PDB entry 1ivb

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HEADER    HYDROLASE (O-GLYCOSYL)                  12-DEC-94   1IVB              
TITLE     STRUCTURES OF AROMATIC INHIBITORS OF INFLUENZA VIRUS                  
TITLE    2 NEURAMINIDASE                                                        
COMPND    MOL_ID: 1;                                                            
COMPND   2 MOLECULE: INFLUENZA VIRUS B/LEE/40 NEURAMINIDASE;                    
COMPND   3 CHAIN: A;                                                            
COMPND   4 EC: 3.2.1.18;                                                        
COMPND   5 ENGINEERED: YES                                                      
SOURCE    MOL_ID: 1;                                                            
SOURCE   2 ORGANISM_SCIENTIFIC: INFLUENZA B VIRUS                               
KEYWDS    HYDROLASE (O-GLYCOSYL)                                                
EXPDTA    X-RAY DIFFRACTION                                                     
AUTHOR    M.J.JEDRZEJAS,M.LUO                                                   
REVDAT   1   31-MAR-95 1IVB    0                                                
JRNL        AUTH   M.J.JEDRZEJAS,S.SINGH,W.J.BROUILLETTE,W.G.LAVER,             
JRNL        AUTH 2 G.M.AIR,M.LUO                                                
JRNL        TITL   STRUCTURES OF AROMATIC INHIBITORS OF INFLUENZA               
JRNL        TITL 2 VIRUS NEURAMINIDASE.                                         
JRNL        REF    BIOCHEMISTRY                  V.  34  3144 1995              
JRNL        REFN   ASTM BICHAW  US ISSN 0006-2960                               
REMARK   1                                                                      
REMARK   1 REFERENCE 1                                                          
REMARK   1  AUTH   S.SINGH,M.J.JEDRZEJAS,G.M.AIR,M.LUO,W.G.LAVER,               
REMARK   1  AUTH 2 W.J.BROUILLETTE                                              
REMARK   1  TITL   STRUCTURE-BASED INHIBITORS OF INFLUENZA VIRAL                
REMARK   1  TITL 2 NEURAMINIDASE. A BENZOIC ACID LEAD WITH NOVEL                
REMARK   1  TITL 3 INTERACTION                                                  
REMARK   1  REF    TO BE PUBLISHED                                              
REMARK   1  REFN                                                                
REMARK   1 REFERENCE 2                                                          
REMARK   1  AUTH   M.N.JANAKIRAMAN,C.L.WHITE,W.G.LAVER,G.M.AIR,M.LUO            
REMARK   1  TITL   STRUCTURE OF INFLUENZA VIRUS NEURAMINIDASE                   
REMARK   1  TITL 2 B(SLASH)LEE(SLASH)40 COMPLEXED WITH SIALIC ACID              
REMARK   1  TITL 3 AND A DEHYDRO ANALOG AT 1.8 ANGSTROMS RESOLUTION:            
REMARK   1  TITL 4 IMPLICATIONS FOR THE CATALYTIC MECHANISM                     
REMARK   1  REF    BIOCHEMISTRY                  V.  33  8172 1994              
REMARK   1  REFN   ASTM BICHAW  US ISSN 0006-2960                               
REMARK   1 REFERENCE 3                                                          
REMARK   1  AUTH   J.N.VARGHESE,P.M.COLMAN                                      
REMARK   1  TITL   THREE-DIMENSIONAL STRUCTURE OF THE NEURAMINIDASE             
REMARK   1  TITL 2 OF INFLUENZA VIRUS A(SLASH)TOKYO(SLASH)3(SLASH)67            
REMARK   1  TITL 3 AT 2.2 ANGSTROMS RESOLUTION                                  
REMARK   1  REF    J.MOL.BIOL.                   V. 221   473 1991              
REMARK   1  REFN   ASTM JMOBAK  UK ISSN 0022-2836                               
REMARK   1 REFERENCE 4                                                          
REMARK   1  AUTH   W.R.TULIP,J.N.VARGHESE,A.T.BAKER,A.VAN DONKELAAR,            
REMARK   1  AUTH 2 W.G.LAVER,R.G.WEBSTER,P.M.COLMAN                             
REMARK   1  TITL   REFINED ATOMIC STRUCTURES OF N9 SUBTYPE INFLUENZA            
REMARK   1  TITL 2 VIRUS NEURAMINIDASE AND ESCAPE MUTANTS                       
REMARK   1  REF    J.MOL.BIOL.                   V. 221   487 1991              
REMARK   1  REFN   ASTM JMOBAK  UK ISSN 0022-2836                               
REMARK   2                                                                      
REMARK   2 RESOLUTION. 2.40 ANGSTROMS.                                          
REMARK   3                                                                      
REMARK   3 REFINEMENT.                                                          
REMARK   3   PROGRAM     : X-PLOR                                               
REMARK   3   AUTHORS     : BRUNGER                                              
REMARK   3                                                                      
REMARK   3  DATA USED IN REFINEMENT.                                            
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.40                           
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 6.50                           
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 4.000                          
REMARK   3   DATA CUTOFF HIGH         (ABS(F)) : NULL                           
REMARK   3   DATA CUTOFF LOW          (ABS(F)) : NULL                           
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : NULL                           
REMARK   3   NUMBER OF REFLECTIONS             : 14129                          
REMARK   3                                                                      
REMARK   3  FIT TO DATA USED IN REFINEMENT.                                     
