PDB entry 1gwb

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HEADER    GLYCOPROTEIN                            14-MAR-02   1GWB              
TITLE     STRUCTURE OF GLYCOPROTEIN 1B                                          
COMPND    MOL_ID: 1;                                                            
COMPND   2 MOLECULE: PLATELET GLYCOPROTEIN IB ALPHA CHAIN;                      
COMPND   3 CHAIN: A;                                                            
COMPND   4 FRAGMENT: N-TERMINAL DOMAIN, RESIDUES 16-296;                        
COMPND   5 SYNONYM: GP-IB ALPHA, GPIBA, CD42B-ALPHA, CD42B;                     
COMPND   6 MOL_ID: 2;                                                           
COMPND   7 MOLECULE:;                                                           
COMPND   8 CHAIN: B;                                                            
COMPND   9 FRAGMENT: N-TERMINAL DOMAIN, RESIDUES 16-296;                        
COMPND  10 ENGINEERED: YES;                                                     
COMPND  11 OTHER_DETAILS: TYR 292,294,295 MODIFIED TO TYROSINE-O-               
COMPND  12 SULPHONIC ACID                                                       
SOURCE    MOL_ID: 1;                                                            
SOURCE   2 ORGANISM_SCIENTIFIC: HOMO SAPIENS;                                   
SOURCE   3 MOL_ID: 2                                                            
KEYWDS    GLYCOPROTEIN 1B, PLATELET, TRANSMEMBRANE, GLYCOPROTEIN,               
KEYWDS   2 HEMOSTASIS, BLOOD COAGULATION, REPEAT, LEUCINE-RICH REPEAT,          
KEYWDS   3 SIGNAL, CELL ADHESION, DISEASE MUTATION, POLYMORPHISM, VON           
KEYWDS   4 WILLEBRAND DISEASE, BERNARD SOULIER SYNDROME                         
EXPDTA    X-RAY DIFFRACTION                                                     
AUTHOR    J.EMSLEY,S.UFF,K.CLEMETSON,J.CLEMETSON,T.HARRISON                     
REVDAT   1   06-FEB-03 1GWB    0                                                
JRNL        AUTH   S.UFF,J.M.CLEMETSON,T.HARRISON,K.J.CLEMETSON,                
JRNL        AUTH 2 J.EMSLEY                                                     
JRNL        TITL   CRYSTAL STRUCTURE OF THE PLATELET GLYCOPROTEIN               
JRNL        TITL 2 IB(ALPHA) N-TERMINAL DOMAIN REVEALS AN UNMASKING             
JRNL        TITL 3 MECHANISM FOR RECEPTOR ACTIVATION.                           
JRNL        REF    J.BIOL.CHEM.                  V. 277 35657 2002              
JRNL        REFN   ASTM JBCHA3  US ISSN 0021-9258                               
REMARK   1                                                                      
REMARK   2                                                                      
REMARK   2 RESOLUTION. 2.80 ANGSTROMS.                                          
REMARK   3                                                                      
REMARK   3 REFINEMENT.                                                          
REMARK   3   PROGRAM     : CNS 1.1                                              
REMARK   3   AUTHORS     : BRUNGER,ADAMS,CLORE,DELANO,GROS,GROSSE-              
REMARK   3               : KUNSTLEVE,JIANG,KUSZEWSKI,NILGES, PANNU,             
REMARK   3               : READ,RICE,SIMONSON,WARREN                            
REMARK   3                                                                      
REMARK   3  REFINEMENT TARGET : NULL                                            
REMARK   3                                                                      
REMARK   3  DATA USED IN REFINEMENT.                                            
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.80                           
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 15.00                          
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 1.000                          
REMARK   3   DATA CUTOFF HIGH         (ABS(F)) : NULL                           
REMARK   3   DATA CUTOFF LOW          (ABS(F)) : NULL                           
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : 98.5                           
REMARK   3   NUMBER OF REFLECTIONS             : 37462                          
REMARK   3                                                                      
REMARK   3  FIT TO DATA USED IN REFINEMENT.                                     
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT                      
REMARK   3   FREE R VALUE TEST SET SELECTION  : NULL                            
REMARK   3   R VALUE            (WORKING SET) : 0.245                           
REMARK   3   FREE R VALUE                     : 0.274                           
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.000                           
REMARK   3   FREE R VALUE TEST SET COUNT      : NULL                            
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : NULL                            
REMARK   3                                                                      
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.                                  
