PDB entry 1e9i

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HEADER    LYASE                                   16-OCT-00   1E9I              
TITLE     ENOLASE FROM E.COLI                                                   
COMPND    MOL_ID: 1;                                                            
COMPND   2 MOLECULE: ENOLASE;                                                   
COMPND   3 CHAIN: A, B, C, D;                                                   
COMPND   4 SYNONYM: 2-PHOSPHOGLYCERATE DEHYDRATASE, 2-PHOSPHO-D-                
COMPND   5 GLYCERATE HYDRO-LYASE;                                               
COMPND   6 EC: 4.2.1.11;                                                        
COMPND   7 ENGINEERED: YES                                                      
SOURCE    MOL_ID: 1;                                                            
SOURCE   2 ORGANISM_SCIENTIFIC: ESCHERICHIA COLI;                               
SOURCE   3 GENE: ENO;                                                           
SOURCE   4 EXPRESSION_SYSTEM: ESCHERICHIA COLI;                                 
SOURCE   5 EXPRESSION_SYSTEM_STRAIN: BL21(DE3);                                 
SOURCE   6 EXPRESSION_SYSTEM_PLASMID: PET11A                                    
KEYWDS    DEGRADOSOME, LYASE                                                    
EXPDTA    X-RAY DIFFRACTION                                                     
AUTHOR    K.KUHNEL,A.J.CARPOUSIS,B.LUISI                                        
REVDAT   3   01-APR-03 1E9I    1       JRNL                                     
REVDAT   2   26-NOV-01 1E9I    1       JRNL                                     
REVDAT   1   15-MAR-01 1E9I    0                                                
JRNL        AUTH   K.KUHNEL,B.F.LUISI                                           
JRNL        TITL   CRYSTAL STRUCTURE OF THE ESCHERICHIA COLI RNA                
JRNL        TITL 2 DEGRADOSOME COMPONENT ENOLASE.                               
JRNL        REF    J.MOL.BIOL.                   V. 313   583 2001              
JRNL        REFN   ASTM JMOBAK  UK ISSN 0022-2836                               
REMARK   1                                                                      
REMARK   2                                                                      
REMARK   2 RESOLUTION. 2.48 ANGSTROMS.                                          
REMARK   3                                                                      
REMARK   3 REFINEMENT.                                                          
REMARK   3   PROGRAM     : CNS 0.9                                              
REMARK   3   AUTHORS     : BRUNGER,ADAMS,CLORE,DELANO,GROS,GROSSE-              
REMARK   3               : KUNSTLEVE,JIANG,KUSZEWSKI,NILGES, PANNU,             
REMARK   3               : READ,RICE,SIMONSON,WARREN                            
REMARK   3                                                                      
REMARK   3  REFINEMENT TARGET : NULL                                            
REMARK   3                                                                      
REMARK   3  DATA USED IN REFINEMENT.                                            
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.48                           
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 27.97                          
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.000                          
REMARK   3   DATA CUTOFF HIGH         (ABS(F)) : NULL                           
REMARK   3   DATA CUTOFF LOW          (ABS(F)) : NULL                           
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : 93.8                           
REMARK   3   NUMBER OF REFLECTIONS             : 63664                          
REMARK   3                                                                      
REMARK   3  FIT TO DATA USED IN REFINEMENT.                                     
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT                      
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM                          
REMARK   3   R VALUE            (WORKING SET) : 0.229                           
REMARK   3   FREE R VALUE                     : 0.276                           
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 10.100                          
REMARK   3   FREE R VALUE TEST SET COUNT      : 6458                            
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : 0.003                           
REMARK   3                                                                      
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.                                  
REMARK   3   TOTAL NUMBER OF BINS USED           : 10                           
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 2.47                         
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 2.56                         
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 72.70                        
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : 4500                         
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2600                       
REMARK   3   BIN FREE R VALUE                    : 0.3180                       
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : 9.40                         
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : 466                          
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : 0.015                        
REMARK   3                                                                      
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.                    
REMARK   3   PROTEIN ATOMS            : 12340                                   
REMARK   3   NUCLEIC ACID ATOMS       : 0                                       
REMARK   3   HETEROGEN ATOMS          : 5                                       
REMARK   3   SOLVENT ATOMS            : NULL                                    
REMARK   3                                                                      
REMARK   3  B VALUES.                                                           
REMARK   3   FROM WILSON PLOT           (A**2) : 33.00                          
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 49.10                          
REMARK   3   OVERALL ANISOTROPIC B VALUE.                                       
REMARK   3    B11 (A**2) : -7.96000                                             
REMARK   3    B22 (A**2) : 19.94000                                             
REMARK   3    B33 (A**2) : -11.98000                                            
REMARK   3    B12 (A**2) : 0.00000                                              
REMARK   3    B13 (A**2) : 6.63000                                              
REMARK   3    B23 (A**2) : 0.00000                                              
REMARK   3                                                                      
REMARK   3  ESTIMATED COORDINATE ERROR.                                         
REMARK   3   ESD FROM LUZZATI PLOT        (A) : 0.34                            
REMARK   3   ESD FROM SIGMAA              (A) : 0.24                            
REMARK   3   LOW RESOLUTION CUTOFF        (A) : 27.00                           
REMARK   3                                                                      
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.                         
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : 0.42                            
REMARK   3   ESD FROM C-V SIGMAA          (A) : 0.38                            
REMARK   3                                                                      
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.                                   
