PDB entry 1gcw

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HEADER    OXYGEN STORAGE/TRANSPORT                08-AUG-00   1GCW              
TITLE     CO FORM HEMOGLOBIN FROM MUSTELUS GRISEUS                              
COMPND    MOL_ID: 1;                                                            
COMPND   2 MOLECULE: PROTEIN (HEMOGLOBIN);                                      
COMPND   3 CHAIN: A, C;                                                         
COMPND   4 FRAGMENT: ALPHA CHAIN;                                               
COMPND   5 MOL_ID: 2;                                                           
COMPND   6 MOLECULE: PROTEIN (HEMOGLOBIN);                                      
COMPND   7 CHAIN: B, D;                                                         
COMPND   8 FRAGMENT: BETA CHAIN                                                 
SOURCE    MOL_ID: 1;                                                            
SOURCE   2 ORGANISM_SCIENTIFIC: MUSTELUS GRISEUS;                               
SOURCE   3 ORGANISM_COMMON: HOUNDSHARK;                                         
SOURCE   4 TISSUE: BLOOD;                                                       
SOURCE   5 CELL: RED BLOOD CELL;                                                
SOURCE   6 MOL_ID: 2;                                                           
SOURCE   7 ORGANISM_SCIENTIFIC: MUSTELUS GRISEUS;                               
SOURCE   8 ORGANISM_COMMON: HOUNDSHARK;                                         
SOURCE   9 TISSUE: BLOOD;                                                       
SOURCE  10 CELL: RED BLOOD CELL                                                 
KEYWDS    HEMOGLOBIN, CARBON MONOXIDE BOUND FORM                                
EXPDTA    X-RAY DIFFRACTION                                                     
AUTHOR    Y.NAOI,K.T.CHONG,K.YOSHIMATSU,G.MIYAZAKI,J.R.H.TAME,                  
AUTHOR   2 S.Y.PARK,S.I.ADACHI,H.MORIMOTO                                       
REVDAT   4   01-APR-03 1GCW    1       JRNL                                     
REVDAT   3   28-JAN-03 1GCW    1       REMARK                                   
REVDAT   2   11-APR-01 1GCW    1       JRNL                                     
REVDAT   1   06-SEP-00 1GCW    0                                                
JRNL        AUTH   Y.NAOI,K.T.CHONG,K.YOSHIMATSU,G.MIYAZAKI,J.R.TAME,           
JRNL        AUTH 2 S.Y.PARK,S.ADACHI,H.MORIMOTO                                 
JRNL        TITL   THE FUNCTIONAL SIMILARITY AND STRUCTURAL DIVERSITY           
JRNL        TITL 2 OF HUMAN AND CARTILAGINOUS FISH HEMOGLOBINS.                 
JRNL        REF    J.MOL.BIOL.                   V. 307   259 2001              
JRNL        REFN   ASTM JMOBAK  UK ISSN 0022-2836                               
REMARK   1                                                                      
REMARK   2                                                                      
REMARK   2 RESOLUTION. 2.00 ANGSTROMS.                                          
REMARK   3                                                                      
REMARK   3 REFINEMENT.                                                          
REMARK   3   PROGRAM     : X-PLOR 3.851                                         
REMARK   3   AUTHORS     : BRUNGER                                              
REMARK   3                                                                      
REMARK   3  DATA USED IN REFINEMENT.                                            
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.00                           
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 30.00                          
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.000                          
REMARK   3   DATA CUTOFF HIGH         (ABS(F)) : NULL                           
REMARK   3   DATA CUTOFF LOW          (ABS(F)) : NULL                           
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : NULL                           
REMARK   3   NUMBER OF REFLECTIONS             : 37342                          
REMARK   3                                                                      
REMARK   3  FIT TO DATA USED IN REFINEMENT.                                     
REMARK   3   CROSS-VALIDATION METHOD          : NULL                            
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM                          
REMARK   3   R VALUE            (WORKING SET) : 0.199                           
REMARK   3   FREE R VALUE                     : 0.255                           
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : NULL                            
REMARK   3   FREE R VALUE TEST SET COUNT      : 1963                            
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : NULL                            
REMARK   3                                                                      
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.                                  
REMARK   3   TOTAL NUMBER OF BINS USED           : NULL                         
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : NULL                         
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : NULL                         
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : NULL                         
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : NULL                         
REMARK   3   BIN R VALUE           (WORKING SET) : NULL                         
REMARK   3   BIN FREE R VALUE                    : NULL                         
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : NULL                         
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : NULL                         
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : NULL                         
REMARK   3                                                                      
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.                    