REMARK   3   TOTAL NUMBER OF BINS USED           : NULL                         
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : NULL                         
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : NULL                         
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : NULL                         
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : NULL                         
REMARK   3   BIN R VALUE           (WORKING SET) : NULL                         
REMARK   3   BIN FREE R VALUE                    : NULL                         
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : NULL                         
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : NULL                         
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : NULL                         
REMARK   3                                                                      
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.                    
REMARK   3   PROTEIN ATOMS            : 4205                                    
REMARK   3   NUCLEIC ACID ATOMS       : 0                                       
REMARK   3   HETEROGEN ATOMS          : 103                                     
REMARK   3   SOLVENT ATOMS            : NULL                                    
REMARK   3                                                                      
REMARK   3  B VALUES.                                                           
REMARK   3   FROM WILSON PLOT           (A**2) : NULL                           
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : NULL                           
REMARK   3   OVERALL ANISOTROPIC B VALUE.                                       
REMARK   3    B11 (A**2) : NULL                                                 
REMARK   3    B22 (A**2) : NULL                                                 
REMARK   3    B33 (A**2) : NULL                                                 
REMARK   3    B12 (A**2) : NULL                                                 
REMARK   3    B13 (A**2) : NULL                                                 
REMARK   3    B23 (A**2) : NULL                                                 
REMARK   3                                                                      
REMARK   3  ESTIMATED COORDINATE ERROR.                                         
REMARK   3   ESD FROM LUZZATI PLOT        (A) : NULL                            
REMARK   3   ESD FROM SIGMAA              (A) : NULL                            
REMARK   3   LOW RESOLUTION CUTOFF        (A) : NULL                            
REMARK   3                                                                      
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.                         
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : NULL                            
REMARK   3   ESD FROM C-V SIGMAA          (A) : NULL                            
REMARK   3                                                                      
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.                                   
REMARK   3   BOND LENGTHS                 (A) : 0.008                           
REMARK   3   BOND ANGLES            (DEGREES) : 1.84                            
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : NULL                            
REMARK   3   IMPROPER ANGLES        (DEGREES) : NULL                            
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL MODEL : NULL                                      
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA                
REMARK   3   MAIN-CHAIN BOND              (A**2) : NULL  ; NULL                 
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : NULL  ; NULL                 
REMARK   3   SIDE-CHAIN BOND              (A**2) : NULL  ; NULL                 
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : NULL  ; NULL                 
REMARK   3                                                                      
REMARK   3  BULK SOLVENT MODELING.                                              
REMARK   3   METHOD USED : NULL                                                 
REMARK   3   KSOL        : NULL                                                 
REMARK   3   BSOL        : NULL                                                 
REMARK   3                                                                      
REMARK   3  NCS MODEL : NULL                                                    
REMARK   3                                                                      
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT          
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL                 
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL                 
REMARK   3                                                                      
REMARK   3  PARAMETER FILE  1  : NULL                                           
REMARK   3  TOPOLOGY FILE  1   : NULL                                           
REMARK   3                                                                      
REMARK   3  OTHER REFINEMENT REMARKS: NULL                                      
REMARK   4                                                                      
REMARK   4 1GWB COMPLIES WITH FORMAT V. 3.0, 1-DEC-2006                         
REMARK   4                                                                      
REMARK   4 THIS IS THE REMEDIATED VERSION OF THIS PDB ENTRY.                    
REMARK   4 REMEDIATED DATA FILE REVISION 3.101 (2007-05-31)                     
REMARK 100                                                                      
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY EBI  .                              
REMARK 100 THE EBI ID CODE IS  EBI-9557  .                                      