REMARK   3   BOND LENGTHS                 (A) : 0.007                           
REMARK   3   BOND ANGLES            (DEGREES) : 1.10                            
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : 22.00                           
REMARK   3   IMPROPER ANGLES        (DEGREES) : 0.76                            
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL MODEL : RESTRAINED                                
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA                
REMARK   3   MAIN-CHAIN BOND              (A**2) : 3.250 ; 1.500                
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : 4.260 ; 2.000                
REMARK   3   SIDE-CHAIN BOND              (A**2) : 0.000 ; 2.000                
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : 0.000 ; 2.500                
REMARK   3                                                                      
REMARK   3  BULK SOLVENT MODELING.                                              
REMARK   3   METHOD USED : FLAT MODEL                                           
REMARK   3   KSOL        : 0.33                                                 
REMARK   3   BSOL        : 37.75                                                
REMARK   3                                                                      
REMARK   3  NCS MODEL : NULL                                                    
REMARK   3                                                                      
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT          
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL                 
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL                 
REMARK   3                                                                      
REMARK   3  PARAMETER FILE  1  : PROTEIN_REP.PARAM                              
REMARK   3  PARAMETER FILE  2  : WATER_REP.PARAM                                
REMARK   3  PARAMETER FILE  3  : ION.PARAM                                      
REMARK   3  PARAMETER FILE  4  : NULL                                           
REMARK   3  TOPOLOGY FILE  1   : PROTEIN.TOP                                    
REMARK   3  TOPOLOGY FILE  2   : WATER.TOP                                      
REMARK   3  TOPOLOGY FILE  3   : ION.TOP                                        
REMARK   3  TOPOLOGY FILE  4   : NULL                                           
REMARK   3                                                                      
REMARK   3  OTHER REFINEMENT REMARKS: NULL                                      
REMARK   4                                                                      
REMARK   4 1E9I COMPLIES WITH FORMAT V. 3.0, 1-DEC-2006                         
REMARK   4                                                                      
REMARK   4 THIS IS THE REMEDIATED VERSION OF THIS PDB ENTRY.                    
REMARK   4 REMEDIATED DATA FILE REVISION 3.101 (2007-05-31)                     
REMARK 100                                                                      
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY EBI  .                              
REMARK 100 THE EBI ID CODE IS  EBI-5431  .                                      
REMARK 200                                                                      
REMARK 200 EXPERIMENTAL DETAILS                                                 
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION                  
REMARK 200  DATE OF DATA COLLECTION        : 15-FEB-1999                        
REMARK 200  TEMPERATURE           (KELVIN) : 100.0                              
REMARK 200  PH                             : 7.00                               
REMARK 200  NUMBER OF CRYSTALS USED        : 1                                  
REMARK 200                                                                      
REMARK 200  SYNCHROTRON              (Y/N) : Y                                  
REMARK 200  RADIATION SOURCE               : ELETTRA                            
REMARK 200  BEAMLINE                       : 5.2R                               
REMARK 200  X-RAY GENERATOR MODEL          : NULL                               
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M                                  
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.472                              
REMARK 200  MONOCHROMATOR                  : NULL                               
REMARK 200  OPTICS                         : MIRRORS                            
REMARK 200                                                                      
REMARK 200  DETECTOR TYPE                  : IP                                 
REMARK 200  DETECTOR MANUFACTURER          : MAR345                             
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : DENZO                              
REMARK 200  DATA SCALING SOFTWARE          : SCALEPACK                          
REMARK 200                                                                      
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 63664                              
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.480                              
REMARK 200  RESOLUTION RANGE LOW       (A) : 27.970                             
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 3.000                              
REMARK 200                                                                      
REMARK 200 OVERALL.                                                             
REMARK 200  COMPLETENESS FOR RANGE     (%) : 98.3                               
REMARK 200  DATA REDUNDANCY                : 4.000                              
REMARK 200  R MERGE                    (I) : 0.05300                            
REMARK 200  R SYM                      (I) : NULL                               
REMARK 200   FOR THE DATA SET  : 33.0000                            
REMARK 200                                                                      
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.                                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.47                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.56                     
REMARK 200  COMPLETENESS FOR SHELL     (%) : 90.0                               
REMARK 200  DATA REDUNDANCY IN SHELL       : 4.00                               
REMARK 200  R MERGE FOR SHELL          (I) : 0.10700                            
REMARK 200  R SYM FOR SHELL            (I) : NULL                               
REMARK 200   FOR SHELL         : 10.000                             
REMARK 200                                                                      
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH                              
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT        
REMARK 200 SOFTWARE USED: AMORE                                                 
REMARK 200 STARTING MODEL: PDB ENTRY 1EBG                                       
REMARK 200                                                                      
REMARK 200 REMARK: NULL                                                         
REMARK 280                                                                      
REMARK 280 CRYSTAL                                                              
REMARK 280 SOLVENT CONTENT, VS   (%): 53.10                                     
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.65                     
REMARK 280                                                                      
REMARK 280 CRYSTALLIZATION CONDITIONS: PH 7                                     
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY                                            
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: C 1 2 1                          
REMARK 290                                                                      
REMARK 290      SYMOP   SYMMETRY                                                
REMARK 290     NNNMMM   OPERATOR                                                
REMARK 290       1555   X,Y,Z                                                   
REMARK 290       2555   -X,Y,-Z                                                 
REMARK 290       3555   1/2+X,1/2+Y,Z                                           
REMARK 290       4555   1/2-X,1/2+Y,-Z                                          
REMARK 290                                                                      
REMARK 290     WHERE NNN -> OPERATOR NUMBER                                     
REMARK 290           MMM -> TRANSLATION VECTOR                                  
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS                            
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM             
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY                
REMARK 290 RELATED MOLECULES.                                                   
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   2  0.000000  1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   2  0.000000  0.000000 -1.000000        0.00000            
REMARK 290   SMTRY1   3  1.000000  0.000000  0.000000       53.94150            
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       75.00300            
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   4 -1.000000  0.000000  0.000000       53.94150            
REMARK 290   SMTRY2   4  0.000000  1.000000  0.000000       75.00300            
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000            
REMARK 290                                                                      
REMARK 290 REMARK: NULL                                                         
REMARK 300                                                                      
REMARK 300 BIOMOLECULE: 1, 2                                                    
REMARK 300 THIS ENTRY CONTAINS THE CRYSTALLOGRAPHIC ASYMMETRIC UNIT             
REMARK 300 WHICH CONSISTS OF 4 CHAIN(S). SEE REMARK 350 FOR                     
REMARK 300 INFORMATION ON GENERATING THE BIOLOGICAL MOLECULE(S).                
REMARK 350                                                                      
REMARK 350 GENERATING THE BIOMOLECULE                                           
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN           
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE                
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS          
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND                          
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.                               