REMARK   3   CROSS-VALIDATION METHOD          : NULL                            
REMARK   3   FREE R VALUE TEST SET SELECTION  : NULL                            
REMARK   3   R VALUE            (WORKING SET) : 0.189                           
REMARK   3   FREE R VALUE                     : NULL                            
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : NULL                            
REMARK   3   FREE R VALUE TEST SET COUNT      : NULL                            
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : NULL                            
REMARK   3                                                                      
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.                                  
REMARK   3   TOTAL NUMBER OF BINS USED           : NULL                         
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : NULL                         
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : NULL                         
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : NULL                         
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : NULL                         
REMARK   3   BIN R VALUE           (WORKING SET) : NULL                         
REMARK   3   BIN FREE R VALUE                    : NULL                         
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : NULL                         
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : NULL                         
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : NULL                         
REMARK   3                                                                      
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.                    
REMARK   3   PROTEIN ATOMS            : 3736                                    
REMARK   3   NUCLEIC ACID ATOMS       : 0                                       
REMARK   3   HETEROGEN ATOMS          : 49                                      
REMARK   3   SOLVENT ATOMS            : 0                                       
REMARK   3                                                                      
REMARK   3  B VALUES.                                                           
REMARK   3   FROM WILSON PLOT           (A**2) : NULL                           
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : NULL                           
REMARK   3   OVERALL ANISOTROPIC B VALUE.                                       
REMARK   3    B11 (A**2) : NULL                                                 
REMARK   3    B22 (A**2) : NULL                                                 
REMARK   3    B33 (A**2) : NULL                                                 
REMARK   3    B12 (A**2) : NULL                                                 
REMARK   3    B13 (A**2) : NULL                                                 
REMARK   3    B23 (A**2) : NULL                                                 
REMARK   3                                                                      
REMARK   3  ESTIMATED COORDINATE ERROR.                                         
REMARK   3   ESD FROM LUZZATI PLOT        (A) : NULL                            
REMARK   3   ESD FROM SIGMAA              (A) : NULL                            
REMARK   3   LOW RESOLUTION CUTOFF        (A) : NULL                            
REMARK   3                                                                      
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.                         
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : NULL                            
REMARK   3   ESD FROM C-V SIGMAA          (A) : NULL                            
REMARK   3                                                                      
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.                                   
REMARK   3   BOND LENGTHS                 (A) : 0.016                           
REMARK   3   BOND ANGLES            (DEGREES) : 2.14                            
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : NULL                            
REMARK   3   IMPROPER ANGLES        (DEGREES) : NULL                            
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL MODEL : NULL                                      
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA                
REMARK   3   MAIN-CHAIN BOND              (A**2) : NULL  ; NULL                 
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : NULL  ; NULL                 
REMARK   3   SIDE-CHAIN BOND              (A**2) : NULL  ; NULL                 
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : NULL  ; NULL                 
REMARK   3                                                                      
REMARK   3  NCS MODEL : NULL                                                    
REMARK   3                                                                      
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT          
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL                 
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL                 
REMARK   3                                                                      
REMARK   3  PARAMETER FILE  1  : NULL                                           
REMARK   3  TOPOLOGY FILE  1   : NULL                                           
REMARK   3                                                                      
REMARK   3  OTHER REFINEMENT REMARKS: NULL                                      
REMARK   4                                                                      
REMARK   4 1IVB COMPLIES WITH FORMAT V. 3.0, 1-DEC-2006                         
REMARK   4                                                                      
REMARK   4 THIS IS THE REMEDIATED VERSION OF THIS PDB ENTRY.                    