REMARK   3   PROTEIN ATOMS            : 4268                                    
REMARK   3   NUCLEIC ACID ATOMS       : 0                                       
REMARK   3   HETEROGEN ATOMS          : 172                                     
REMARK   3   SOLVENT ATOMS            : 63                                      
REMARK   3                                                                      
REMARK   3  B VALUES.                                                           
REMARK   3   FROM WILSON PLOT           (A**2) : 37229.00                       
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : NULL                           
REMARK   3   OVERALL ANISOTROPIC B VALUE.                                       
REMARK   3    B11 (A**2) : NULL                                                 
REMARK   3    B22 (A**2) : NULL                                                 
REMARK   3    B33 (A**2) : NULL                                                 
REMARK   3    B12 (A**2) : NULL                                                 
REMARK   3    B13 (A**2) : NULL                                                 
REMARK   3    B23 (A**2) : NULL                                                 
REMARK   3                                                                      
REMARK   3  ESTIMATED COORDINATE ERROR.                                         
REMARK   3   ESD FROM LUZZATI PLOT        (A) : NULL                            
REMARK   3   ESD FROM SIGMAA              (A) : NULL                            
REMARK   3   LOW RESOLUTION CUTOFF        (A) : NULL                            
REMARK   3                                                                      
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.                         
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : NULL                            
REMARK   3   ESD FROM C-V SIGMAA          (A) : NULL                            
REMARK   3                                                                      
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.                                   
REMARK   3   BOND LENGTHS                 (A) : 0.008                           
REMARK   3   BOND ANGLES            (DEGREES) : 1.09                            
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : NULL                            
REMARK   3   IMPROPER ANGLES        (DEGREES) : NULL                            
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL MODEL : NULL                                      
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA                
REMARK   3   MAIN-CHAIN BOND              (A**2) : NULL  ; NULL                 
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : NULL  ; NULL                 
REMARK   3   SIDE-CHAIN BOND              (A**2) : NULL  ; NULL                 
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : NULL  ; NULL                 
REMARK   3                                                                      
REMARK   3  NCS MODEL : NULL                                                    
REMARK   3                                                                      
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT          
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL                 
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL                 
REMARK   3                                                                      
REMARK   3  PARAMETER FILE  1  : NULL                                           
REMARK   3  TOPOLOGY FILE  1   : NULL                                           
REMARK   3                                                                      
REMARK   3  OTHER REFINEMENT REMARKS: NULL                                      
REMARK   4                                                                      
REMARK   4 1GCW COMPLIES WITH FORMAT V. 3.0, 1-DEC-2006                         
REMARK   4                                                                      
REMARK   4 THIS IS THE REMEDIATED VERSION OF THIS PDB ENTRY.                    
REMARK   4 REMEDIATED DATA FILE REVISION 3.101 (2007-05-01)                     
REMARK 100                                                                      
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBJ .                              
REMARK 100 THE RCSB ID CODE IS RCSB005039.                                      
REMARK 200                                                                      
REMARK 200 EXPERIMENTAL DETAILS                                                 
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION                  
REMARK 200  DATE OF DATA COLLECTION        : NULL                               
REMARK 200  TEMPERATURE           (KELVIN) : 298.0                              
REMARK 200  PH                             : 7.00                               
REMARK 200  NUMBER OF CRYSTALS USED        : 3                                  
REMARK 200                                                                      
REMARK 200  SYNCHROTRON              (Y/N) : Y                                  