REMARK 350                                                                      
REMARK 350 BIOMOLECULE: 1                                                       
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B, W, X, Y, Z                      
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 350 BIOMOLECULE: 2                                                       
REMARK 350 APPLY THE FOLLOWING TO CHAINS: C, D                                  
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 465                                                                      
REMARK 465 MISSING RESIDUES                                                     
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE                       
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)                
REMARK 465                                                                      
REMARK 465   M RES C SSSEQI                                                     
REMARK 465     ALA A   431                                                      
REMARK 465     ALA B   431                                                      
REMARK 465     PHE C   251                                                      
REMARK 465     TYR C   252                                                      
REMARK 465     LYS C   253                                                      
REMARK 465     GLY C   429                                                      
REMARK 465     GLN C   430                                                      
REMARK 465     ALA C   431                                                      
REMARK 470                                                                      
REMARK 470 MISSING ATOM                                                         
REMARK 470 THE FOLLOWING RESIDUES HAVE MISSING ATOMS(M=MODEL NUMBER;            
REMARK 470 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;          
REMARK 470 I=INSERTION CODE):                                                   
REMARK 470   M RES CSSEQI  ATOMS                                                
REMARK 470     LYS A   2    CG    CD    CE    NZ                                
REMARK 470     GLN A  76    CG    CD    OE1   NE2                               
REMARK 470     ILE A  94    CG1   CG2   CD1                                     
REMARK 470     LYS A 197    CG    CD    CE    NZ                                
REMARK 470     GLU A 235    CG    CD    OE1   OE2                               
REMARK 470     GLU A 270    CG    CD    OE1   OE2                               
REMARK 470     GLU A 278    CG    CD    OE1   OE2                               
REMARK 470     LYS A 324    CG    CD    CE    NZ                                
REMARK 470     GLU A 328    CG    CD    OE1   OE2                               
REMARK 470     GLU A 417    CG    CD    OE1   OE2                               
REMARK 470     LYS B  55    CG    CD    CE    NZ                                
REMARK 470     LYS B 142    CG    CD    CE    NZ                                
REMARK 470     ASN B 162    CG    OD1   ND2                                     
REMARK 470     LYS B 176    CG    CD    CE    NZ                                
REMARK 470     GLU B 180    CG    CD    OE1   OE2                               
REMARK 470     LYS B 194    CG    CD    CE    NZ                                
REMARK 470     LYS B 197    CG    CD    CE    NZ                                
REMARK 470     LYS B 199    CG    CD    CE    NZ                                
REMARK 470     GLU B 220    CG    CD    OE1   OE2                               
REMARK 470     LYS B 230    CG    CD    CE    NZ                                
REMARK 470     LYS B 238    CG    CD    CE    NZ                                
REMARK 470     SER B 249    OG                                                  
REMARK 470     LYS B 253    CG    CD    CE    NZ                                
REMARK 470     ASP B 254    CG    OD1   OD2                                     
REMARK 470     LYS B 256    CG    CD    CE    NZ                                
REMARK 470     VAL B 258    CG1   CG2                                           
REMARK 470     ASN B 264    CG    OD1   ND2                                     
REMARK 470     LYS B 265    CG    CD    CE    NZ                                
REMARK 470     PHE B 267    CG    CD1   CD2   CE1   CE2   CZ                    
REMARK 470     THR B 268    OG1   CG2                                           
REMARK 470     GLU B 270    CG    CD    OE1   OE2                               
REMARK 470     GLU B 271    CG    CD    OE1   OE2                               
REMARK 470     THR B 273    OG1   CG2                                           
REMARK 470     GLU B 278    CG    CD    OE1   OE2                               
REMARK 470     LYS B 281    CG    CD    CE    NZ                                
REMARK 470     TYR B 302    CG    CD1   CD2   CE1   CE2   CZ    OH              
REMARK 470     LYS B 305    CG    CD    CE    NZ                                
REMARK 470     LYS B 310    CG    CD    CE    NZ                                
REMARK 470     LYS B 327    CG    CD    CE    NZ                                
REMARK 470     GLU B 328    CG    CD    OE1   OE2                               
REMARK 470     LYS B 332    CG    CD    CE    NZ                                
REMARK 470     LYS B 359    CG    CD    CE    NZ                                