REMARK   4 REMEDIATED DATA FILE REVISION 3.100 (2007-03-17)                     
REMARK 200                                                                      
REMARK 200 EXPERIMENTAL DETAILS                                                 
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION                  
REMARK 200  DATE OF DATA COLLECTION        : NULL                               
REMARK 200  TEMPERATURE           (KELVIN) : NULL                               
REMARK 200  PH                             : NULL                               
REMARK 200  NUMBER OF CRYSTALS USED        : NULL                               
REMARK 200                                                                      
REMARK 200  SYNCHROTRON              (Y/N) : NULL                               
REMARK 200  RADIATION SOURCE               : NULL                               
REMARK 200  BEAMLINE                       : NULL                               
REMARK 200  X-RAY GENERATOR MODEL          : NULL                               
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : NULL                               
REMARK 200  WAVELENGTH OR RANGE        (A) : NULL                               
REMARK 200  MONOCHROMATOR                  : NULL                               
REMARK 200  OPTICS                         : NULL                               
REMARK 200                                                                      
REMARK 200  DETECTOR TYPE                  : NULL                               
REMARK 200  DETECTOR MANUFACTURER          : NULL                               
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : NULL                               
REMARK 200  DATA SCALING SOFTWARE          : NULL                               
REMARK 200                                                                      
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 15271                              
REMARK 200  RESOLUTION RANGE HIGH      (A) : NULL                               
REMARK 200  RESOLUTION RANGE LOW       (A) : NULL                               
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 1.000                              
REMARK 200                                                                      
REMARK 200 OVERALL.                                                             
REMARK 200  COMPLETENESS FOR RANGE     (%) : NULL                               
REMARK 200  DATA REDUNDANCY                : NULL                               
REMARK 200  R MERGE                    (I) : NULL                               
REMARK 200  R SYM                      (I) : NULL                               
REMARK 200   FOR THE DATA SET  : NULL                               
REMARK 200                                                                      
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.                                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : NULL                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : NULL                     
REMARK 200  COMPLETENESS FOR SHELL     (%) : NULL                               
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL                               
REMARK 200  R MERGE FOR SHELL          (I) : NULL                               
REMARK 200  R SYM FOR SHELL            (I) : NULL                               
REMARK 200   FOR SHELL         : NULL                               
REMARK 200                                                                      
REMARK 200 DIFFRACTION PROTOCOL: NULL                                           
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: NULL                         
REMARK 200 SOFTWARE USED: X-PLOR                                                
REMARK 200 STARTING MODEL: NULL                                                 
REMARK 200                                                                      
REMARK 200 REMARK: NULL                                                         
REMARK 280                                                                      
REMARK 280 CRYSTAL                                                              
REMARK 280 SOLVENT CONTENT, VS   (%): 61.59                                     
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 3.20                     
REMARK 280                                                                      
REMARK 280 CRYSTALLIZATION CONDITIONS: NULL                                     
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY                                            
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 4 21 2                         
REMARK 290                                                                      
REMARK 290      SYMOP   SYMMETRY                                                
REMARK 290     NNNMMM   OPERATOR                                                
REMARK 290       1555   X,Y,Z                                                   
REMARK 290       2555   -X,-Y,Z                                                 
REMARK 290       3555   1/2-Y,1/2+X,Z                                           
REMARK 290       4555   1/2+Y,1/2-X,Z                                           
REMARK 290       5555   1/2-X,1/2+Y,-Z                                          
REMARK 290       6555   1/2+X,1/2-Y,-Z                                          
REMARK 290       7555   Y,X,-Z                                                  
REMARK 290       8555   -Y,-X,-Z                                                
REMARK 290                                                                      
REMARK 290     WHERE NNN -> OPERATOR NUMBER                                     
REMARK 290           MMM -> TRANSLATION VECTOR                                  
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS                            
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM             
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY                
REMARK 290 RELATED MOLECULES.                                                   
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   3  0.000000 -1.000000  0.000000       62.27000            
REMARK 290   SMTRY2   3  1.000000  0.000000  0.000000       62.27000            
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   4  0.000000  1.000000  0.000000       62.27000            
REMARK 290   SMTRY2   4 -1.000000  0.000000  0.000000       62.27000            
REMARK 290   SMTRY3   4  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   5 -1.000000  0.000000  0.000000       62.27000            
REMARK 290   SMTRY2   5  0.000000  1.000000  0.000000       62.27000            
REMARK 290   SMTRY3   5  0.000000  0.000000 -1.000000        0.00000            
REMARK 290   SMTRY1   6  1.000000  0.000000  0.000000       62.27000            
REMARK 290   SMTRY2   6  0.000000 -1.000000  0.000000       62.27000            
REMARK 290   SMTRY3   6  0.000000  0.000000 -1.000000        0.00000            
REMARK 290   SMTRY1   7  0.000000  1.000000  0.000000        0.00000            
REMARK 290   SMTRY2   7  1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY3   7  0.000000  0.000000 -1.000000        0.00000            
REMARK 290   SMTRY1   8  0.000000 -1.000000  0.000000        0.00000            
REMARK 290   SMTRY2   8 -1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY3   8  0.000000  0.000000 -1.000000        0.00000            
REMARK 290                                                                      
REMARK 290 REMARK: NULL                                                         
REMARK 300                                                                      
REMARK 300 BIOMOLECULE: 1                                                       
REMARK 300 THIS ENTRY CONTAINS THE CRYSTALLOGRAPHIC ASYMMETRIC UNIT             
REMARK 300 WHICH CONSISTS OF 1 CHAIN(S). SEE REMARK 350 FOR                     
REMARK 300 INFORMATION ON GENERATING THE BIOLOGICAL MOLECULE(S).                
