PDB entry 1fsx

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HEADER    OXYGEN STORAGE/TRANSPORT                11-SEP-00   1FSX              
TITLE     THE X-RAY STRUCTURE DETERMINATION OF BOVINE CARBONMONOXY HB           
TITLE    2 AT 2.1 A RESOLUTION AND ITS RELATIONSHIP TO THE QUATERNARY           
TITLE    3 STRUCTURE OF OTHER HB CRYSTAL FORMS                                  
COMPND    MOL_ID: 1;                                                            
COMPND   2 MOLECULE: HEMOGLOBIN ALPHA CHAIN;                                    
COMPND   3 CHAIN: A, C;                                                         
COMPND   4 MOL_ID: 2;                                                           
COMPND   5 MOLECULE: HEMOGLOBIN BETA CHAIN;                                     
COMPND   6 CHAIN: B, D                                                          
SOURCE    MOL_ID: 1;                                                            
SOURCE   2 ORGANISM_SCIENTIFIC: BOS TAURUS;                                     
SOURCE   3 ORGANISM_COMMON: BOVINE;                                             
SOURCE   4 TISSUE: BLOOD;                                                       
SOURCE   5 MOL_ID: 2;                                                           
SOURCE   6 ORGANISM_SCIENTIFIC: BOS TAURUS;                                     
SOURCE   7 ORGANISM_COMMON: BOVINE;                                             
SOURCE   8 TISSUE: BLOOD                                                        
KEYWDS    HEMOGLOBIN TETRAMER, R-STATE, QUATERNARY STRUCTURE                    
EXPDTA    X-RAY DIFFRACTION                                                     
AUTHOR    M.K.SAFO,D.J.ABRAHAM                                                  
REVDAT   2   01-APR-03 1FSX    1       JRNL                                     
REVDAT   1   06-JUN-01 1FSX    0                                                
JRNL        AUTH   M.K.SAFO,D.J.ABRAHAM                                         
JRNL        TITL   THE X-RAY STRUCTURE DETERMINATION OF BOVINE                  
JRNL        TITL 2 CARBONMONOXY HEMOGLOBIN AT 2.1 A RESOULTION AND              
JRNL        TITL 3 ITS RELATIONSHIP TO THE QUATERNARY STRUCTURES OF             
JRNL        TITL 4 OTHER HEMOGLOBIN CRYSTAL FROMS.                              
JRNL        REF    PROTEIN SCI.                  V.  10  1091 2001              
JRNL        REFN   ASTM PRCIEI  US ISSN 0961-8368                               
REMARK   1                                                                      
REMARK   2                                                                      
REMARK   2 RESOLUTION. 2.10 ANGSTROMS.                                          
REMARK   3                                                                      
REMARK   3 REFINEMENT.                                                          
REMARK   3   PROGRAM     : X-PLOR 3.843                                         
REMARK   3   AUTHORS     : BRUNGER                                              
REMARK   3                                                                      
REMARK   3  DATA USED IN REFINEMENT.                                            
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.10                           
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 64.50                          
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.000                          
REMARK   3   DATA CUTOFF HIGH         (ABS(F)) : NULL                           
REMARK   3   DATA CUTOFF LOW          (ABS(F)) : NULL                           
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : 100.0                          
REMARK   3   NUMBER OF REFLECTIONS             : 32949                          
REMARK   3                                                                      
REMARK   3  FIT TO DATA USED IN REFINEMENT.                                     
REMARK   3   CROSS-VALIDATION METHOD          : NULL                            
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM                          
REMARK   3   R VALUE            (WORKING SET) : 0.205                           
REMARK   3   FREE R VALUE                     : 0.254                           
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : NULL                            
REMARK   3   FREE R VALUE TEST SET COUNT      : 1647                            
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : NULL                            
REMARK   3                                                                      
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.                                  
REMARK   3   TOTAL NUMBER OF BINS USED           : NULL                         
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : NULL                         
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : NULL                         
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : NULL                         
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : NULL                         
REMARK   3   BIN R VALUE           (WORKING SET) : NULL                         
REMARK   3   BIN FREE R VALUE                    : NULL                         
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : NULL                         
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : NULL                         
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : NULL                         
REMARK   3                                                                      
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.                    