REMARK 470     GLU B 417    CG    CD    OE1   OE2                               
REMARK 470     LYS C  55    CG    CD    CE    NZ                                
REMARK 470     ASP C 160    CG    OD1   OD2                                     
REMARK 470     ASN C 161    CG    OD1   ND2                                     
REMARK 470     ASN C 162    CG    OD1   ND2                                     
REMARK 470     ASP C 164    CG    OD1   OD2                                     
REMARK 470     ILE C 165    CG1   CG2   CD1                                     
REMARK 470     GLN C 166    CG    CD    OE1   NE2                               
REMARK 470     LYS C 176    CG    CD    CE    NZ                                
REMARK 470     LYS C 197    CG    CD    CE    NZ                                
REMARK 470     LYS C 199    CG    CD    CE    NZ                                
REMARK 470     ASN C 214    CG    OD1   ND2                                     
REMARK 470     SER C 217    OG                                                  
REMARK 470     LYS C 230    CG    CD    CE    NZ                                
REMARK 470     LEU C 236    CG    CD1   CD2                                     
REMARK 470     LYS C 238    CG    CD    CE    NZ                                
REMARK 470     CYS C 246    SG                                                  
REMARK 470     ASP C 254    CG    OD1   OD2                                     
REMARK 470     LYS C 256    CG    CD    CE    NZ                                
REMARK 470     TYR C 257    CG    CD1   CD2   CE1   CE2   CZ    OH              
REMARK 470     VAL C 258    CG1   CG2                                           
REMARK 470     GLU C 262    CG    CD    OE1   OE2                               
REMARK 470     ASN C 264    CG    OD1   ND2                                     
REMARK 470     LYS C 265    CG    CD    CE    NZ                                
REMARK 470     PHE C 267    CG    CD1   CD2   CE1   CE2   CZ                    
REMARK 470     SER C 269    OG                                                  
REMARK 470     GLU C 270    CG    CD    OE1   OE2                               
REMARK 470     GLU C 271    CG    CD    OE1   OE2                               
REMARK 470     THR C 273    OG1   CG2                                           
REMARK 470     HIS C 274    CG    ND1   CD2   CE1   NE2                         
REMARK 470     GLU C 277    CG    CD    OE1   OE2                               
REMARK 470     GLU C 278    CG    CD    OE1   OE2                               
REMARK 470     LYS C 281    CG    CD    CE    NZ                                
REMARK 470     GLN C 282    CG    CD    OE1   NE2                               
REMARK 470     LEU C 292    CG    CD1   CD2                                     
REMARK 470     GLU C 294    CG    CD    OE1   OE2                               
REMARK 470     TRP C 297    CG    CD1   CD2   NE1   CE2   CE3   CZ2             
REMARK 470     TRP C 297    CZ3   CH2                                           
REMARK 470     PHE C 300    CG    CD1   CD2   CE1   CE2   CZ                    
REMARK 470     TYR C 302    CG    CD1   CD2   CE1   CE2   CZ    OH              
REMARK 470     GLN C 303    CG    CD    OE1   NE2                               
REMARK 470     LYS C 305    CG    CD    CE    NZ                                
REMARK 470     VAL C 306    CG1   CG2                                           
REMARK 470     LEU C 313    CG    CD1   CD2                                     
REMARK 470     GLU C 331    CG    CD    OE1   OE2                               
REMARK 470     ASN C 336    CG    OD1   ND2                                     
REMARK 470     LYS C 359    CG    CD    CE    NZ                                
REMARK 470     LYS C 425    CG    CD    CE    NZ                                
REMARK 470     LYS D   2    CG    CD    CE    NZ                                
REMARK 470     GLU D  27    CG    CD    OE1   OE2                               
REMARK 470     LYS D  55    CG    CD    CE    NZ                                
REMARK 470     LYS D 142    CG    CD    CE    NZ                                
REMARK 470     ASP D 160    CG    OD1   OD2                                     
REMARK 470     LYS D 197    CG    CD    CE    NZ                                
REMARK 470     LYS D 230    CG    CD    CE    NZ                                
REMARK 470     LYS D 253    CG    CD    CE    NZ                                
REMARK 470     LYS D 256    CG    CD    CE    NZ                                
REMARK 470     GLU D 262    CG    CD    OE1   OE2                               
REMARK 470     LYS D 265    CG    CD    CE    NZ                                
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT                     
REMARK 500                                                                      
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.                            
REMARK 500                                                                      
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI                             
REMARK 500   O    HOH X    81     O    HOH Z   142              2.08            
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: CLOSE CONTACTS                                             
REMARK 500                                                                      
