465     SER H   136                                                      
REMARK 465     THR H   231                                                      
REMARK 465     ILE H   232                                                      
REMARK 465     LYS H   233                                                      
REMARK 465     PRO H   234                                                      
REMARK 465     CYS H   235                                                      
REMARK 465     PRO H   236                                                      
REMARK 465     PRO H   237                                                      
REMARK 465     CYS H   238                                                      
REMARK 465     LYS H   239                                                      
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT                     
REMARK 500                                                                      
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.                            
REMARK 500                                                                      
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI                             
REMARK 500   O    HOH     852     O    HOH     933              1.52            
REMARK 500   O    HOH     374     O    HOH     893              1.55            
REMARK 500   O    HOH     199     O    HOH     598              1.56            
REMARK 500   C    CYS H   128     O    HOH     894              1.58            
REMARK 500   O    HOH     178     O    HOH     927              1.58            
REMARK 500   OE1  GLN H   105     O    HOH     928              1.59            
REMARK 500   NE2  GLN H   203     O    HOH     852              1.62            
REMARK 500   O    HOH     237     O    HOH     930              1.62            
REMARK 500   O    HOH     138     O    HOH     909              1.65            
REMARK 500   O    HOH     241     O    HOH     826              1.66            
REMARK 500   O    HOH     410     O    HOH     675              1.73            
REMARK 500   O    HOH     341     O    HOH     456              1.74            
REMARK 500   O    HOH     312     O    HOH     561              1.76            
REMARK 500   O    HOH     886     O    HOH     887              1.77            
REMARK 500   O    HOH     460     O    HOH     571              1.83            
REMARK 500   O    HOH     221     O    HOH     416              1.85            
REMARK 500   O    HOH     201     O    HOH     920              1.86            
REMARK 500   O    HOH     438     O    HOH     757              1.86            
REMARK 500   NH2  ARG L    24     O    HOH     828              1.87            
REMARK 500   O    HOH     306     O    HOH     935              1.88            
REMARK 500   O    HOH     182     O    HOH     889              1.91            
REMARK 500   O    HOH     177     O    HOH     930              1.92            
REMARK 500   O    HOH     533     O    HOH     794              1.92            
REMARK 500   O    HOH     188     O    HOH     906              1.93            
REMARK 500   O    GLN H   179     O    HOH     572              1.96            
REMARK 500   O    HOH     433     O    HOH     503              1.96            
REMARK 500   O    HOH     729     O    HOH     900              1.97            
REMARK 500   CG2  THR H   198     O    HOH     932              1.98            
REMARK 500   O    HOH     213     O    HOH     890              2.01            
REMARK 500   O    HOH     267     O    HOH     763              2.01            
REMARK 500   O    HOH     407     O    HOH     896              2.01            
REMARK 500   N    ALA L     7     O    HOH     906              2.02            
REMARK 500   OE2  GLU L   154     O    HOH     405              2.04            
REMARK 500   O    HOH     226     O    HOH     456              2.04            
REMARK 500   O    HOH     546     O    HOH     897              2.04            
REMARK 500   NH2  ARG L    77     O    HOH     727              2.07            
REMARK 500   OE2  GLU L    79     O    HOH     406              2.07            
REMARK 500   O    HOH     620     O    HOH     919              2.07            
REMARK 500   O    HOH     137     O    HOH     769              2.08            
REMARK 500   O    HOH     659     O    HOH     792              2.08            
REMARK 500   O2   GOL    1402     O    HOH     486              2.09            
REMARK 500   O    HOH     524     O    HOH     922              2.09            
REMARK 500   O    HOH     552     O    HOH     577              2.09            
REMARK 500   OE1  GLU H    46     O    HOH     459              2.11            
REMARK 500   N    SER H   137     O    HOH     929              2.12            
REMARK 500   O    HOH     631     O    HOH     732              2.13            
REMARK 500   O    HOH     750     O    HOH     809              2.16            
REMARK 500   NZ   LYS L   183     OE2  GLU L   187              2.17            
REMARK 500   O    HOH     344     O    HOH     663              2.19            
REMARK 500   O    HOH     402     O    HOH     539              2.19            
REMARK 500   O    HOH     849     O    HOH     871              2.19            
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: CLOSE CONTACTS                                             
REMARK 500                                                                      
REMARK 500 THE FOLLOWING ATOMS THAT ARE RELATED BY CRYSTALLOGRAPHIC             
REMARK 500 SYMMETRY ARE IN CLOSE CONTACT.  AN ATOM LOCATED WITHIN 0.15          
REMARK 500 ANGSTROMS OF A SYMMETRY RELATED ATOM IS ASSUMED TO BE ON A           
REMARK 500 SPECIAL POSITION AND IS, THEREFORE, LISTED IN REMARK 375             
REMARK 500 INSTEAD OF REMARK 500.  ATOMS WITH NON-BLANK ALTERNATE               
REMARK 500 LOCATION INDICATORS ARE NOT INCLUDED IN THE CALCULATIONS.            