REMARK 200  RADIATION SOURCE               : PHOTON FACTORY                     
REMARK 200  BEAMLINE                       : BL-18B                             
REMARK 200  X-RAY GENERATOR MODEL          : NULL                               
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M                                  
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.0                                
REMARK 200  MONOCHROMATOR                  : NULL                               
REMARK 200  OPTICS                         : NULL                               
REMARK 200                                                                      
REMARK 200  DETECTOR TYPE                  : IMAGE PLATE                        
REMARK 200  DETECTOR MANUFACTURER          : FUJI                               
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : DENZO                              
REMARK 200  DATA SCALING SOFTWARE          : SCALEPACK                          
REMARK 200                                                                      
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 255405                             
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.000                              
REMARK 200  RESOLUTION RANGE LOW       (A) : 100.000                            
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 1.000                              
REMARK 200                                                                      
REMARK 200 OVERALL.                                                             
REMARK 200  COMPLETENESS FOR RANGE     (%) : 91.5                               
REMARK 200  DATA REDUNDANCY                : 6.500                              
REMARK 200  R MERGE                    (I) : 0.07500                            
REMARK 200  R SYM                      (I) : NULL                               
REMARK 200   FOR THE DATA SET  : NULL                               
REMARK 200                                                                      
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.                                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.00                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.07                     
REMARK 200  COMPLETENESS FOR SHELL     (%) : 66.4                               
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL                               
REMARK 200  R MERGE FOR SHELL          (I) : 0.26500                            
REMARK 200  R SYM FOR SHELL            (I) : NULL                               
REMARK 200   FOR SHELL         : NULL                               
REMARK 200                                                                      
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH                              
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: NULL                         
REMARK 200 SOFTWARE USED: X-PLOR                                                
REMARK 200 STARTING MODEL: NULL                                                 
REMARK 200                                                                      
REMARK 200 REMARK: NULL                                                         
REMARK 280                                                                      
REMARK 280 CRYSTAL                                                              
REMARK 280 SOLVENT CONTENT, VS   (%): 50.24                                     
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.47                     
REMARK 280                                                                      
REMARK 280 CRYSTALLIZATION CONDITIONS: PEG2000, SODIUM PHOSPHATE, HYDRO         
REMARK 280  SULFIDE, PH 7.00, SMALL TUBES, TEMPERATURE 298K                     
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY                                            
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 21                       
REMARK 290                                                                      
REMARK 290      SYMOP   SYMMETRY                                                
REMARK 290     NNNMMM   OPERATOR                                                
REMARK 290       1555   X,Y,Z                                                   
REMARK 290       2555   1/2-X,-Y,1/2+Z                                          
REMARK 290       3555   -X,1/2+Y,1/2-Z                                          
REMARK 290       4555   1/2+X,1/2-Y,-Z                                          
REMARK 290                                                                      
REMARK 290     WHERE NNN -> OPERATOR NUMBER                                     
REMARK 290           MMM -> TRANSLATION VECTOR                                  
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS                            
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM             
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY                
REMARK 290 RELATED MOLECULES.                                                   
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000       33.35000            