REMARK 500 THE FOLLOWING ATOMS THAT ARE RELATED BY CRYSTALLOGRAPHIC             
REMARK 500 SYMMETRY ARE IN CLOSE CONTACT.  AN ATOM LOCATED WITHIN 0.15          
REMARK 500 ANGSTROMS OF A SYMMETRY RELATED ATOM IS ASSUMED TO BE ON A           
REMARK 500 SPECIAL POSITION AND IS, THEREFORE, LISTED IN REMARK 375             
REMARK 500 INSTEAD OF REMARK 500.  ATOMS WITH NON-BLANK ALTERNATE               
REMARK 500 LOCATION INDICATORS ARE NOT INCLUDED IN THE CALCULATIONS.            
REMARK 500                                                                      
REMARK 500 DISTANCE CUTOFF:                                                     
REMARK 500 2.2 ANGSTROMS FOR CONTACTS NOT INVOLVING HYDROGEN ATOMS              
REMARK 500 1.6 ANGSTROMS FOR CONTACTS INVOLVING HYDROGEN ATOMS                  
REMARK 500                                                                      
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI  SSYMOP   DISTANCE          
REMARK 500   O    HOH W    14     O    HOH W    14     2656     1.17            
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND LENGTHS                                      
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,2(A3,1X,A1,I4,A1,1X,A4,3X),F6.3)                  
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   RES CSSEQI ATM2   DEVIATION                     
REMARK 500    MET A 150   SD    MET A 150   CE     0.044                        
REMARK 500    MET A 357   SD    MET A 357   CE    -0.047                        
REMARK 500    MET A 396   SD    MET A 396   CE    -0.045                        
REMARK 500    MET B  33   SD    MET B  33   CE    -0.042                        
REMARK 500    MET C 357   SD    MET C 357   CE    -0.059                        
REMARK 500    MET D 184   SD    MET D 184   CE    -0.059                        
REMARK 500    MET D 244   CG    MET D 244   SD     0.040                        
REMARK 500    MET D 396   CG    MET D 396   SD     0.045                        
REMARK 500    MET D 396   SD    MET D 396   CE    -0.051                        
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES                                       
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)              
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3                                     
REMARK 500    PRO A 140   N   -  CA  -  C   ANGL. DEV. =  5.9 DEGREES           
REMARK 500    GLY A 141   N   -  CA  -  C   ANGL. DEV. =  5.6 DEGREES           
REMARK 500    VAL A 147   N   -  CA  -  C   ANGL. DEV. = -6.5 DEGREES           
REMARK 500    PRO A 148   N   -  CA  -  C   ANGL. DEV. =  5.8 DEGREES           
REMARK 500    MET A 149   N   -  CA  -  C   ANGL. DEV. = -9.4 DEGREES           
REMARK 500    MET A 150   N   -  CA  -  C   ANGL. DEV. =  6.0 DEGREES           
REMARK 500    GLY A 216   N   -  CA  -  C   ANGL. DEV. =  5.4 DEGREES           
REMARK 500    ALA A 243   N   -  CA  -  C   ANGL. DEV. = -5.6 DEGREES           
REMARK 500    ALA A 247   N   -  CA  -  C   ANGL. DEV. = -5.4 DEGREES           
REMARK 500    THR A 268   N   -  CA  -  C   ANGL. DEV. = -7.7 DEGREES           
REMARK 500    GLY A 308   N   -  CA  -  C   ANGL. DEV. =  6.2 DEGREES           
REMARK 500    GLY A 346   N   -  CA  -  C   ANGL. DEV. =  6.4 DEGREES           
REMARK 500    GLY A 394   N   -  CA  -  C   ANGL. DEV. =  6.7 DEGREES           
REMARK 500    TYR A 421   N   -  CA  -  C   ANGL. DEV. = -6.5 DEGREES           
REMARK 500    GLY B  72   N   -  CA  -  C   ANGL. DEV. =  5.1 DEGREES           
REMARK 500    MET B 149   N   -  CA  -  C   ANGL. DEV. = -5.2 DEGREES           
REMARK 500    ASN B 151   N   -  CA  -  C   ANGL. DEV. = -5.3 DEGREES           
REMARK 500    GLY B 346   N   -  CA  -  C   ANGL. DEV. =  5.5 DEGREES           
REMARK 500    GLU C  49   N   -  CA  -  C   ANGL. DEV. = -5.2 DEGREES           
REMARK 500    PRO C  73   N   -  CA  -  C   ANGL. DEV. =  5.1 DEGREES           
REMARK 500    GLY C  98   N   -  CA  -  C   ANGL. DEV. =  5.0 DEGREES           
REMARK 500    VAL C 147   N   -  CA  -  C   ANGL. DEV. = -5.0 DEGREES           
REMARK 500    PRO C 148   N   -  CA  -  C   ANGL. DEV. =  5.9 DEGREES           
REMARK 500    MET C 149   N   -  CA  -  C   ANGL. DEV. = -5.7 DEGREES           
REMARK 500    GLY C 346   N   -  CA  -  C   ANGL. DEV. =  5.9 DEGREES           
REMARK 500    SER C 368   N   -  CA  -  C   ANGL. DEV. =  5.0 DEGREES           
REMARK 500    TYR C 421   N   -  CA  -  C   ANGL. DEV. = -5.5 DEGREES           
REMARK 500    VAL D 147   N   -  CA  -  C   ANGL. DEV. = -6.8 DEGREES           
REMARK 500    PRO D 148   N   -  CA  -  C   ANGL. DEV. =  6.6 DEGREES           
REMARK 500    MET D 149   N   -  CA  -  C   ANGL. DEV. = -7.2 DEGREES           
REMARK 500    MET D 150   N   -  CA  -  C   ANGL. DEV. =  5.0 DEGREES           
REMARK 500    GLN D 166   N   -  CA  -  C   ANGL. DEV. =  5.8 DEGREES           
REMARK 500    GLU D 167   N   -  CA  -  C   ANGL. DEV. =  5.1 DEGREES           
REMARK 500    PHE D 168   N   -  CA  -  C   ANGL. DEV. = -6.1 DEGREES           
REMARK 500    MET D 169   N   -  CA  -  C   ANGL. DEV. =  5.3 DEGREES           
REMARK 500    THR D 241   N   -  CA  -  C   ANGL. DEV. = -5.1 DEGREES           
REMARK 500    THR D 268   N   -  CA  -  C   ANGL. DEV. = -6.7 DEGREES           
REMARK 500    GLY D 308   N   -  CA  -  C   ANGL. DEV. =  6.6 DEGREES           
REMARK 500    ASP D 309   N   -  CA  -  C   ANGL. DEV. =  5.7 DEGREES           