REMARK 200                                                                      
REMARK 200 EXPERIMENTAL DETAILS                                                 
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION                  
REMARK 200  DATE OF DATA COLLECTION        : NULL                               
REMARK 200  TEMPERATURE           (KELVIN) : 100.0                              
REMARK 200  PH                             : 6.50                               
REMARK 200  NUMBER OF CRYSTALS USED        : 10                                 
REMARK 200                                                                      
REMARK 200  SYNCHROTRON              (Y/N) : Y                                  
REMARK 200  RADIATION SOURCE               : ESRF                               
REMARK 200  BEAMLINE                       : ID14-4                             
REMARK 200  X-RAY GENERATOR MODEL          : NULL                               
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M                                  
REMARK 200  WAVELENGTH OR RANGE        (A) : 1                                  
REMARK 200  MONOCHROMATOR                  : NULL                               
REMARK 200  OPTICS                         : NULL                               
REMARK 200                                                                      
REMARK 200  DETECTOR TYPE                  : NULL                               
REMARK 200  DETECTOR MANUFACTURER          : NULL                               
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : NULL                               
REMARK 200  DATA SCALING SOFTWARE          : NULL                               
REMARK 200                                                                      
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 37462                              
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.800                              
REMARK 200  RESOLUTION RANGE LOW       (A) : 20.000                             
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 0.000                              
REMARK 200                                                                      
REMARK 200 OVERALL.                                                             
REMARK 200  COMPLETENESS FOR RANGE     (%) : 98.9                               
REMARK 200  DATA REDUNDANCY                : 4.300                              
REMARK 200  R MERGE                    (I) : 0.09900                            
REMARK 200  R SYM                      (I) : NULL                               
REMARK 200   FOR THE DATA SET  : 9.1000                             
REMARK 200                                                                      
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.                                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : NULL                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : NULL                     
REMARK 200  COMPLETENESS FOR SHELL     (%) : NULL                               
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL                               
REMARK 200  R MERGE FOR SHELL          (I) : NULL                               
REMARK 200  R SYM FOR SHELL            (I) : NULL                               
REMARK 200   FOR SHELL         : NULL                               
REMARK 200                                                                      
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH                              
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MIR                          
REMARK 200 SOFTWARE USED: NULL                                                  
REMARK 200 STARTING MODEL: NULL                                                 
REMARK 200                                                                      
REMARK 200 REMARK: NULL                                                         
REMARK 280                                                                      
REMARK 280 CRYSTAL                                                              
REMARK 280 SOLVENT CONTENT, VS   (%): NULL                                      
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): NULL                     
REMARK 280                                                                      
REMARK 280 CRYSTALLIZATION CONDITIONS: PH 6.5                                   
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY                                            
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 64 2 2                         
REMARK 290                                                                      
REMARK 290      SYMOP   SYMMETRY                                                
REMARK 290     NNNMMM   OPERATOR                                                
REMARK 290       1555   X,Y,Z                                                   
REMARK 290       2555   -Y,X-Y,1/3+Z                                            
REMARK 290       3555   -X+Y,-X,2/3+Z                                           
REMARK 290       4555   -X,-Y,Z                                                 
REMARK 290       5555   Y,-X+Y,1/3+Z                                            
REMARK 290       6555   X-Y,X,2/3+Z                                             
REMARK 290       7555   Y,X,1/3-Z                                               
REMARK 290       8555   X-Y,-Y,-Z                                               
REMARK 290       9555   -X,-X+Y,2/3-Z                                           
REMARK 290      10555   -Y,-X,1/3-Z                                             
REMARK 290      11555   -X+Y,Y,-Z                                               
REMARK 290      12555   X,X-Y,2/3-Z                                             
REMARK 290                                                                      
REMARK 290     WHERE NNN -> OPERATOR NUMBER                                     
REMARK 290           MMM -> TRANSLATION VECTOR                                  
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS                            
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM             
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY                
REMARK 290 RELATED MOLECULES.                                                   
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   2 -0.500000 -0.866025  0.000000        0.00000            
REMARK 290   SMTRY2   2  0.866025 -0.500000  0.000000        0.00000            
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       42.66667            
REMARK 290   SMTRY1   3 -0.500000  0.866025  0.000000        0.00000            
REMARK 290   SMTRY2   3 -0.866025 -0.500000  0.000000        0.00000            