REMARK 500                                                                      
REMARK 500 DISTANCE CUTOFF:                                                     
REMARK 500 2.2 ANGSTROMS FOR CONTACTS NOT INVOLVING HYDROGEN ATOMS              
REMARK 500 1.6 ANGSTROMS FOR CONTACTS INVOLVING HYDROGEN ATOMS                  
REMARK 500                                                                      
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI  SSYMOP   DISTANCE          
REMARK 500   O    HOH     391     O    HOH     915     2555     1.67            
REMARK 500   O    HOH     384     O    HOH     926     2555     1.88            
REMARK 500   O    HOH     850     O    HOH     908     2554     1.95            
REMARK 500   O    HOH     880     O    HOH     916     3655     2.03            
REMARK 500   O    HOH     306     O    HOH     926     2555     2.04            
REMARK 500   O    HOH     850     O    HOH     935     2554     2.06            
REMARK 500   O    HOH     303     O    HOH     489     3645     2.09            
REMARK 500   CG2  THR L   202     O    HOH     927     3645     2.15            
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND LENGTHS                                      
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,2(A3,1X,A1,I4,A1,1X,A4,3X),F6.3)                  
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   RES CSSEQI ATM2   DEVIATION                     
REMARK 500    ASN L 190   CG    ASN L 190   OD1    0.079                        
REMARK 500    VAL H 152   CA    VAL H 152   CB     0.088                        
REMARK 500    VAL H 191   CA    VAL H 191   CB     0.079                        
REMARK 500    SER H 196   C     THR H 198   N      0.090                        
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES                                       
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)              
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3                                     
REMARK 500    GLN L   6   N   -  CA  -  C   ANGL. DEV. =-12.0 DEGREES           
REMARK 500    LEU L 104   CA  -  CB  -  CG  ANGL. DEV. = 11.7 DEGREES           
REMARK 500    ASN L 145   N   -  CA  -  C   ANGL. DEV. =-10.8 DEGREES           
REMARK 500    VAL L 146   N   -  CA  -  CB  ANGL. DEV. =-11.7 DEGREES           
REMARK 500    VAL L 146   CG1 -  CB  -  CG2 ANGL. DEV. = 13.2 DEGREES           
REMARK 500    MET L 175   CG  -  SD  -  CE  ANGL. DEV. =-18.0 DEGREES           
REMARK 500    VAL H 177   CG1 -  CB  -  CG2 ANGL. DEV. =-16.2 DEGREES           
REMARK 500    VAL H 177   CG1 -  CB  -  CG2 ANGL. DEV. = 13.3 DEGREES           
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: TORSION ANGLES                                             
REMARK 500                                                                      
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:            
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;               
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                             
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)                    
REMARK 500                                                                      
REMARK 500  M RES CSSEQI        PSI       PHI                                   
REMARK 500    MET L  51      -38.81     74.07                                   
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: NON-CIS, NON-TRANS                                         
REMARK 500                                                                      
REMARK 500 THE FOLLOWING PEPTIDE BONDS DEVIATE SIGNIFICANTLY FROM BOTH          
REMARK 500 CIS AND TRANS CONFORMATION.  CIS BONDS, IF ANY, ARE LISTED           
REMARK 500 ON CISPEP RECORDS.  TRANS IS DEFINED AS 180 +/- 30 AND               
REMARK 500 CIS IS DEFINED AS 0 +/- 30 DEGREES.                                  