REMARK 500    GLY D 346   N   -  CA  -  C   ANGL. DEV. =  6.0 DEGREES           
REMARK 500    HIS D 369   N   -  CA  -  C   ANGL. DEV. = -5.1 DEGREES           
REMARK 500    GLY D 394   N   -  CA  -  C   ANGL. DEV. =  6.1 DEGREES           
REMARK 500    TYR D 421   N   -  CA  -  C   ANGL. DEV. = -5.6 DEGREES           
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: TORSION ANGLES                                             
REMARK 500                                                                      
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:            
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;               
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                             
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)                    
REMARK 500                                                                      
REMARK 500  M RES CSSEQI        PSI       PHI                                   
REMARK 500    ARG C 398      120.07     76.58                                   
DBREF  1E9I A    1   431  UNP    P0A6P9   ENO_ECOLI        1    431             
DBREF  1E9I B    1   431  UNP    P0A6P9   ENO_ECOLI        1    431             
DBREF  1E9I C    1   431  UNP    P0A6P9   ENO_ECOLI        1    431             
DBREF  1E9I D    1   431  UNP    P0A6P9   ENO_ECOLI        1    431             
SEQRES   1 A  431  SER LYS ILE VAL LYS ILE ILE GLY ARG GLU ILE ILE ASP          
SEQRES   2 A  431  SER ARG GLY ASN PRO THR VAL GLU ALA GLU VAL HIS LEU          
SEQRES   3 A  431  GLU GLY GLY PHE VAL GLY MET ALA ALA ALA PRO SER GLY          
SEQRES   4 A  431  ALA SER THR GLY SER ARG GLU ALA LEU GLU LEU ARG ASP          
SEQRES   5 A  431  GLY ASP LYS SER ARG PHE LEU GLY LYS GLY VAL THR LYS          
SEQRES   6 A  431  ALA VAL ALA ALA VAL ASN GLY PRO ILE ALA GLN ALA LEU          
SEQRES   7 A  431  ILE GLY LYS ASP ALA LYS ASP GLN ALA GLY ILE ASP LYS          
SEQRES   8 A  431  ILE MET ILE ASP LEU ASP GLY THR GLU ASN LYS SER LYS          
SEQRES   9 A  431  PHE GLY ALA ASN ALA ILE LEU ALA VAL SER LEU ALA ASN          
SEQRES  10 A  431  ALA LYS ALA ALA ALA ALA ALA LYS GLY MET PRO LEU TYR          
SEQRES  11 A  431  GLU HIS ILE ALA GLU LEU ASN GLY THR PRO GLY LYS TYR          
SEQRES  12 A  431  SER MET PRO VAL PRO MET MET ASN ILE ILE ASN GLY GLY          
SEQRES  13 A  431  GLU HIS ALA ASP ASN ASN VAL ASP ILE GLN GLU PHE MET          
SEQRES  14 A  431  ILE GLN PRO VAL GLY ALA LYS THR VAL LYS GLU ALA ILE          
SEQRES  15 A  431  ARG MET GLY SER GLU VAL PHE HIS HIS LEU ALA LYS VAL          
SEQRES  16 A  431  LEU LYS ALA LYS GLY MET ASN THR ALA VAL GLY ASP GLU          
SEQRES  17 A  431  GLY GLY TYR ALA PRO ASN LEU GLY SER ASN ALA GLU ALA          
SEQRES  18 A  431  LEU ALA VAL ILE ALA GLU ALA VAL LYS ALA ALA GLY TYR          
SEQRES  19 A  431  GLU LEU GLY LYS ASP ILE THR LEU ALA MET ASP CYS ALA          
SEQRES  20 A  431  ALA SER GLU PHE TYR LYS ASP GLY LYS TYR VAL LEU ALA          
SEQRES  21 A  431  GLY GLU GLY ASN LYS ALA PHE THR SER GLU GLU PHE THR          
SEQRES  22 A  431  HIS PHE LEU GLU GLU LEU THR LYS GLN TYR PRO ILE VAL          
SEQRES  23 A  431  SER ILE GLU ASP GLY LEU ASP GLU SER ASP TRP ASP GLY          
SEQRES  24 A  431  PHE ALA TYR GLN THR LYS VAL LEU GLY ASP LYS ILE GLN          
SEQRES  25 A  431  LEU VAL GLY ASP ASP LEU PHE VAL THR ASN THR LYS ILE          
SEQRES  26 A  431  LEU LYS GLU GLY ILE GLU LYS GLY ILE ALA ASN SER ILE          
SEQRES  27 A  431  LEU ILE LYS PHE ASN GLN ILE GLY SER LEU THR GLU THR          
SEQRES  28 A  431  LEU ALA ALA ILE LYS MET ALA LYS ASP ALA GLY TYR THR          
SEQRES  29 A  431  ALA VAL ILE SER HIS ARG SER GLY GLU THR GLU ASP ALA          
SEQRES  30 A  431  THR ILE ALA ASP LEU ALA VAL GLY THR ALA ALA GLY GLN          
SEQRES  31 A  431  ILE LYS THR GLY SER MET SER ARG SER ASP ARG VAL ALA          
SEQRES  32 A  431  LYS TYR ASN GLN LEU ILE ARG ILE GLU GLU ALA LEU GLY          
SEQRES  33 A  431  GLU LYS ALA PRO TYR ASN GLY ARG LYS GLU ILE LYS GLY          
SEQRES  34 A  431  GLN ALA                                                      
SEQRES   1 B  431  SER LYS ILE VAL LYS ILE ILE GLY ARG GLU ILE ILE ASP          
SEQRES   2 B  431  SER ARG GLY ASN PRO THR VAL GLU ALA GLU VAL HIS LEU          
SEQRES   3 B  431  GLU GLY GLY PHE VAL GLY MET ALA ALA ALA PRO SER GLY          
SEQRES   4 B  431  ALA SER THR GLY SER ARG GLU ALA LEU GLU LEU ARG ASP          
SEQRES   5 B  431  GLY ASP LYS SER ARG PHE LEU GLY LYS GLY VAL THR LYS          
SEQRES   6 B  431  ALA VAL ALA ALA VAL ASN GLY PRO ILE ALA GLN ALA LEU          
SEQRES   7 B  431  ILE GLY LYS ASP ALA LYS ASP GLN ALA GLY ILE ASP LYS          
SEQRES   8 B  431  ILE MET ILE ASP LEU ASP GLY THR GLU ASN LYS SER LYS          
SEQRES   9 B  431  PHE GLY ALA ASN ALA ILE LEU ALA VAL SER LEU ALA ASN          
SEQRES  10 B  431  ALA LYS ALA ALA ALA ALA ALA LYS GLY MET PRO LEU TYR          
SEQRES  11 B  431  GLU HIS ILE ALA GLU LEU ASN GLY THR PRO GLY LYS TYR          
SEQRES  12 B  431  SER MET PRO VAL PRO MET MET ASN ILE ILE ASN GLY GLY          
SEQRES  13 B  431  GLU HIS ALA ASP ASN ASN VAL ASP ILE GLN GLU PHE MET          
SEQRES  14 B  431  ILE GLN PRO VAL GLY ALA LYS THR VAL LYS GLU ALA ILE          
SEQRES  15 B  431  ARG MET GLY SER GLU VAL PHE HIS HIS LEU ALA LYS VAL          
SEQRES  16 B  431  LEU LYS ALA LYS GLY MET ASN THR ALA VAL GLY ASP GLU          
SEQRES  17 B  431  GLY GLY TYR ALA PRO ASN LEU GLY SER ASN ALA GLU ALA          