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000       85.33333            
REMARK 290   SMTRY1   4 -1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   4  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   5  0.500000  0.866025  0.000000        0.00000            
REMARK 290   SMTRY2   5 -0.866025  0.500000  0.000000        0.00000            
REMARK 290   SMTRY3   5  0.000000  0.000000  1.000000       42.66667            
REMARK 290   SMTRY1   6  0.500000 -0.866025  0.000000        0.00000            
REMARK 290   SMTRY2   6  0.866025  0.500000  0.000000        0.00000            
REMARK 290   SMTRY3   6  0.000000  0.000000  1.000000       85.33333            
REMARK 290   SMTRY1   7 -0.500000  0.866025  0.000000        0.00000            
REMARK 290   SMTRY2   7  0.866025  0.500000  0.000000        0.00000            
REMARK 290   SMTRY3   7  0.000000  0.000000 -1.000000       42.66667            
REMARK 290   SMTRY1   8  1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   8  0.000000 -1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   8  0.000000  0.000000 -1.000000        0.00000            
REMARK 290   SMTRY1   9 -0.500000 -0.866025  0.000000        0.00000            
REMARK 290   SMTRY2   9 -0.866025  0.500000  0.000000        0.00000            
REMARK 290   SMTRY3   9  0.000000  0.000000 -1.000000       85.33333            
REMARK 290   SMTRY1  10  0.500000 -0.866025  0.000000        0.00000            
REMARK 290   SMTRY2  10 -0.866025 -0.500000  0.000000        0.00000            
REMARK 290   SMTRY3  10  0.000000  0.000000 -1.000000       42.66667            
REMARK 290   SMTRY1  11 -1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2  11  0.000000  1.000000  0.000000        0.00000            
REMARK 290   SMTRY3  11  0.000000  0.000000 -1.000000        0.00000            
REMARK 290   SMTRY1  12  0.500000  0.866025  0.000000        0.00000            
REMARK 290   SMTRY2  12  0.866025 -0.500000  0.000000        0.00000            
REMARK 290   SMTRY3  12  0.000000  0.000000 -1.000000       85.33333            
REMARK 290                                                                      
REMARK 290 REMARK: NULL                                                         
REMARK 300                                                                      
REMARK 300 BIOMOLECULE: 1                                                       
REMARK 300 THIS ENTRY CONTAINS THE CRYSTALLOGRAPHIC ASYMMETRIC UNIT             
REMARK 300 WHICH CONSISTS OF 2 CHAIN(S). SEE REMARK 350 FOR                     
REMARK 300 INFORMATION ON GENERATING THE BIOLOGICAL MOLECULE(S).                
REMARK 350                                                                      
REMARK 350 GENERATING THE BIOMOLECULE                                           
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN           
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE                
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS          
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND                          
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.                               
REMARK 350                                                                      
REMARK 350 BIOMOLECULE: 1                                                       
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B, Y, Z                            
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 465                                                                      
REMARK 465 MISSING RESIDUES                                                     
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE                       
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)                
REMARK 465                                                                      
REMARK 465   M RES C SSSEQI                                                     
REMARK 465     PRO A    16                                                      
REMARK 465     HIS A    17                                                      
REMARK 465     GLY A   284                                                      
REMARK 465     ASP A   285                                                      
REMARK 465     GLU A   286                                                      
REMARK 465     GLY A   287                                                      
REMARK 465     ASP A   288                                                      
REMARK 465     THR A   289                                                      
REMARK 465     ASP A   290                                                      
REMARK 465     LEU A   291                                                      
REMARK 465     STY A   292                                                      
REMARK 465     ASP A   293                                                      
REMARK 465     STY A   294                                                      
REMARK 465     STY A   295                                                      
REMARK 465     PRO A   296                                                      
REMARK 465     PRO B    16                                                      
REMARK 465     HIS B    17                                                      
REMARK 470                                                                      
REMARK 470 MISSING ATOM                                                         
REMARK 470 THE FOLLOWING RESIDUES HAVE MISSING ATOMS(M=MODEL NUMBER;            
REMARK 470 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;          
REMARK 470 I=INSERTION CODE):                                                   
REMARK 470   M RES CSSEQI  ATOMS                                                
REMARK 470     MET A  68    CG    SD    CE                                      
REMARK 470     MET A 255    CG    SD    CE                                      
REMARK 470     LEU A 283    CA    C     O     CB    CG    CD1   CD2             
REMARK 470     MET B  68    CG    SD    CE                                      
REMARK 470     MET B 255    CG    SD    CE                                      
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: CLOSE CONTACTS                                             
REMARK 500                                                                      
REMARK 500 THE FOLLOWING ATOMS THAT ARE RELATED BY CRYSTALLOGRAPHIC             
REMARK 500 SYMMETRY ARE IN CLOSE CONTACT.  AN ATOM LOCATED WITHIN 0.15          
REMARK 500 ANGSTROMS OF A SYMMETRY RELATED ATOM IS ASSUMED TO BE ON A           
REMARK 500 SPECIAL POSITION AND IS, THEREFORE, LISTED IN REMARK 375             
REMARK 500 INSTEAD OF REMARK 500.  ATOMS WITH NON-BLANK ALTERNATE               
REMARK 500 LOCATION INDICATORS ARE NOT INCLUDED IN THE CALCULATIONS.            