REMARK   3   PROTEIN ATOMS            : 4386                                    
REMARK   3   NUCLEIC ACID ATOMS       : 0                                       
REMARK   3   HETEROGEN ATOMS          : 180                                     
REMARK   3   SOLVENT ATOMS            : 136                                     
REMARK   3                                                                      
REMARK   3  B VALUES.                                                           
REMARK   3   FROM WILSON PLOT           (A**2) : 33.40                          
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : NULL                           
REMARK   3   OVERALL ANISOTROPIC B VALUE.                                       
REMARK   3    B11 (A**2) : NULL                                                 
REMARK   3    B22 (A**2) : NULL                                                 
REMARK   3    B33 (A**2) : NULL                                                 
REMARK   3    B12 (A**2) : NULL                                                 
REMARK   3    B13 (A**2) : NULL                                                 
REMARK   3    B23 (A**2) : NULL                                                 
REMARK   3                                                                      
REMARK   3  ESTIMATED COORDINATE ERROR.                                         
REMARK   3   ESD FROM LUZZATI PLOT        (A) : NULL                            
REMARK   3   ESD FROM SIGMAA              (A) : NULL                            
REMARK   3   LOW RESOLUTION CUTOFF        (A) : NULL                            
REMARK   3                                                                      
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.                         
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : NULL                            
REMARK   3   ESD FROM C-V SIGMAA          (A) : NULL                            
REMARK   3                                                                      
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.                                   
REMARK   3   BOND LENGTHS                 (A) : 0.013                           
REMARK   3   BOND ANGLES            (DEGREES) : NULL                            
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : NULL                            
REMARK   3   IMPROPER ANGLES        (DEGREES) : NULL                            
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL MODEL : NULL                                      
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA                
REMARK   3   MAIN-CHAIN BOND              (A**2) : NULL  ; NULL                 
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : NULL  ; NULL                 
REMARK   3   SIDE-CHAIN BOND              (A**2) : NULL  ; NULL                 
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : NULL  ; NULL                 
REMARK   3                                                                      
REMARK   3  NCS MODEL : NULL                                                    
REMARK   3                                                                      
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT          
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL                 
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL                 
REMARK   3                                                                      
REMARK   3  PARAMETER FILE  1  : NULL                                           
REMARK   3  TOPOLOGY FILE  1   : NULL                                           
REMARK   3                                                                      
REMARK   3  OTHER REFINEMENT REMARKS: 1. ANOMALOUS DATA USED FOR                
REMARK   3  REFINEMENT. 2. BULK SOLVENT CORRECTION DURING REFINEMENT            
REMARK   4                                                                      
REMARK   4 1FSX COMPLIES WITH FORMAT V. 3.0, 1-DEC-2006                         
REMARK   4                                                                      
REMARK   4 THIS IS THE REMEDIATED VERSION OF THIS PDB ENTRY.                    
REMARK   4 REMEDIATED DATA FILE REVISION 3.101 (2007-05-29)                     
REMARK 100                                                                      
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB .                              
REMARK 100 THE RCSB ID CODE IS RCSB011873.                                      