SEQRES  18 B  431  LEU ALA VAL ILE ALA GLU ALA VAL LYS ALA ALA GLY TYR          
SEQRES  19 B  431  GLU LEU GLY LYS ASP ILE THR LEU ALA MET ASP CYS ALA          
SEQRES  20 B  431  ALA SER GLU PHE TYR LYS ASP GLY LYS TYR VAL LEU ALA          
SEQRES  21 B  431  GLY GLU GLY ASN LYS ALA PHE THR SER GLU GLU PHE THR          
SEQRES  22 B  431  HIS PHE LEU GLU GLU LEU THR LYS GLN TYR PRO ILE VAL          
SEQRES  23 B  431  SER ILE GLU ASP GLY LEU ASP GLU SER ASP TRP ASP GLY          
SEQRES  24 B  431  PHE ALA TYR GLN THR LYS VAL LEU GLY ASP LYS ILE GLN          
SEQRES  25 B  431  LEU VAL GLY ASP ASP LEU PHE VAL THR ASN THR LYS ILE          
SEQRES  26 B  431  LEU LYS GLU GLY ILE GLU LYS GLY ILE ALA ASN SER ILE          
SEQRES  27 B  431  LEU ILE LYS PHE ASN GLN ILE GLY SER LEU THR GLU THR          
SEQRES  28 B  431  LEU ALA ALA ILE LYS MET ALA LYS ASP ALA GLY TYR THR          
SEQRES  29 B  431  ALA VAL ILE SER HIS ARG SER GLY GLU THR GLU ASP ALA          
SEQRES  30 B  431  THR ILE ALA ASP LEU ALA VAL GLY THR ALA ALA GLY GLN          
SEQRES  31 B  431  ILE LYS THR GLY SER MET SER ARG SER ASP ARG VAL ALA          
SEQRES  32 B  431  LYS TYR ASN GLN LEU ILE ARG ILE GLU GLU ALA LEU GLY          
SEQRES  33 B  431  GLU LYS ALA PRO TYR ASN GLY ARG LYS GLU ILE LYS GLY          
SEQRES  34 B  431  GLN ALA                                                      
SEQRES   1 C  431  SER LYS ILE VAL LYS ILE ILE GLY ARG GLU ILE ILE ASP          
SEQRES   2 C  431  SER ARG GLY ASN PRO THR VAL GLU ALA GLU VAL HIS LEU          
SEQRES   3 C  431  GLU GLY GLY PHE VAL GLY MET ALA ALA ALA PRO SER GLY          
SEQRES   4 C  431  ALA SER THR GLY SER ARG GLU ALA LEU GLU LEU ARG ASP          
SEQRES   5 C  431  GLY ASP LYS SER ARG PHE LEU GLY LYS GLY VAL THR LYS          
SEQRES   6 C  431  ALA VAL ALA ALA VAL ASN GLY PRO ILE ALA GLN ALA LEU          
SEQRES   7 C  431  ILE GLY LYS ASP ALA LYS ASP GLN ALA GLY ILE ASP LYS          
SEQRES   8 C  431  ILE MET ILE ASP LEU ASP GLY THR GLU ASN LYS SER LYS          
SEQRES   9 C  431  PHE GLY ALA ASN ALA ILE LEU ALA VAL SER LEU ALA ASN          
SEQRES  10 C  431  ALA LYS ALA ALA ALA ALA ALA LYS GLY MET PRO LEU TYR          
SEQRES  11 C  431  GLU HIS ILE ALA GLU LEU ASN GLY THR PRO GLY LYS TYR          
SEQRES  12 C  431  SER MET PRO VAL PRO MET MET ASN ILE ILE ASN GLY GLY          
SEQRES  13 C  431  GLU HIS ALA ASP ASN ASN VAL ASP ILE GLN GLU PHE MET          
SEQRES  14 C  431  ILE GLN PRO VAL GLY ALA LYS THR VAL LYS GLU ALA ILE          
SEQRES  15 C  431  ARG MET GLY SER GLU VAL PHE HIS HIS LEU ALA LYS VAL          
SEQRES  16 C  431  LEU LYS ALA LYS GLY MET ASN THR ALA VAL GLY ASP GLU          
SEQRES  17 C  431  GLY GLY TYR ALA PRO ASN LEU GLY SER ASN ALA GLU ALA          
SEQRES  18 C  431  LEU ALA VAL ILE ALA GLU ALA VAL LYS ALA ALA GLY TYR          
SEQRES  19 C  431  GLU LEU GLY LYS ASP ILE THR LEU ALA MET ASP CYS ALA          
SEQRES  20 C  431  ALA SER GLU PHE TYR LYS ASP GLY LYS TYR VAL LEU ALA          
SEQRES  21 C  431  GLY GLU GLY ASN LYS ALA PHE THR SER GLU GLU PHE THR          
SEQRES  22 C  431  HIS PHE LEU GLU GLU LEU THR LYS GLN TYR PRO ILE VAL          
SEQRES  23 C  431  SER ILE GLU ASP GLY LEU ASP GLU SER ASP TRP ASP GLY          
SEQRES  24 C  431  PHE ALA TYR GLN THR LYS VAL LEU GLY ASP LYS ILE GLN          
SEQRES  25 C  431  LEU VAL GLY ASP ASP LEU PHE VAL THR ASN THR LYS ILE          
SEQRES  26 C  431  LEU LYS GLU GLY ILE GLU LYS GLY ILE ALA ASN SER ILE          
SEQRES  27 C  431  LEU ILE LYS PHE ASN GLN ILE GLY SER LEU THR GLU THR          
SEQRES  28 C  431  LEU ALA ALA ILE LYS MET ALA LYS ASP ALA GLY TYR THR          
SEQRES  29 C  431  ALA VAL ILE SER HIS ARG SER GLY GLU THR GLU ASP ALA          
SEQRES  30 C  431  THR ILE ALA ASP LEU ALA VAL GLY THR ALA ALA GLY GLN          
SEQRES  31 C  431  ILE LYS THR GLY SER MET SER ARG SER ASP ARG VAL ALA          
SEQRES  32 C  431  LYS TYR ASN GLN LEU ILE ARG ILE GLU GLU ALA LEU GLY          
SEQRES  33 C  431  GLU LYS ALA PRO TYR ASN GLY ARG LYS GLU ILE LYS GLY          
SEQRES  34 C  431  GLN ALA                                                      
SEQRES   1 D  431  SER LYS ILE VAL LYS ILE ILE GLY ARG GLU ILE ILE ASP          
SEQRES   2 D  431  SER ARG GLY ASN PRO THR VAL GLU ALA GLU VAL HIS LEU          
SEQRES   3 D  431  GLU GLY GLY PHE VAL GLY MET ALA ALA ALA PRO SER GLY          
SEQRES   4 D  431  ALA SER THR GLY SER ARG GLU ALA LEU GLU LEU ARG ASP          
SEQRES   5 D  431  GLY ASP LYS SER ARG PHE LEU GLY LYS GLY VAL THR LYS          
SEQRES   6 D  431  ALA VAL ALA ALA VAL ASN GLY PRO ILE ALA GLN ALA LEU          
SEQRES   7 D  431  ILE GLY LYS ASP ALA LYS ASP GLN ALA GLY ILE ASP LYS          
SEQRES   8 D  431  ILE MET ILE ASP LEU ASP GLY THR GLU ASN LYS SER LYS          
SEQRES   9 D  431  PHE GLY ALA ASN ALA ILE LEU ALA VAL SER LEU ALA ASN          
SEQRES  10 D  431  ALA LYS ALA ALA ALA ALA ALA LYS GLY MET PRO LEU TYR          
SEQRES  11 D  431  GLU HIS ILE ALA GLU LEU ASN GLY THR PRO GLY LYS TYR          
SEQRES  12 D  431  SER MET PRO VAL PRO MET MET ASN ILE ILE ASN GLY GLY          
SEQRES  13 D  431  GLU HIS ALA ASP ASN ASN VAL ASP ILE GLN GLU PHE MET          
SEQRES  14 D  431  ILE GLN PRO VAL GLY ALA LYS THR VAL LYS GLU ALA ILE          
SEQRES  15 D  431  ARG MET GLY SER GLU VAL PHE HIS HIS LEU ALA LYS VAL          
SEQRES  16 D  431  LEU LYS ALA LYS GLY MET ASN THR ALA VAL GLY ASP GLU          
SEQRES  17 D  431  GLY GLY TYR ALA PRO ASN LEU GLY SER ASN ALA GLU ALA          