REMARK 350                                                                      
REMARK 350 GENERATING THE BIOMOLECULE                                           
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN           
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE                
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS          
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND                          
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.                               
REMARK 350                                                                      
REMARK 350 BIOMOLECULE: 1                                                       
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A                                     
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 350   BIOMT1   2 -1.000000  0.000000  0.000000      124.54000            
REMARK 350   BIOMT2   2  0.000000 -1.000000  0.000000        0.00000            
REMARK 350   BIOMT3   2  0.000000  0.000000  1.000000        0.00000            
REMARK 350   BIOMT1   3  0.000000 -1.000000  0.000000       62.27000            
REMARK 350   BIOMT2   3  1.000000  0.000000  0.000000      -62.27000            
REMARK 350   BIOMT3   3  0.000000  0.000000  1.000000        0.00000            
REMARK 350   BIOMT1   4  0.000000  1.000000  0.000000       62.27000            
REMARK 350   BIOMT2   4 -1.000000  0.000000  0.000000       62.27000            
REMARK 350   BIOMT3   4  0.000000  0.000000  1.000000        0.00000            
REMARK 375                                                                      
REMARK 375 SPECIAL POSITION                                                     
REMARK 375 THE FOLLOWING ATOMS ARE FOUND TO BE WITHIN 0.15 ANGSTROMS            
REMARK 375 OF A SYMMETRY RELATED ATOM AND ARE ASSUMED TO BE ON SPECIAL          
REMARK 375 POSITIONS.                                                           
REMARK 375                                                                      
REMARK 375 ATOM RES CSSEQI                                                      
REMARK 375 CA    CA   501   LIES ON A SPECIAL POSITION.                         
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND LENGTHS                                      
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,2(A3,1X,A1,I4,A1,1X,A4,3X),F6.3)                  
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   RES CSSEQI ATM2   DEVIATION                     
REMARK 500    MET A 239   SD    MET A 239   CE     0.108                        
REMARK 500    MET A 357   SD    MET A 357   CE     0.107                        
REMARK 500    MET A 429   SD    MET A 429   CE     0.109                        
REMARK 500    MET A 464   SD    MET A 464   CE     0.132                        
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES                                       
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)              
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3                                     
REMARK 500    THR A 189   N   -  CA  -  C   ANGL. DEV. =-13.9 DEGREES           
REMARK 500    ALA A 290   N   -  CA  -  C   ANGL. DEV. =-17.4 DEGREES           
REMARK 500    TYR A 296   N   -  CA  -  C   ANGL. DEV. = 16.4 DEGREES           
REMARK 500    GLN A 355   N   -  CA  -  C   ANGL. DEV. =-12.8 DEGREES           
REMARK 500    LEU A 379   CA  -  CB  -  CG  ANGL. DEV. = 21.8 DEGREES           
REMARK 500    LYS A 435   N   -  CA  -  C   ANGL. DEV. = 13.9 DEGREES           
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: TORSION ANGLES                                             
REMARK 500                                                                      
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:            
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;               