REMARK 200                                                                      
REMARK 200 EXPERIMENTAL DETAILS                                                 
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION                  
REMARK 200  DATE OF DATA COLLECTION        : 20-APR-1993                        
REMARK 200  TEMPERATURE           (KELVIN) : 298.0                              
REMARK 200  PH                             : 7.30                               
REMARK 200  NUMBER OF CRYSTALS USED        : 1                                  
REMARK 200                                                                      
REMARK 200  SYNCHROTRON              (Y/N) : N                                  
REMARK 200  RADIATION SOURCE               : ROTATING ANODE                     
REMARK 200  BEAMLINE                       : NULL                               
REMARK 200  X-RAY GENERATOR MODEL          : RIGAKU 200                         
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M                                  
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.5418                             
REMARK 200  MONOCHROMATOR                  : NULL                               
REMARK 200  OPTICS                         : NULL                               
REMARK 200                                                                      
REMARK 200  DETECTOR TYPE                  : IMAGE PLATE                        
REMARK 200  DETECTOR MANUFACTURER          : RIGAKU RAXIS II                    
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : R-AXIS                             
REMARK 200  DATA SCALING SOFTWARE          : CCP4 (AGROVATA, ROTAVATA)          
REMARK 200                                                                      
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 32949                              
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.100                              
REMARK 200  RESOLUTION RANGE LOW       (A) : 64.500                             
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 0.000                              
REMARK 200                                                                      
REMARK 200 OVERALL.                                                             
REMARK 200  COMPLETENESS FOR RANGE     (%) : 96.2                               
REMARK 200  DATA REDUNDANCY                : 3.700                              
REMARK 200  R MERGE                    (I) : 0.10300                            
REMARK 200  R SYM                      (I) : NULL                               
REMARK 200   FOR THE DATA SET  : 5.7000                             
REMARK 200                                                                      
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.                                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.10                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.15                     
REMARK 200  COMPLETENESS FOR SHELL     (%) : 92.7                               
REMARK 200  DATA REDUNDANCY IN SHELL       : 3.60                               
REMARK 200  R MERGE FOR SHELL          (I) : 0.42000                            
REMARK 200  R SYM FOR SHELL            (I) : NULL                               
REMARK 200   FOR SHELL         : NULL                               
REMARK 200                                                                      
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH                              
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: NULL                         
REMARK 200 SOFTWARE USED: AMORE                                                 
REMARK 200 STARTING MODEL: NULL                                                 
REMARK 200                                                                      
REMARK 200 REMARK: NULL                                                         
REMARK 280                                                                      
REMARK 280 CRYSTAL                                                              
REMARK 280 SOLVENT CONTENT, VS   (%): 46.80                                     
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.31                     
REMARK 280                                                                      
REMARK 280 CRYSTALLIZATION CONDITIONS: POTASSIUM PHOSPHATE, SODIUM              
REMARK 280  PHOSPHATE, PYRIDINE, BENZENE, PH 7.3, LIQUID DIFFUSION,             
REMARK 280  TEMPERATURE 298.0K                                                  
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY                                            
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 21                       
REMARK 290                                                                      
REMARK 290      SYMOP   SYMMETRY                                                
REMARK 290     NNNMMM   OPERATOR                                                
REMARK 290       1555   X,Y,Z                                                   
REMARK 290       2555   1/2-X,-Y,1/2+Z                                          
REMARK 290       3555   -X,1/2+Y,1/2-Z                                          
REMARK 290       4555   1/2+X,1/2-Y,-Z                                          
REMARK 290                                                                      
REMARK 290     WHERE NNN -> OPERATOR NUMBER                                     
REMARK 290           MMM -> TRANSLATION VECTOR                                  
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS                            
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM             
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY                
REMARK 290 RELATED MOLECULES.                                                   
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000       39.70500            
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       32.70000            
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       55.31500            
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000       32.70000            
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       39.70500            
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       55.31500            
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000            
REMARK 290                                                                      
REMARK 290 REMARK: NULL                                                         
REMARK 300                                                                      
REMARK 300 BIOMOLECULE: 1                                                       
REMARK 300 THIS ENTRY CONTAINS THE CRYSTALLOGRAPHIC ASYMMETRIC UNIT             
REMARK 300 WHICH CONSISTS OF 4 CHAIN(S). SEE REMARK 350 FOR                     
REMARK 300 INFORMATION ON GENERATING THE BIOLOGICAL MOLECULE(S).                
REMARK 350                                                                      
REMARK 350 GENERATING THE BIOMOLECULE                                           
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN           
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE                
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS          
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND                          
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.                               
REMARK 350                                                                      
REMARK 350 BIOMOLECULE: 1                                                       
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B, C, D                            
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT                     
REMARK 500                                                                      
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.                            