REMARK 500                                                                      
REMARK 500 DISTANCE CUTOFF:                                                     
REMARK 500 2.2 ANGSTROMS FOR CONTACTS NOT INVOLVING HYDROGEN ATOMS              
REMARK 500 1.6 ANGSTROMS FOR CONTACTS INVOLVING HYDROGEN ATOMS                  
REMARK 500                                                                      
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI  SSYMOP   DISTANCE          
REMARK 500   O    HOH Z    27     O    HOH Z    27    12565     1.97            
REMARK 500   O    HOH Y    57     O    HOH Y    57    11556     2.05            
REMARK 500   CG2  THR A    39     CG2  THR A    39     9555     2.12            
REMARK 500   OE1  GLU B   188     OE1  GLU B   188    11556     2.18            
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND LENGTHS                                      
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,2(A3,1X,A1,I4,A1,1X,A4,3X),F6.3)                  
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   RES CSSEQI ATM2   DEVIATION                     
REMARK 500    PRO A  18   CB    PRO A  18   CG     0.052                        
REMARK 500    THR A 282   C     LEU A 283   N     -0.078                        
REMARK 500    PRO B  42   CB    PRO B  42   CG     0.067                        
REMARK 500    PRO B  42   CG    PRO B  42   CD     0.054                        
REMARK 500    LEU B 139   CG    LEU B 139   CD2   -0.052                        
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES                                       
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)              
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3                                     
REMARK 500    GLY A  90   N   -  CA  -  C   ANGL. DEV. = 14.9 DEGREES           
REMARK 500    VAL A 272   N   -  CA  -  C   ANGL. DEV. =-14.7 DEGREES           
REMARK 500    PRO B  42   C   -  N   -  CA  ANGL. DEV. = 25.0 DEGREES           
REMARK 500    PRO B  42   C   -  N   -  CD  ANGL. DEV. =-34.9 DEGREES           
REMARK 500    GLY B  90   N   -  CA  -  C   ANGL. DEV. = 19.2 DEGREES           
REMARK 500    LEU B 139   CA  -  CB  -  CG  ANGL. DEV. = 13.0 DEGREES           
REMARK 500    PRO B 296   C   -  N   -  CA  ANGL. DEV. = 52.5 DEGREES           
REMARK 500    PRO B 296   C   -  N   -  CD  ANGL. DEV. =-53.6 DEGREES           
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: TORSION ANGLES                                             
REMARK 500                                                                      
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:            
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;               
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                             
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)                    
REMARK 500                                                                      
REMARK 500  M RES CSSEQI        PSI       PHI                                   
REMARK 500    ILE A  19      -65.45     50.09                                   
REMARK 500    ALA A  26     -111.98     65.54                                   
REMARK 500    THR A  39      -37.07     69.15                                   
REMARK 500    TYR A  60      -36.71     60.58                                   
REMARK 500    LEU A 152      125.11     62.93                                   
REMARK 500    ALA B  26     -108.04     54.48                                   
REMARK 500    TYR B  60      -52.01     64.92                                   
REMARK 500    GLN B 248     -115.44     58.96                                   
REMARK 500    TYR B 273      -25.38    124.70                                   
REMARK 525                                                                      
REMARK 525 SOLVENT                                                              
REMARK 525 THE FOLLOWING SOLVENT MOLECULES LIE FARTHER THAN EXPECTED            
REMARK 525 FROM THE PROTEIN OR NUCLEIC ACID MOLECULE AND MAY BE                 