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       57.41500            
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       40.22500            
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000       57.41500            
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       33.35000            
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       40.22500            
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000            
REMARK 290                                                                      
REMARK 290 REMARK: NULL                                                         
REMARK 300                                                                      
REMARK 300 BIOMOLECULE: 1                                                       
REMARK 300 THIS ENTRY CONTAINS THE CRYSTALLOGRAPHIC ASYMMETRIC UNIT             
REMARK 300 WHICH CONSISTS OF 4 CHAIN(S). SEE REMARK 350 FOR                     
REMARK 300 INFORMATION ON GENERATING THE BIOLOGICAL MOLECULE(S).                
REMARK 350                                                                      
REMARK 350 GENERATING THE BIOMOLECULE                                           
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN           
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE                
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS          
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND                          
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.                               
REMARK 350                                                                      
REMARK 350 BIOMOLECULE: 1                                                       
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B, C, D                            
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 465                                                                      
REMARK 465 MISSING RESIDUES                                                     
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE                       
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)                
REMARK 465                                                                      
REMARK 465   M RES C SSSEQI                                                     
REMARK 465     LYS B    44                                                      
REMARK 465     ARG B    45                                                      
REMARK 465     THR B    46                                                      
REMARK 465     ASP B    47                                                      
REMARK 465     GLN B   134                                                      
REMARK 465     TYR B   135                                                      
REMARK 465     LYS D    44                                                      
REMARK 465     ARG D    45                                                      
REMARK 465     THR D    46                                                      
REMARK 465     ASP D    47                                                      
REMARK 465     GLN D   134                                                      
REMARK 465     TYR D   135                                                      
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND LENGTHS                                      
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,2(A3,1X,A1,I4,A1,1X,A4,3X),F6.3)                  
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   RES CSSEQI ATM2   DEVIATION                     
REMARK 500    HIS A  58   CG    HIS A  58   CD2    0.059                        
REMARK 500    MET B  31   SD    MET B  31   CE    -0.097                        
REMARK 500    PHE B  48   N     PHE B  48   CA     0.043                        
REMARK 500    HIS B  53   CG    HIS B  53   CD2    0.042                        
REMARK 500    HIS B  82   CG    HIS B  82   CD2    0.063                        
REMARK 500    HIS B  82   NE2   HIS B  82   CD2    0.048                        
REMARK 500    HIS C  58   CG    HIS C  58   CD2    0.053                        
REMARK 500    HIS C  87   CG    HIS C  87   CD2    0.048                        
REMARK 500    PHE D  48   N     PHE D  48   CA     0.041                        
REMARK 500    HIS D  82   CG    HIS D  82   CD2    0.047                        
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES                                       
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)              
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3                                     
REMARK 500    HIS A  58   CA  -  CB  -  CG  ANGL. DEV. = -9.2 DEGREES           
REMARK 500    HIS A  58   ND1 -  CG  -  CD2 ANGL. DEV. =  7.6 DEGREES           
REMARK 500    HIS A  58   CG  -  ND1 -  CE1 ANGL. DEV. = -8.8 DEGREES           
REMARK 500    HIS A  58   ND1 -  CE1 -  NE2 ANGL. DEV. =  7.1 DEGREES           
REMARK 500    HIS A  58   CG  -  CD2 -  NE2 ANGL. DEV. = -7.5 DEGREES           
REMARK 500    HIS A  87   ND1 -  CG  -  CD2 ANGL. DEV. =  6.9 DEGREES           
REMARK 500    HIS A  87   CG  -  ND1 -  CE1 ANGL. DEV. = -7.0 DEGREES           