REMARK 500                                 MODEL     OMEGA                      
REMARK 500 ASN L  145    VAL L  146                  149.94                     
REMARK 500 PRO H  149    GLU H  150                  144.16                     
REMARK 500 PRO H  151    VAL H  152                  134.57                     
REMARK 525                                                                      
REMARK 525 SOLVENT                                                              
REMARK 525 THE FOLLOWING SOLVENT MOLECULES LIE FARTHER THAN EXPECTED            
REMARK 525 FROM THE PROTEIN OR NUCLEIC ACID MOLECULE AND MAY BE                 
REMARK 525 ASSOCIATED WITH A SYMMETRY RELATED MOLECULE (M=MODEL                 
REMARK 525 NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE          
REMARK 525 NUMBER; I=INSERTION CODE):                                           
REMARK 525                                                                      
REMARK 525  M RES CSSEQI                                                        
REMARK 525    HOH   825        DISTANCE =  5.02 ANGSTROMS                       
REMARK 900                                                                      
REMARK 900 RELATED ENTRIES                                                      
REMARK 900 RELATED ID: 1MH5   RELATED DB: PDB                                   
REMARK 900 THE STRUCTURE OF THE COMPLEX OF THE FAB FRAGMENT OF                  
REMARK 900 THEESTEROLYTIC ANTIBODY MS6-164 AND A TRANSITION-STATE               
REMARK 900 ANALOG                                                               
REMARK 900 RELATED ID: 1MIE   RELATED DB: PDB                                   
REMARK 900 CRYSTAL STRUCTURE OF THE FAB FRAGMENT OF ESTEROLYTIC                 
REMARK 900 ANTIBODY MS5-393                                                     
REMARK 900 RELATED ID: 1MJ7   RELATED DB: PDB                                   
REMARK 900 CRYSTAL STRUCTURE OF THE COMPLEX OF THE FAB FRAGMENT OF              
REMARK 900 ESTEROLYTIC ANTIBODY MS5-393D9 AND A TRANSITION-STATE ANALOG         
REMARK 900 RELATED ID: 1MJ8   RELATED DB: PDB                                   
REMARK 900 HIGH RESOLUTION CRYSTAL STRUCTURE OF THE COMPLEX OF THE FAB          
REMARK 900 FRAGMENT OF THE ESTEROLYTIC ANTIBODY MS6-126 AND A                   
REMARK 900 TRANSITION-STATE ANALOG                                              
REMARK 900 RELATED ID: 1MJJ   RELATED DB: PDB                                   
REMARK 900 HIGH RESOLUTION CRYSTAL STRUCTURE OF THE COMPLEX OF THE FAB          
REMARK 900 FRAGMENT OF ESTEROLYTIC ANTIBODY MS6-12 AND A TRANSITION-            
REMARK 900 STATE ANALOG                                                         
REMARK 999                                                                      
REMARK 999 SEQUENCE                                                             
REMARK 999 AN APPROPRIATE SEQUENCE DATABASE REFERENCE WAS                       
REMARK 999 NOT AVAILABLE AT THE TIME OF PROCESSING.                             