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                             
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)                    
REMARK 500                                                                      
REMARK 500  M RES CSSEQI        PSI       PHI                                   
REMARK 500    SER A 283     -169.54    148.83                                   
DBREF  1IVB A   77   466  UNP    P03474   NRAM_INBLE      77    466             
SEQADV 1IVB ARG A  382  UNP  P03474    LYS   382 CONFLICT                       
SEQRES   1 A  390  GLU PRO GLU TRP THR TYR PRO ARG LEU SER CYS GLN GLY          
SEQRES   2 A  390  SER THR PHE GLN LYS ALA LEU LEU ILE SER PRO HIS ARG          
SEQRES   3 A  390  PHE GLY GLU ILE LYS GLY ASN SER ALA PRO LEU ILE ILE          
SEQRES   4 A  390  ARG GLU PRO PHE VAL ALA CYS GLY PRO LYS GLU CYS ARG          
SEQRES   5 A  390  HIS PHE ALA LEU THR HIS TYR ALA ALA GLN PRO GLY GLY          
SEQRES   6 A  390  TYR TYR ASN GLY THR ARG LYS ASP ARG ASN LYS LEU ARG          
SEQRES   7 A  390  HIS LEU VAL SER VAL LYS LEU GLY LYS ILE PRO THR VAL          
SEQRES   8 A  390  GLU ASN SER ILE PHE HIS MET ALA ALA TRP SER GLY SER          
SEQRES   9 A  390  ALA CYS HIS ASP GLY ARG GLU TRP THR TYR ILE GLY VAL          
SEQRES  10 A  390  ASP GLY PRO ASP ASN ASP ALA LEU VAL LYS ILE LYS TYR          
SEQRES  11 A  390  GLY GLU ALA TYR THR ASP THR TYR HIS SER TYR ALA HIS          
SEQRES  12 A  390  ASN ILE LEU ARG THR GLN GLU SER ALA CYS ASN CYS ILE          
SEQRES  13 A  390  GLY GLY ASP CYS TYR LEU MET ILE THR ASP GLY SER ALA          
SEQRES  14 A  390  SER GLY ILE SER LYS CYS ARG PHE LEU LYS ILE ARG GLU          
SEQRES  15 A  390  GLY ARG ILE ILE LYS GLU ILE LEU PRO THR GLY ARG VAL          
SEQRES  16 A  390  GLU HIS THR GLU GLU CYS THR CYS GLY PHE ALA SER ASN          
SEQRES  17 A  390  LYS THR ILE GLU CYS ALA CYS ARG ASP ASN SER TYR THR          
SEQRES  18 A  390  ALA LYS ARG PRO PHE VAL LYS LEU ASN VAL GLU THR ASP          
SEQRES  19 A  390  THR ALA GLU ILE ARG LEU MET CYS THR LYS THR TYR LEU          
SEQRES  20 A  390  ASP THR PRO ARG PRO ASP ASP GLY SER ILE ALA GLY PRO          
SEQRES  21 A  390  CYS GLU SER ASN GLY ASP LYS TRP LEU GLY GLY ILE LYS          
SEQRES  22 A  390  GLY GLY PHE VAL HIS GLN ARG MET ALA SER LYS ILE GLY          
SEQRES  23 A  390  ARG TRP TYR SER ARG THR MET SER LYS THR ASN ARG MET          
SEQRES  24 A  390  GLY MET GLU LEU TYR VAL ARG TYR ASP GLY ASP PRO TRP          
SEQRES  25 A  390  THR ASP SER ASP ALA LEU THR LEU SER GLY VAL MET VAL          
SEQRES  26 A  390  SER ILE GLU GLU PRO GLY TRP TYR SER PHE GLY PHE GLU          
SEQRES  27 A  390  ILE LYS ASP LYS LYS CYS ASP VAL PRO CYS ILE GLY ILE          
SEQRES  28 A  390  GLU MET VAL HIS ASP GLY GLY LYS ASP THR TRP HIS SER          
SEQRES  29 A  390  ALA ALA THR ALA ILE TYR CYS LEU MET GLY SER GLY GLN          
SEQRES  30 A  390  LEU LEU TRP ASP THR VAL THR GLY VAL ASP MET ALA LEU          
MODRES 1IVB ASN A  284  ASN  GLYCOSYLATION SITE                                 
FTNOTE   1 CIS PROLINE - PRO 139                                                
FTNOTE   2 CIS PROLINE - PRO 326                                                
HET    NAG    467      28                                                       
HET     CA    500       1                                                       
HET     CA    501       1                                                       
HET    ST1    471      19                                                       
HETNAM     NAG N-ACETYL-D-GLUCOSAMINE                                           
HETNAM      CA CALCIUM ION                                                      
HETNAM     ST1 4-(ACETYLAMINO)-3-HYDROXY-5-NITROBENZOIC ACID                    
HETSYN     NAG NAG                                                              