REMARK 500    HIS A 112   N   -  CA  -  C   ANGL. DEV. =  8.2 DEGREES           
REMARK 500    HIS B  53   CA  -  CB  -  CG  ANGL. DEV. = -7.5 DEGREES           
REMARK 500    HIS B  82   ND1 -  CE1 -  NE2 ANGL. DEV. =  6.8 DEGREES           
REMARK 500    LEU B 107   CA  -  CB  -  CG  ANGL. DEV. =  6.8 DEGREES           
REMARK 500    HIS C  58   CA  -  CB  -  CG  ANGL. DEV. = -7.4 DEGREES           
REMARK 500    HIS C  87   ND1 -  CG  -  CD2 ANGL. DEV. =  7.2 DEGREES           
REMARK 500    HIS C  87   CG  -  ND1 -  CE1 ANGL. DEV. = -8.1 DEGREES           
REMARK 500    HIS C  87   ND1 -  CE1 -  NE2 ANGL. DEV. =  6.8 DEGREES           
REMARK 500    HIS D  53   CA  -  CB  -  CG  ANGL. DEV. = -7.5 DEGREES           
REMARK 500    HIS D  53   CB  -  CG  -  CD2 ANGL. DEV. = -6.7 DEGREES           
REMARK 500    HIS D  53   CG  -  CD2 -  NE2 ANGL. DEV. = -6.8 DEGREES           
REMARK 900                                                                      
REMARK 900 RELATED ENTRIES                                                      
REMARK 900 RELATED ID: 1GCV   RELATED DB: PDB                                   
REMARK 900 1GCV CONTAINS THE DEOXY FORM OF HEMOGLOBIN.                          
DBREF  1GCW A    1   140  UNP    Q9YGW2   HBA_MUSGR        1    140             
DBREF  1GCW B    1   136  UNP    Q9YGW1   HBB_MUSGR        1    136             
DBREF  1GCW C    1   140  UNP    Q9YGW2   HBA_MUSGR        1    140             
DBREF  1GCW D    1   136  UNP    Q9YGW1   HBB_MUSGR        1    136             
SEQRES   1 A  140  ALA PHE THR ALA CYS GLU LYS GLN THR ILE GLY LYS ILE          
SEQRES   2 A  140  ALA GLN VAL LEU ALA LYS SER PRO GLU ALA TYR GLY ALA          
SEQRES   3 A  140  GLU CYS LEU ALA ARG LEU PHE VAL THR HIS PRO GLY SER          
SEQRES   4 A  140  LYS SER TYR PHE GLU TYR LYS ASP TYR SER ALA ALA GLY          
SEQRES   5 A  140  ALA LYS VAL GLN VAL HIS GLY GLY LYS VAL ILE ARG ALA          
SEQRES   6 A  140  VAL VAL LYS ALA ALA GLU HIS VAL ASP ASP LEU HIS SER          
SEQRES   7 A  140  HIS LEU GLU THR LEU ALA LEU THR HIS GLY LYS LYS LEU          
SEQRES   8 A  140  LEU VAL ASP PRO GLN ASN PHE PRO MET LEU SER GLU CYS          
SEQRES   9 A  140  ILE ILE VAL THR LEU ALA THR HIS LEU THR GLU PHE SER          
SEQRES  10 A  140  PRO ASP THR HIS CYS ALA VAL ASP LYS LEU LEU SER ALA          
SEQRES  11 A  140  ILE CYS GLN GLU LEU SER SER ARG TYR ARG                      
SEQRES   1 B  135  VAL HIS TRP THR GLN GLU GLU ARG ASP GLU ILE SER LYS          
SEQRES   2 B  135  THR PHE GLN GLY THR ASP MET LYS THR VAL VAL THR GLN          
SEQRES   3 B  135  ALA LEU ASP ARG MET PHE LYS VAL TYR PRO TRP THR ASN          
SEQRES   4 B  135  ARG TYR PHE GLN LYS ARG THR ASP PHE ARG SER SER ILE          
SEQRES   5 B  135  HIS ALA GLY ILE VAL VAL GLY ALA LEU GLN ASP ALA VAL          
SEQRES   6 B  135  LYS HIS MET ASP ASP VAL LYS THR LEU PHE LYS ASP LEU          
SEQRES   7 B  135  SER LYS LYS HIS ALA ASP ASP LEU HIS VAL ASP PRO GLY          
SEQRES   8 B  135  SER PHE HIS LEU LEU THR ASP CYS ILE ILE VAL GLU LEU          
SEQRES   9 B  135  ALA TYR LEU ARG LYS ASP CYS PHE THR PRO HIS ILE GLN          
SEQRES  10 B  135  GLY ILE TRP ASP LYS PHE PHE GLU VAL VAL ILE ASP ALA          
SEQRES  11 B  135  ILE SER LYS GLN TYR                                          
SEQRES   1 C  140  ALA PHE THR ALA CYS GLU LYS GLN THR ILE GLY LYS ILE          
SEQRES   2 C  140  ALA GLN VAL LEU ALA LYS SER PRO GLU ALA TYR GLY ALA          
SEQRES   3 C  140  GLU CYS LEU ALA ARG LEU PHE VAL THR HIS PRO GLY SER          
SEQRES   4 C  140  LYS SER TYR PHE GLU TYR LYS ASP TYR SER ALA ALA GLY          
SEQRES   5 C  140  ALA LYS VAL GLN VAL HIS GLY GLY LYS VAL ILE ARG ALA          
SEQRES   6 C  140  VAL VAL LYS ALA ALA GLU HIS VAL ASP ASP LEU HIS SER          
SEQRES   7 C  140  HIS LEU GLU THR LEU ALA LEU THR HIS GLY LYS LYS LEU          
SEQRES   8 C  140  LEU VAL ASP PRO GLN ASN PHE PRO MET LEU SER GLU CYS          
SEQRES   9 C  140  ILE ILE VAL THR LEU ALA THR HIS LEU THR GLU PHE SER          
SEQRES  10 C  140  PRO ASP THR HIS CYS ALA VAL ASP LYS LEU LEU SER ALA          
SEQRES  11 C  140  ILE CYS GLN GLU LEU SER SER ARG TYR ARG                      
SEQRES   1 D  135  VAL HIS TRP THR GLN GLU GLU ARG ASP GLU ILE SER LYS          
SEQRES   2 D  135  THR PHE GLN GLY THR ASP MET LYS THR VAL VAL THR GLN          
SEQRES   3 D  135  ALA LEU ASP ARG MET PHE LYS VAL TYR PRO TRP THR ASN          
SEQRES   4 D  135  ARG TYR PHE GLN LYS ARG THR ASP PHE ARG SER SER ILE          
SEQRES   5 D  135  HIS ALA GLY ILE VAL VAL GLY ALA LEU GLN ASP ALA VAL          
SEQRES   6 D  135  LYS HIS MET ASP ASP VAL LYS THR LEU PHE LYS ASP LEU          
SEQRES   7 D  135  SER LYS LYS HIS ALA ASP ASP LEU HIS VAL ASP PRO GLY          
SEQRES   8 D  135  SER PHE HIS LEU LEU THR ASP CYS ILE ILE VAL GLU LEU          
SEQRES   9 D  135  ALA TYR LEU ARG LYS ASP CYS PHE THR PRO HIS ILE GLN          
SEQRES  10 D  135  GLY ILE TRP ASP LYS PHE PHE GLU VAL VAL ILE ASP ALA          
SEQRES  11 D  135  ILE SER LYS GLN TYR                                          
HET    HEM  A 141      43                                                       