SEQRES   1 L  219  ASP ILE VAL MET THR GLN ALA ALA PRO SER VAL PRO VAL          
SEQRES   2 L  219  THR PRO GLY GLU SER VAL SER ILE SER CYS ARG SER SER          
SEQRES   3 L  219  LYS SER LEU LEU HIS SER ASN GLY ASN THR TYR LEU TYR          
SEQRES   4 L  219  TRP PHE LEU GLN ARG PRO GLY GLN SER PRO GLN LEU LEU          
SEQRES   5 L  219  ILE TYR ARG MET SER ASN LEU ALA SER GLY VAL PRO ASP          
SEQRES   6 L  219  ARG PHE SER GLY SER GLY SER GLY THR ALA PHE THR LEU          
SEQRES   7 L  219  ARG ILE SER ARG VAL GLU ALA GLU ASP VAL GLY VAL TYR          
SEQRES   8 L  219  TYR CYS LEU GLN HIS LEU GLU TYR PRO PHE THR PHE GLY          
SEQRES   9 L  219  ALA GLY THR LYS LEU GLU LEU LYS ARG ALA ASP ALA ALA          
SEQRES  10 L  219  PRO THR VAL SER ILE PHE PRO PRO SER SER GLU GLN LEU          
SEQRES  11 L  219  THR SER GLY GLY ALA SER VAL VAL CYS PHE LEU ASN ASN          
SEQRES  12 L  219  PHE TYR PRO LYS ASP ILE ASN VAL LYS TRP LYS ILE ASP          
SEQRES  13 L  219  GLY SER GLU ARG GLN ASN GLY VAL LEU ASN SER TRP THR          
SEQRES  14 L  219  ASP GLN ASP SER LYS ASP SER THR TYR SER MET SER SER          
SEQRES  15 L  219  THR LEU THR LEU THR LYS ASP GLU TYR GLU ARG HIS ASN          
SEQRES  16 L  219  SER TYR THR CYS GLU ALA THR HIS LYS THR SER THR SER          
SEQRES  17 L  219  PRO ILE VAL LYS SER PHE ASN ARG ASN GLU CYS                  
SEQRES   1 H  227  PCA VAL GLN LEU GLN GLN PRO GLY ALA GLU LEU VAL LYS          
SEQRES   2 H  227  PRO GLY ALA SER VAL LYS LEU SER CYS LYS ALA SER GLY          
SEQRES   3 H  227  TYR THR PHE THR ASN TYR TRP ILE ASN TRP VAL LYS GLN          
SEQRES   4 H  227  ARG PRO GLY GLN GLY LEU GLU TRP ILE GLY ASN ILE TYR          
SEQRES   5 H  227  PRO GLY SER SER TYR THR HIS TYR ASN GLU LYS PHE LYS          
SEQRES   6 H  227  ASN LYS ALA THR LEU THR VAL ASP THR SER SER SER THR          
SEQRES   7 H  227  ALA TYR MET GLN LEU SER SER LEU THR SER ASP ASP SER          
SEQRES   8 H  227  ALA VAL TYR TYR CYS ALA ASN LYS LEU GLY TRP PHE PRO          
SEQRES   9 H  227  TYR TRP GLY GLN GLY THR LEU VAL THR VAL SER ALA ALA          
SEQRES  10 H  227  LYS THR THR ALA PRO SER VAL TYR PRO LEU ALA PRO VAL          
SEQRES  11 H  227  CYS GLY ASP THR SER GLY SER SER VAL THR LEU GLY CYS          
SEQRES  12 H  227  LEU VAL LYS GLY TYR PHE PRO GLU PRO VAL THR LEU THR          
SEQRES  13 H  227  TRP ASN SER GLY SER LEU SER SER GLY VAL HIS THR PHE          
SEQRES  14 H  227  PRO ALA VAL LEU GLN SER ASP LEU TYR THR LEU SER SER          
SEQRES  15 H  227  SER VAL THR VAL THR SER SER THR TRP PRO SER GLN SER          
SEQRES  16 H  227  ILE THR CYS ASN VAL ALA HIS PRO ALA SER SER THR LYS          
SEQRES  17 H  227  VAL ASP LYS LYS ILE GLU PRO ARG GLY PRO THR ILE LYS          
SEQRES  18 H  227  PRO CYS PRO PRO CYS LYS                                      
MODRES 1MJU PCA H    1  GLU  PYROGLUTAMIC ACID                                  
HET    PCA  H   1       8                                                       
HET    GOL   1401       6                                                       
HET    GOL   1402       6                                                       
HET    GOL   1403       6                                                       
HET    GOL   1404       6                                                       