REMARK 500                                                                      
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI                             
REMARK 500   C    CMO     747    FE    HEM     748              1.74            
REMARK 500   C    CMO     142    FE    HEM     143              1.80            
REMARK 500   C    CMO     347    FE    HEM     348              1.80            
REMARK 500   C    CMO     542    FE    HEM     543              1.81            
REMARK 500   O    THR C   537     O    ARG C   541              1.97            
REMARK 500   NE2  HIS A    87    FE    HEM     143              2.04            
REMARK 500   NE2  HIS D   692    FE    HEM     748              2.06            
REMARK 500   NE2  HIS C   487    FE    HEM     543              2.11            
REMARK 500   NE2  HIS B   292    FE    HEM     348              2.14            
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND LENGTHS                                      
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,2(A3,1X,A1,I4,A1,1X,A4,3X),F6.3)                  
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   RES CSSEQI ATM2   DEVIATION                     
REMARK 500    MET B 255   SD    MET B 255   CE    -0.172                        
REMARK 500    CYS D 693   CB    CYS D 693   SG     0.083                        
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES                                       
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)              
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3                                     
REMARK 500    VAL A   1   N   -  CA  -  C   ANGL. DEV. = -8.7 DEGREES           
REMARK 500    ASP A  47   N   -  CA  -  C   ANGL. DEV. =-15.2 DEGREES           
REMARK 500    SER A 138   N   -  CA  -  C   ANGL. DEV. =  9.6 DEGREES           
REMARK 500    LEU B 278   N   -  CA  -  C   ANGL. DEV. =  9.7 DEGREES           
REMARK 500    LEU C 402   CA  -  CB  -  CG  ANGL. DEV. =  8.5 DEGREES           
REMARK 500    ASP C 447   N   -  CA  -  C   ANGL. DEV. =-13.8 DEGREES           
REMARK 500    TYR C 540   N   -  CA  -  C   ANGL. DEV. = 16.0 DEGREES           
REMARK 500    TYR C 540   C   -  N   -  CA  ANGL. DEV. = 12.6 DEGREES           
REMARK 500    PRO D 636   N   -  CA  -  C   ANGL. DEV. =  9.2 DEGREES           
REMARK 500    SER D 649   N   -  CA  -  C   ANGL. DEV. = 10.3 DEGREES           
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: TORSION ANGLES                                             
REMARK 500                                                                      
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:            
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;               
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                             
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)                    
REMARK 500                                                                      
REMARK 500  M RES CSSEQI        PSI       PHI                                   
REMARK 500    LEU D 603      154.26     59.46                                   
DBREF  1FSX A    1   141  UNP    P01966   HBA_BOVIN        1    141             
DBREF  1FSX C  401   541  UNP    P01966   HBA_BOVIN        1    141             
DBREF  1FSX B  202   346  UNP    P02070   HBB_BOVIN        1    145             
DBREF  1FSX D  602   746  UNP    P02070   HBB_BOVIN        1    145             
SEQRES   1 A  141  VAL LEU SER ALA ALA ASP LYS GLY ASN VAL LYS ALA ALA          
SEQRES   2 A  141  TRP GLY LYS VAL GLY GLY HIS ALA ALA GLU TYR GLY ALA          
SEQRES   3 A  141  GLU ALA LEU GLU ARG MET PHE LEU SER PHE PRO THR THR          
SEQRES   4 A  141  LYS THR TYR PHE PRO HIS PHE ASP LEU SER HIS GLY SER          
SEQRES   5 A  141  ALA GLN VAL LYS GLY HIS GLY ALA LYS VAL ALA ALA ALA          
SEQRES   6 A  141  LEU THR LYS ALA VAL GLU HIS LEU ASP ASP LEU PRO GLY          