HET    CMO  A 142       2                                                       
HET    HEM  B 134      43                                                       
HET    CMO  B 135       2                                                       
HET    HEM  C 141      43                                                       
HET    CMO  C 142       2                                                       
HET    HEM  D 134      43                                                       
HET    CMO  D 135       2                                                       
HETNAM     HEM PROTOPORPHYRIN IX CONTAINING FE                                  
HETNAM     CMO CARBON MONOXIDE                                                  
HETSYN     HEM HEME                                                             
FORMUL   5  HEM    4(C34 H32 FE N4 O4)                                          
FORMUL   6  CMO    4(C O)                                                       
FORMUL  13  HOH   *63(H2 O)                                                     
HELIX    1   1 CYS A    5  SER A   20  1                                  16    
HELIX    2   2 SER A   20  HIS A   36  1                                  17    
HELIX    3   3 PRO A   37  PHE A   43  5                                   7    
HELIX    4   4 GLY A   52  GLU A   71  1                                  20    
HELIX    5   5 HIS A   72  ASP A   74  5                                   3    
HELIX    6   6 ASP A   75  LEU A   80  1                                   6    
HELIX    7   7 LEU A   80  LYS A   89  1                                  10    
HELIX    8   8 GLN A   96  LEU A  113  1                                  18    
HELIX    9   9 SER A  117  LEU A  135  1                                  19    
HELIX   10  10 THR B    4  THR B   18  1                                  15    
HELIX   11  11 ASP B   19  TYR B   35  1                                  17    
HELIX   12  12 PRO B   36  GLN B   43  5                                   8    
HELIX   13  13 ARG B   49  HIS B   67  1                                  19    
HELIX   14  14 ASP B   70  PHE B   75  1                                   6    
HELIX   15  15 PHE B   75  ASP B   84  1                                  10    
HELIX   16  16 ASP B   89  GLY B   91  5                                   3    
HELIX   17  17 SER B   92  LYS B  109  1                                  18    
HELIX   18  18 ASP B  110  PHE B  112  5                                   3    
HELIX   19  19 THR B  113  SER B  132  1                                  20    
HELIX   20  20 THR C    3  SER C   20  1                                  18    
HELIX   21  21 SER C   20  HIS C   36  1                                  17    
HELIX   22  22 PRO C   37  PHE C   43  5                                   7    
HELIX   23  23 GLY C   52  GLU C   71  1                                  20    
HELIX   24  24 HIS C   72  ASP C   74  5                                   3    
HELIX   25  25 ASP C   75  LEU C   80  1                                   6    
HELIX   26  26 LEU C   80  GLY C   88  1                                   9    
HELIX   27  27 GLN C   96  LEU C  113  1                                  18    
HELIX   28  28 SER C  117  SER C  136  1                                  20    
HELIX   29  29 THR D    4  THR D   18  1                                  15    
HELIX   30  30 ASP D   19  TYR D   35  1                                  17    
HELIX   31  31 PRO D   36  GLN D   43  5                                   8    
HELIX   32  32 ARG D   49  HIS D   67  1                                  19    
HELIX   33  33 ASP D   70  PHE D   75  1                                   6    
HELIX   34  34 PHE D   75  ASP D   85  1                                  11    
HELIX   35  35 SER D   92  ARG D  108  1                                  17    
HELIX   36  36 LYS D  109  PHE D  112  5                                   4    
HELIX   37  37 THR D  113  ALA D  130  1                                  18    
LINK        FE   HEM A 141                 NE2 HIS A  87                        
LINK        FE   HEM A 141                 C   CMO A 142                        
LINK        FE   HEM B 134                 NE2 HIS B  82                        
LINK        FE   HEM B 134                 C   CMO B 135                        
LINK        FE   HEM C 141                 NE2 HIS C  87                        
LINK        FE   HEM C 141                 C   CMO C 142                        
LINK        FE   HEM D 134                 NE2 HIS D  82                        
LINK        FE   HEM D 134                 C   CMO D 135                        
CRYST1   66.700   80.450  114.830  90.00  90.00  90.00 P 21 21 21    8          
ORIGX1      1.000000  0.000000  0.000000        0.00000                         
ORIGX2      0.000000  1.000000  0.000000        0.00000                         
ORIGX3      0.000000  0.000000  1.000000        0.00000                         
SCALE1      0.014993  0.000000  0.000000        0.00000                         
SCALE2      0.000000  0.012430  0.000000        0.00000                         
SCALE3      0.000000  0.000000  0.008709        0.00000