HET    GOL   1405       6                                                       
HET    GOL   1406       6                                                       
HETNAM     PCA PYROGLUTAMIC ACID                                                
HETNAM     GOL GLYCEROL                                                         
FORMUL   2  PCA    C5 H7 N O3                                                   
FORMUL   3  GOL    6(C3 H8 O3)                                                  
FORMUL   9  HOH   *935(H2 O)                                                    
HELIX    1   1 GLU L   79  VAL L   83  5                                   5    
HELIX    2   2 SER L  121  THR L  126  1                                   6    
HELIX    3   3 LYS L  183  ARG L  188  1                                   6    
HELIX    4   4 THR H   28  TYR H   32  5                                   5    
HELIX    5   5 GLU H   61  LYS H   64  5                                   4    
HELIX    6   6 THR H   83  SER H   87  5                                   5    
HELIX    7   7 SER H  163  SER H  165  5                                   3    
HELIX    8   8 PRO H  213  SER H  216  5                                   4    
SHEET    1   A 4 MET L   4  THR L   5  0                                        
SHEET    2   A 4 VAL L  19  SER L  25 -1  O  ARG L  24   N  THR L   5           
SHEET    3   A 4 ALA L  70  ILE L  75 -1  O  PHE L  71   N  CYS L  23           
SHEET    4   A 4 PHE L  62  GLY L  66 -1  N  SER L  63   O  ARG L  74           
SHEET    1   B 5 ASN L  53  LEU L  54  0                                        
SHEET    2   B 5 GLN L  45  TYR L  49 -1  N  TYR L  49   O  ASN L  53           
SHEET    3   B 5 LEU L  33  GLN L  38 -1  N  LEU L  37   O  GLN L  45           
SHEET    4   B 5 GLY L  84  GLN L  90 -1  O  VAL L  85   N  GLN L  38           
SHEET    5   B 5 THR L  97  PHE L  98 -1  O  THR L  97   N  GLN L  90           
SHEET    1   C 6 ASN L  53  LEU L  54  0                                        
SHEET    2   C 6 GLN L  45  TYR L  49 -1  N  TYR L  49   O  ASN L  53           
SHEET    3   C 6 LEU L  33  GLN L  38 -1  N  LEU L  37   O  GLN L  45           
SHEET    4   C 6 GLY L  84  GLN L  90 -1  O  VAL L  85   N  GLN L  38           
SHEET    5   C 6 THR L 102  LEU L 106 -1  O  THR L 102   N  TYR L  86           
SHEET    6   C 6 SER L  10  VAL L  13  1  N  VAL L  11   O  GLU L 105           
SHEET    1   D 4 THR L 114  PHE L 118  0                                        
SHEET    2   D 4 GLY L 129  PHE L 139 -1  O  ASN L 137   N  THR L 114           
SHEET    3   D 4 TYR L 173  THR L 182 -1  O  TYR L 173   N  PHE L 139           
SHEET    4   D 4 VAL L 159  TRP L 163 -1  N  LEU L 160   O  THR L 178           
SHEET    1   E 4 SER L 153  ARG L 155  0                                        
SHEET    2   E 4 ASN L 145  ILE L 150 -1  N  ILE L 150   O  SER L 153           
SHEET    3   E 4 SER L 191  THR L 197 -1  O  GLU L 195   N  LYS L 147           
SHEET    4   E 4 ILE L 205  ASN L 210 -1  O  ILE L 205   N  ALA L 196           
SHEET    1   F 4 GLN H   3  GLN H   5  0                                        
SHEET    2   F 4 VAL H  18  SER H  25 -1  O  LYS H  23   N  GLN H   5           