REMARK 525 ASSOCIATED WITH A SYMMETRY RELATED MOLECULE (M=MODEL                 
REMARK 525 NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE          
REMARK 525 NUMBER; I=INSERTION CODE):                                           
REMARK 525                                                                      
REMARK 525  M RES CSSEQI                                                        
REMARK 525    HOH Y   6        DISTANCE =  6.86 ANGSTROMS                       
REMARK 525    HOH Y   9        DISTANCE =  5.17 ANGSTROMS                       
REMARK 525    HOH Y  12        DISTANCE =  7.04 ANGSTROMS                       
REMARK 800                                                                      
REMARK 800 SITE                                                                 
REMARK 800 SITE_IDENTIFIER: AC2                                                 
REMARK 800 SITE_DESCRIPTION: SO4 BINDING SITE FOR CHAIN B                       
DBREF  1GWB A   16   296  UNP    P07359   GPBA_HUMAN      16    296             
DBREF  1GWB B   16   296  UNP    P07359   GPBA_HUMAN      16    296             
SEQRES   1 A  281  PRO HIS PRO ILE CYS GLU VAL SER LYS VAL ALA SER HIS          
SEQRES   2 A  281  LEU GLU VAL ASN CYS ASP LYS ARG ASN LEU THR ALA LEU          
SEQRES   3 A  281  PRO PRO ASP LEU PRO LYS ASP THR THR ILE LEU HIS LEU          
SEQRES   4 A  281  SER GLU ASN LEU LEU TYR THR PHE SER LEU ALA THR LEU          
SEQRES   5 A  281  MET PRO TYR THR ARG LEU THR GLN LEU ASN LEU ASP ARG          
SEQRES   6 A  281  CYS GLU LEU THR LYS LEU GLN VAL ASP GLY THR LEU PRO          
SEQRES   7 A  281  VAL LEU GLY THR LEU ASP LEU SER HIS ASN GLN LEU GLN          
SEQRES   8 A  281  SER LEU PRO LEU LEU GLY GLN THR LEU PRO ALA LEU THR          
SEQRES   9 A  281  VAL LEU ASP VAL SER PHE ASN ARG LEU THR SER LEU PRO          
SEQRES  10 A  281  LEU GLY ALA LEU ARG GLY LEU GLY GLU LEU GLN GLU LEU          
SEQRES  11 A  281  TYR LEU LYS GLY ASN GLU LEU LYS THR LEU PRO PRO GLY          
SEQRES  12 A  281  LEU LEU THR PRO THR PRO LYS LEU GLU LYS LEU SER LEU          
SEQRES  13 A  281  ALA ASN ASN ASN LEU THR GLU LEU PRO ALA GLY LEU LEU          
SEQRES  14 A  281  ASN GLY LEU GLU ASN LEU ASP THR LEU LEU LEU GLN GLU          
SEQRES  15 A  281  ASN SER LEU TYR THR ILE PRO LYS GLY PHE PHE GLY SER          
SEQRES  16 A  281  HIS LEU LEU PRO PHE ALA PHE LEU HIS GLY ASN PRO TRP          
SEQRES  17 A  281  LEU CYS ASN CYS GLU ILE LEU TYR PHE ARG ARG TRP LEU          
SEQRES  18 A  281  GLN ASP ASN ALA GLU ASN VAL TYR VAL TRP LYS GLN GLY          
SEQRES  19 A  281  VAL ASP VAL LYS ALA MET THR SER ASN VAL ALA SER VAL          
SEQRES  20 A  281  GLN CYS ASP ASN SER ASP LYS PHE PRO VAL TYR LYS TYR          
SEQRES  21 A  281  PRO GLY LYS GLY CYS PRO THR LEU GLY ASP GLU GLY ASP          
SEQRES  22 A  281  THR ASP LEU STY ASP STY STY PRO                              
SEQRES   1 B  281  PRO HIS PRO ILE CYS GLU VAL SER LYS VAL ALA SER HIS          
SEQRES   2 B  281  LEU GLU VAL ASN CYS ASP LYS ARG ASN LEU THR ALA LEU          
SEQRES   3 B  281  PRO PRO ASP LEU PRO LYS ASP THR THR ILE LEU HIS LEU          
SEQRES   4 B  281  SER GLU ASN LEU LEU TYR THR PHE SER LEU ALA THR LEU          
SEQRES   5 B  281  MET PRO TYR THR ARG LEU THR GLN LEU ASN LEU ASP ARG          
SEQRES   6 B  281  CYS GLU LEU THR LYS LEU GLN VAL ASP GLY THR LEU PRO          
SEQRES   7 B  281  VAL LEU GLY THR LEU ASP LEU SER HIS ASN GLN LEU GLN          
SEQRES   8 B  281  SER LEU PRO LEU LEU GLY GLN THR LEU PRO ALA LEU THR          
SEQRES   9 B  281  VAL LEU ASP VAL SER PHE ASN ARG LEU THR SER LEU PRO          
SEQRES  10 B  281  LEU GLY ALA LEU ARG GLY LEU GLY GLU LEU GLN GLU LEU          
SEQRES  11 B  281  TYR LEU LYS GLY ASN GLU LEU LYS THR LEU PRO PRO GLY          
SEQRES  12 B  281  LEU LEU THR PRO THR PRO LYS LEU GLU LYS LEU SER LEU          
SEQRES  13 B  281  ALA ASN ASN ASN LEU THR GLU LEU PRO ALA GLY LEU LEU          
SEQRES  14 B  281  ASN GLY LEU GLU ASN LEU ASP THR LEU LEU LEU GLN GLU          
SEQRES  15 B  281  ASN SER LEU TYR THR ILE PRO LYS GLY PHE PHE GLY SER          
SEQRES  16 B  281  HIS LEU LEU PRO PHE ALA PHE LEU HIS GLY ASN PRO TRP          