SEQRES   7 A  141  ALA LEU SER GLU LEU SER ASP LEU HIS ALA HIS LYS LEU          
SEQRES   8 A  141  ARG VAL ASP PRO VAL ASN PHE LYS LEU LEU SER HIS SER          
SEQRES   9 A  141  LEU LEU VAL THR LEU ALA SER HIS LEU PRO SER ASP PHE          
SEQRES  10 A  141  THR PRO ALA VAL HIS ALA SER LEU ASP LYS PHE LEU ALA          
SEQRES  11 A  141  ASN VAL SER THR VAL LEU THR SER LYS TYR ARG                  
SEQRES   1 B  145  MET LEU THR ALA GLU GLU LYS ALA ALA VAL THR ALA PHE          
SEQRES   2 B  145  TRP GLY LYS VAL LYS VAL ASP GLU VAL GLY GLY GLU ALA          
SEQRES   3 B  145  LEU GLY ARG LEU LEU VAL VAL TYR PRO TRP THR GLN ARG          
SEQRES   4 B  145  PHE PHE GLU SER PHE GLY ASP LEU SER THR ALA ASP ALA          
SEQRES   5 B  145  VAL MET ASN ASN PRO LYS VAL LYS ALA HIS GLY LYS LYS          
SEQRES   6 B  145  VAL LEU ASP SER PHE SER ASN GLY MET LYS HIS LEU ASP          
SEQRES   7 B  145  ASP LEU LYS GLY THR PHE ALA ALA LEU SER GLU LEU HIS          
SEQRES   8 B  145  CYS ASP LYS LEU HIS VAL ASP PRO GLU ASN PHE LYS LEU          
SEQRES   9 B  145  LEU GLY ASN VAL LEU VAL VAL VAL LEU ALA ARG ASN PHE          
SEQRES  10 B  145  GLY LYS GLU PHE THR PRO VAL LEU GLN ALA ASP PHE GLN          
SEQRES  11 B  145  LYS VAL VAL ALA GLY VAL ALA ASN ALA LEU ALA HIS ARG          
SEQRES  12 B  145  TYR HIS                                                      
SEQRES   1 C  141  VAL LEU SER ALA ALA ASP LYS GLY ASN VAL LYS ALA ALA          
SEQRES   2 C  141  TRP GLY LYS VAL GLY GLY HIS ALA ALA GLU TYR GLY ALA          
SEQRES   3 C  141  GLU ALA LEU GLU ARG MET PHE LEU SER PHE PRO THR THR          
SEQRES   4 C  141  LYS THR TYR PHE PRO HIS PHE ASP LEU SER HIS GLY SER          
SEQRES   5 C  141  ALA GLN VAL LYS GLY HIS GLY ALA LYS VAL ALA ALA ALA          
SEQRES   6 C  141  LEU THR LYS ALA VAL GLU HIS LEU ASP ASP LEU PRO GLY          
SEQRES   7 C  141  ALA LEU SER GLU LEU SER ASP LEU HIS ALA HIS LYS LEU          
SEQRES   8 C  141  ARG VAL ASP PRO VAL ASN PHE LYS LEU LEU SER HIS SER          
SEQRES   9 C  141  LEU LEU VAL THR LEU ALA SER HIS LEU PRO SER ASP PHE          
SEQRES  10 C  141  THR PRO ALA VAL HIS ALA SER LEU ASP LYS PHE LEU ALA          
SEQRES  11 C  141  ASN VAL SER THR VAL LEU THR SER LYS TYR ARG                  
SEQRES   1 D  145  MET LEU THR ALA GLU GLU LYS ALA ALA VAL THR ALA PHE          
SEQRES   2 D  145  TRP GLY LYS VAL LYS VAL ASP GLU VAL GLY GLY GLU ALA          
SEQRES   3 D  145  LEU GLY ARG LEU LEU VAL VAL TYR PRO TRP THR GLN ARG          
SEQRES   4 D  145  PHE PHE GLU SER PHE GLY ASP LEU SER THR ALA ASP ALA          
SEQRES   5 D  145  VAL MET ASN ASN PRO LYS VAL LYS ALA HIS GLY LYS LYS          
SEQRES   6 D  145  VAL LEU ASP SER PHE SER ASN GLY MET LYS HIS LEU ASP          
SEQRES   7 D  145  ASP LEU LYS GLY THR PHE ALA ALA LEU SER GLU LEU HIS          
SEQRES   8 D  145  CYS ASP LYS LEU HIS VAL ASP PRO GLU ASN PHE LYS LEU          
SEQRES   9 D  145  LEU GLY ASN VAL LEU VAL VAL VAL LEU ALA ARG ASN PHE          
SEQRES  10 D  145  GLY LYS GLU PHE THR PRO VAL LEU GLN ALA ASP PHE GLN          
SEQRES  11 D  145  LYS VAL VAL ALA GLY VAL ALA ASN ALA LEU ALA HIS ARG          
SEQRES  12 D  145  TYR HIS                                                      
HET    CMO    142       2                                                       
HET    HEM    143      43                                                       
HET    CMO    347       2                                                       
HET    HEM    348      43                                                       
HET    CMO    542       2                                                       
HET    HEM    543      43                                                       
HET    CMO    747       2                                                       
HET    HEM    748      43                                                       
HETNAM     CMO CARBON MONOXIDE                                                  
HETNAM     HEM PROTOPORPHYRIN IX CONTAINING FE                                  