SHEET    3   F 4 THR H  77  LEU H  82 -1  O  MET H  80   N  LEU H  20           
SHEET    4   F 4 ALA H  67  ASP H  72 -1  N  THR H  70   O  TYR H  79           
SHEET    1   G 5 THR H  57  TYR H  59  0                                        
SHEET    2   G 5 LEU H  45  ILE H  51 -1  N  ASN H  50   O  HIS H  58           
SHEET    3   G 5 ILE H  34  GLN H  39 -1  N  TRP H  36   O  ILE H  48           
SHEET    4   G 5 ALA H  88  LEU H  96 -1  O  ALA H  93   N  ASN H  35           
SHEET    5   G 5 PHE H  99  TRP H 103 -1  O  PHE H  99   N  LEU H  96           
SHEET    1   H 6 THR H  57  TYR H  59  0                                        
SHEET    2   H 6 LEU H  45  ILE H  51 -1  N  ASN H  50   O  HIS H  58           
SHEET    3   H 6 ILE H  34  GLN H  39 -1  N  TRP H  36   O  ILE H  48           
SHEET    4   H 6 ALA H  88  LEU H  96 -1  O  ALA H  93   N  ASN H  35           
SHEET    5   H 6 THR H 107  VAL H 111 -1  O  VAL H 109   N  ALA H  88           
SHEET    6   H 6 ALA H   9  VAL H  12  1  N  VAL H  12   O  THR H 110           
SHEET    1   I 4 SER H 120  LEU H 124  0                                        
SHEET    2   I 4 VAL H 138  TYR H 147 -1  O  LEU H 143   N  TYR H 122           
SHEET    3   I 4 LEU H 184  VAL H 193 -1  O  TYR H 185   N  TYR H 147           
SHEET    4   I 4 VAL H 171  THR H 173 -1  N  HIS H 172   O  SER H 190           
SHEET    1   J 4 SER H 120  LEU H 124  0                                        
SHEET    2   J 4 VAL H 138  TYR H 147 -1  O  LEU H 143   N  TYR H 122           
SHEET    3   J 4 LEU H 184  VAL H 193 -1  O  TYR H 185   N  TYR H 147           
SHEET    4   J 4 VAL H 177  GLN H 179 -1  N  GLN H 179   O  LEU H 184           
SHEET    1   K 3 THR H 153  TRP H 158  0                                        
SHEET    2   K 3 THR H 206  HIS H 212 -1  O  ASN H 209   N  THR H 157           
SHEET    3   K 3 THR H 217  LYS H 222 -1  O  VAL H 219   N  VAL H 210           
SSBOND   1 CYS L   23    CYS L   88                                             
SSBOND   2 CYS L  134    CYS L  194                                             
SSBOND   3 CYS L  214    CYS H  128                                             
SSBOND   4 CYS H   22    CYS H   92                                             
SSBOND   5 CYS H  142    CYS H  208                                             
CISPEP   1 TYR L   94    PRO L   95          0        -4.34                     
CISPEP   2 TYR L  140    PRO L  141          0         3.07                     
CISPEP   3 PHE H  148    PRO H  149          0        -8.52                     
CISPEP   4 GLU H  150    PRO H  151          0        -4.98                     
CISPEP   5 TRP H  199    PRO H  200          0         6.00                     
CRYST1   56.046   65.349  138.563  90.00  90.00  90.00 P 21 21 21    4          
ORIGX1      1.000000  0.000000  0.000000        0.00000                         
ORIGX2      0.000000  1.000000  0.000000        0.00000                         
ORIGX3      0.000000  0.000000  1.000000        0.00000                         
SCALE1      0.017842  0.000000  0.000000        0.00000                         
SCALE2      0.000000  0.015302  0.000000        0.00000                         
SCALE3      0.000000  0.000000  0.007217        0.00000