SEQRES  17 B  281  LEU CYS ASN CYS GLU ILE LEU TYR PHE ARG ARG TRP LEU          
SEQRES  18 B  281  GLN ASP ASN ALA GLU ASN VAL TYR VAL TRP LYS GLN GLY          
SEQRES  19 B  281  VAL ASP VAL LYS ALA MET THR SER ASN VAL ALA SER VAL          
SEQRES  20 B  281  GLN CYS ASP ASN SER ASP LYS PHE PRO VAL TYR LYS TYR          
SEQRES  21 B  281  PRO GLY LYS GLY CYS PRO THR LEU GLY ASP GLU GLY ASP          
SEQRES  22 B  281  THR ASP LEU TYS ASP TYS TYS PRO                              
MODRES 1GWB ASN A  175  ASN  GLYCOSYLATION SITE                                 
MODRES 1GWB ASN B  175  ASN  GLYCOSYLATION SITE                                 
MODRES 1GWB ASN A   37  ASN  GLYCOSYLATION SITE                                 
MODRES 1GWB TYS B  292  TYR  SULFONATED TYROSINE                                
MODRES 1GWB TYS B  294  TYR  SULFONATED TYROSINE                                
MODRES 1GWB TYS B  295  TYR  SULFONATED TYROSINE                                
HET    TYS  B 292      16                                                       
HET    TYS  B 294      16                                                       
HET    TYS  B 295      16                                                       
HET    NAG  A 600      14                                                       
HET    NAG  A1175      14                                                       
HET    NDG  A1137      14                                                       
HET     PT  A 400       1                                                       
HET     PT  A 402       1                                                       
HET     PT  A 403       1                                                       
HET     PT  A 404       1                                                       
HET    SO4  B 504       5                                                       
HET    ACY  A 510       4                                                       
HETNAM     TYS SULFONATED TYROSINE                                              
HETNAM     NAG N-ACETYL-D-GLUCOSAMINE                                           
HETNAM     NDG 2-(ACETYLAMINO)-2-DEOXY-A-D-GLUCOPYRANOSE                        
HETNAM      PT PLATINUM (II) ION                                                
HETNAM     SO4 SULFATE ION                                                      
HETNAM     ACY ACETIC ACID                                                      
HETSYN     NAG NAG                                                              
FORMUL   2  TYS    3(C9 H11 N O6 S)                                             
FORMUL   3  NAG    2(C8 H15 N O6)                                               
FORMUL   5  NDG    C8 H15 N O6                                                  
FORMUL   6   PT    4(PT 2+)                                                     
FORMUL  10  SO4    O4 S 2-                                                      
FORMUL  11  ACY    C2 H4 O2                                                     
FORMUL  12  HOH   *143(H2 O)                                                    
HELIX    1   1 ALA A   65  MET A   68  5                                   4    
HELIX    2   2 GLU A  228  ASN A  239  1                                  12    
HELIX    3   3 ASN A  258  SER A  261  5                                   4    
HELIX    4   4 ALA B   65  MET B   68  5                                   4    
HELIX    5   5 ILE B  229  ASN B  239  1                                  11    
HELIX    6   6 ASN B  258  VAL B  262  5                                   5    
HELIX    7   7 ASP B  265  SER B  267  5                                   3    
SHEET    1  AA11 VAL A  22  SER A  23  0                                        
SHEET    2  AA11 GLU A  30  ASN A  32 -1  O  GLU A  30   N  SER A  23           
SHEET    3  AA11 ILE A  51  HIS A  53  1  O  ILE A  51   N  VAL A  31           
SHEET    4  AA11 GLN A  75  ASN A  77  1  O  GLN A  75   N  LEU A  52           
SHEET    5  AA11 THR A  97  ASP A  99  1  O  THR A  97   N  LEU A  76           
SHEET    6  AA11 VAL A 120  ASP A 122  1  O  VAL A 120   N  LEU A  98           