FORMUL   2  NAG    C8 H15 N O6                                                  
FORMUL   3   CA    2(CA 2+)                                                     
FORMUL   5  ST1    C9 H8 N2 O6                                                  
HELIX    1   1 PRO A  100  PHE A  103  5                                   4    
SHEET    1   A 4 PHE A 119  CYS A 122  0                                        
SHEET    2   A 4 CYS A 127  THR A 133 -1  N  PHE A 130   O  PHE A 119           
SHEET    3   A 4 HIS A 155  LYS A 160 -1  N  VAL A 159   O  HIS A 129           
SHEET    4   A 4 ILE A 171  ALA A 175 -1  N  MET A 174   O  LEU A 156           
SHEET    1   B 4 SER A 178  HIS A 183  0                                        
SHEET    2   B 4 TRP A 188  PRO A 196 -1  N  VAL A 193   O  SER A 178           
SHEET    3   B 4 ASP A 199  TYR A 206 -1  N  LYS A 205   O  TYR A 190           
SHEET    4   B 4 ALA A 209  HIS A 215 -1  N  TYR A 214   O  VAL A 202           
SHEET    1   C 4 ASN A 230  ILE A 232  0                                        
SHEET    2   C 4 ASP A 235  ILE A 240 -1  N  TYR A 237   O  ASN A 230           
SHEET    3   C 4 ARG A 252  ILE A 256 -1  N  ILE A 256   O  CYS A 236           
SHEET    4   C 4 ILE A 261  LEU A 266 -1  N  ILE A 265   O  PHE A 253           
SHEET    1   D 4 THR A 278  PHE A 281  0                                        
SHEET    2   D 4 THR A 286  ALA A 290 -1  N  ALA A 290   O  THR A 278           
SHEET    3   D 4 PRO A 301  ASN A 306 -1  N  LEU A 305   O  ILE A 287           
SHEET    4   D 4 THR A 311  LEU A 316 -1  N  ARG A 315   O  PHE A 302           
SHEET    1   E 3 PHE A 352  ARG A 356  0                                        
SHEET    2   E 3 ILE A 361  ARG A 367 -1  N  TRP A 364   O  VAL A 353           
SHEET    3   E 3 GLU A 378  TYR A 383 -1  N  ARG A 382   O  ARG A 363           
SHEET    1   F 4 SER A 410  LYS A 416  0                                        
SHEET    2   F 4 ASP A 421  ASP A 432 -1  N  GLY A 426   O  PHE A 411           
SHEET    3   F 4 HIS A 439  LEU A 448 -1  N  TYR A 446   O  ILE A 425           
SHEET    4   F 4 PHE A  92  ILE A  98 -1  N  ILE A  98   O  THR A 443           
SSBOND   1 CYS A   87    CYS A  420                                             
SSBOND   2 CYS A  122    CYS A  127                                             
SSBOND   3 CYS A  182    CYS A  229                                             
SSBOND   4 CYS A  231    CYS A  236                                             
SSBOND   5 CYS A  277    CYS A  291                                             
SSBOND   6 CYS A  279    CYS A  289                                             
SSBOND   7 CYS A  318    CYS A  337                                             
SSBOND   8 CYS A  424    CYS A  447                                             
LINK         ND2 ASN A 284                 C1  NAG   467                        
CISPEP   1 GLN A  138    PRO A  139          0         0.00                     
CISPEP   2 THR A  325    PRO A  326          0         0.49                     
SITE     1 CAT 11 ARG A 116  GLU A 117  ASP A 149  ARG A 150                    
SITE     2 CAT 11 TRP A 177  ILE A 221  ARG A 223  GLU A 275                    
SITE     3 CAT 11 ARG A 292  ARG A 374  TYR A 409                               
CRYST1  124.540  124.540   71.820  90.00  90.00  90.00 P 4 21 2      8          
ORIGX1      1.000000  0.000000  0.000000        0.00000                         
ORIGX2      0.000000  1.000000  0.000000        0.00000                         
ORIGX3      0.000000  0.000000  1.000000        0.00000                         
SCALE1      0.008030  0.000000  0.000000        0.00000                         
SCALE2      0.000000  0.008030  0.000000        0.00000                         
SCALE3      0.000000  0.000000  0.013924        0.00000