HETSYN     HEM HEME                                                             
FORMUL   5  CMO    4(C O)                                                       
FORMUL   6  HEM    4(C34 H32 FE N4 O4)                                          
FORMUL  13  HOH   *136(H2 O)                                                    
HELIX    1   1 SER A    3  GLY A   18  1                                  16    
HELIX    2   2 HIS A   20  PHE A   36  1                                  17    
HELIX    3   3 PRO A   37  PHE A   43  5                                   7    
HELIX    4   4 SER A   52  HIS A   72  1                                  21    
HELIX    5   5 ASP A   75  LEU A   80  1                                   6    
HELIX    6   6 LEU A   80  HIS A   89  1                                  10    
HELIX    7   7 PRO A   95  LEU A  113  1                                  19    
HELIX    8   8 THR A  118  THR A  137  1                                  20    
HELIX    9   9 THR B  204  GLY B  216  1                                  13    
HELIX   10  10 LYS B  219  TYR B  235  1                                  17    
HELIX   11  11 PRO B  236  PHE B  242  5                                   7    
HELIX   12  12 PHE B  242  GLY B  246  5                                   5    
HELIX   13  13 THR B  250  ASN B  256  1                                   7    
HELIX   14  14 ASN B  257  LYS B  276  1                                  20    
HELIX   15  15 ASP B  280  PHE B  285  1                                   6    
HELIX   16  16 PHE B  285  ASP B  294  1                                  10    
HELIX   17  17 PRO B  300  GLY B  319  1                                  20    
HELIX   18  18 LYS B  320  PHE B  322  5                                   3    
HELIX   19  19 THR B  323  HIS B  343  1                                  21    
HELIX   20  20 SER C  403  GLY C  418  1                                  16    
HELIX   21  21 HIS C  420  PHE C  436  1                                  17    
HELIX   22  22 PRO C  437  PHE C  443  5                                   7    
HELIX   23  23 SER C  452  GLU C  471  1                                  20    
HELIX   24  24 ASP C  475  LEU C  480  1                                   6    
HELIX   25  25 LEU C  480  HIS C  489  1                                  10    
HELIX   26  26 PRO C  495  LEU C  513  1                                  19    
HELIX   27  27 THR C  518  THR C  537  1                                  20    
HELIX   28  28 THR D  604  GLY D  616  1                                  13    
HELIX   29  29 LYS D  619  TYR D  635  1                                  17    
HELIX   30  30 PRO D  636  GLU D  643  5                                   8    
HELIX   31  31 PHE D  642  GLY D  646  5                                   5    
HELIX   32  32 THR D  650  ASN D  656  1                                   7    
HELIX   33  33 ASN D  657  MET D  675  1                                  19    
HELIX   34  34 ASP D  680  PHE D  685  1                                   6    
HELIX   35  35 PHE D  685  LYS D  695  1                                  11    
HELIX   36  36 PRO D  700  GLY D  719  1                                  20    
HELIX   37  37 LYS D  720  PHE D  722  5                                   3    
HELIX   38  38 THR D  723  HIS D  743  1                                  21    
HELIX   39  39 ARG D  744  HIS D  746  5                                   3    
CRYST1   79.410  110.630   65.400  90.00  90.00  90.00 P 21 21 21    8          
ORIGX1      1.000000  0.000000  0.000000        0.00000                         
ORIGX2      0.000000  1.000000  0.000000        0.00000                         
ORIGX3      0.000000  0.000000  1.000000        0.00000                         
SCALE1      0.012593  0.000000  0.000000        0.00000                         
SCALE2      0.000000  0.009039  0.000000        0.00000                         
SCALE3      0.000000  0.000000  0.015291        0.00000