SHEET    7  AA11 GLU A 144  TYR A 146  1  O  GLU A 144   N  LEU A 121           
SHEET    8  AA11 LYS A 168  SER A 170  1  O  LYS A 168   N  LEU A 145           
SHEET    9  AA11 THR A 192  LEU A 194  1  O  THR A 192   N  LEU A 169           
SHEET   10  AA11 PHE A 215  PHE A 217  1  O  PHE A 215   N  LEU A 193           
SHEET   11  AA11 VAL A 243  TYR A 244  1  N  TYR A 244   O  ALA A 216           
SHEET    1  AB 2 THR A  61  SER A  63  0                                        
SHEET    2  AB 2 LYS A  85  GLN A  87  1  O  LYS A  85   N  PHE A  62           
SHEET    1  AC 2 GLN A 263  CYS A 264  0                                        
SHEET    2  AC 2 PHE A 270  PRO A 271 -1  O  PHE A 270   N  CYS A 264           
SHEET    1  BA11 GLU B  21  SER B  23  0                                        
SHEET    2  BA11 GLU B  30  ASN B  32 -1  O  GLU B  30   N  SER B  23           
SHEET    3  BA11 ILE B  51  HIS B  53  1  O  ILE B  51   N  VAL B  31           
SHEET    4  BA11 GLN B  75  ASN B  77  1  O  GLN B  75   N  LEU B  52           
SHEET    5  BA11 THR B  97  ASP B  99  1  O  THR B  97   N  LEU B  76           
SHEET    6  BA11 VAL B 120  ASP B 122  1  O  VAL B 120   N  LEU B  98           
SHEET    7  BA11 GLU B 144  TYR B 146  1  O  GLU B 144   N  LEU B 121           
SHEET    8  BA11 LYS B 168  SER B 170  1  O  LYS B 168   N  LEU B 145           
SHEET    9  BA11 THR B 192  LEU B 194  1  O  THR B 192   N  LEU B 169           
SHEET   10  BA11 PHE B 215  PHE B 217  1  O  PHE B 215   N  LEU B 193           
SHEET   11  BA11 VAL B 243  TYR B 244  1  N  TYR B 244   O  ALA B 216           
SHEET    1  BB 2 THR B  61  SER B  63  0                                        
SHEET    2  BB 2 LYS B  85  GLN B  87  1  O  LYS B  85   N  PHE B  62           
SSBOND   1 CYS A   20    CYS A   33                                             
SSBOND   2 CYS A  225    CYS A  264                                             
SSBOND   3 CYS A  227    CYS A  280                                             
SSBOND   4 CYS B   20    CYS B   33                                             
SSBOND   5 CYS B  225    CYS B  264                                             
SSBOND   6 CYS B  227    CYS B  280                                             
LINK         ND2 ASN A  37                 C1  NDG A1137                        
LINK         ND2 ASN A 175                 C1  NAG A1175                        
LINK        PT    PT A 400                 O   HOH Y  14                        
LINK        PT    PT A 404                 O   HOH Y  27                        
LINK         ND2 ASN B 175                 C1  NAG A 600                        
LINK         C   LEU B 291                 N   TYS B 292                        
LINK         C   TYS B 292                 N   ASP B 293                        
LINK         C   ASP B 293                 N   TYS B 294                        
LINK         C   TYS B 294                 N   TYS B 295                        
LINK         C   TYS B 295                 N   PRO B 296                        
CISPEP   1 LEU B   41    PRO B   42          0         1.11                     
SITE     1 AC2  2 MET A  68  THR A  91                                          
SITE     1 AC5  4 ASN A 185  GLY A 186  SER A 210  HIS A 211                    
SITE     1 AC6  3 GLU B 151  ASN B 175  HOH Z  54                               
SITE     1 AC8  3 GLU A 151  ASN A 175  HOH Z  55                               
SITE     1 AC9  5 ARG B 137  GLY B 138  GLU B 141  PRO B 162                    
SITE     2 AC9  5 LYS B 165                                                     
CRYST1  202.000  202.000  128.000  90.00  90.00 120.00 P 64 2 2     24          
ORIGX1      1.000000  0.000000  0.000000        0.00000                         
ORIGX2      0.000000  1.000000  0.000000        0.00000                         
ORIGX3      0.000000  0.000000  1.000000        0.00000                         
SCALE1      0.004950  0.002858  0.000000        0.00000                         
SCALE2      0.000000  0.005716  0.000000        0.00000                         
SCALE3      0.000000  0.000000  0.007812        0.00000