SEQRES  18 D  431  LEU ALA VAL ILE ALA GLU ALA VAL LYS ALA ALA GLY TYR          
SEQRES  19 D  431  GLU LEU GLY LYS ASP ILE THR LEU ALA MET ASP CYS ALA          
SEQRES  20 D  431  ALA SER GLU PHE TYR LYS ASP GLY LYS TYR VAL LEU ALA          
SEQRES  21 D  431  GLY GLU GLY ASN LYS ALA PHE THR SER GLU GLU PHE THR          
SEQRES  22 D  431  HIS PHE LEU GLU GLU LEU THR LYS GLN TYR PRO ILE VAL          
SEQRES  23 D  431  SER ILE GLU ASP GLY LEU ASP GLU SER ASP TRP ASP GLY          
SEQRES  24 D  431  PHE ALA TYR GLN THR LYS VAL LEU GLY ASP LYS ILE GLN          
SEQRES  25 D  431  LEU VAL GLY ASP ASP LEU PHE VAL THR ASN THR LYS ILE          
SEQRES  26 D  431  LEU LYS GLU GLY ILE GLU LYS GLY ILE ALA ASN SER ILE          
SEQRES  27 D  431  LEU ILE LYS PHE ASN GLN ILE GLY SER LEU THR GLU THR          
SEQRES  28 D  431  LEU ALA ALA ILE LYS MET ALA LYS ASP ALA GLY TYR THR          
SEQRES  29 D  431  ALA VAL ILE SER HIS ARG SER GLY GLU THR GLU ASP ALA          
SEQRES  30 D  431  THR ILE ALA ASP LEU ALA VAL GLY THR ALA ALA GLY GLN          
SEQRES  31 D  431  ILE LYS THR GLY SER MET SER ARG SER ASP ARG VAL ALA          
SEQRES  32 D  431  LYS TYR ASN GLN LEU ILE ARG ILE GLU GLU ALA LEU GLY          
SEQRES  33 D  431  GLU LYS ALA PRO TYR ASN GLY ARG LYS GLU ILE LYS GLY          
SEQRES  34 D  431  GLN ALA                                                      
HET     MG  A1431       1                                                       
HET     MG  B1431       1                                                       
HET     MG  C1429       1                                                       
HET     MG  D1432       1                                                       
HET     MG  D1433       1                                                       
HET    SO4  D1434       5                                                       
HETNAM      MG MAGNESIUM ION                                                    
HETNAM     SO4 SULFATE ION                                                      
FORMUL   5   MG    5(MG 2+)                                                     
FORMUL  10  SO4    O4 S 2-                                                      
FORMUL  11  HOH   *506(H2 O)                                                    
HELIX    1   1 ARG A   57  LYS A   61  5                                   5    
HELIX    2   2 VAL A   63  GLY A   72  1                                  10    
HELIX    3   3 GLY A   72  ILE A   79  1                                   8    
HELIX    4   4 ASP A   85  GLY A   98  1                                  14    
HELIX    5   5 GLY A  106  LYS A  125  1                                  20    
HELIX    6   6 PRO A  128  GLY A  138  1                                  11    
HELIX    7   7 GLY A  156  ALA A  159  5                                   4    
HELIX    8   8 THR A  177  LYS A  199  1                                  23    
HELIX    9   9 SER A  217  ALA A  232  1                                  16    
HELIX   10  10 ALA A  247  GLU A  250  5                                   4    
HELIX   11  11 THR A  268  TYR A  283  1                                  16    
HELIX   12  12 ASP A  296  GLY A  308  1                                  13    
HELIX   13  13 ASN A  322  LYS A  332  1                                  11    
HELIX   14  14 LYS A  341  ILE A  345  5                                   5    
HELIX   15  15 SER A  347  GLY A  362  1                                  16    
HELIX   16  16 ALA A  377  THR A  386  1                                  10    
HELIX   17  17 ARG A  398  GLY A  416  1                                  19    
HELIX   18  18 GLU A  417  ALA A  419  5                                   3    
HELIX   19  19 ASN A  422  ILE A  427  5                                   6    
HELIX   20  20 ARG B   57  LYS B   61  5                                   5    
HELIX   21  21 VAL B   63  GLY B   72  1                                  10    
HELIX   22  22 GLY B   72  ALA B   77  1                                   6    
HELIX   23  23 ASP B   85  GLY B   98  1                                  14    
HELIX   24  24 GLY B  106  LYS B  125  1                                  20    
HELIX   25  25 PRO B  128  ASN B  137  1                                  10    
HELIX   26  26 GLY B  156  ALA B  159  5                                   4    
HELIX   27  27 THR B  177  LYS B  199  1                                  23    
HELIX   28  28 SER B  217  ALA B  232  1                                  16    
HELIX   29  29 ALA B  247  GLU B  250  5                                   4    
HELIX   30  30 ALA B  260  GLY B  263  5                                   4    
HELIX   31  31 THR B  268  TYR B  283  1                                  16    
HELIX   32  32 ASP B  296  LEU B  307  1                                  12    
HELIX   33  33 ASN B  322  LYS B  332  1                                  11    
HELIX   34  34 LYS B  341  GLY B  346  1                                   6    
HELIX   35  35 SER B  347  ASP B  360  1                                  14    
HELIX   36  36 ALA B  377  THR B  386  1                                  10    
HELIX   37  37 ARG B  398  GLY B  416  1                                  19    
HELIX   38  38 GLU B  417  ALA B  419  5                                   3    
HELIX   39  39 ASN B  422  GLU B  426  5                                   5    
HELIX   40  40 ARG C   57  LYS C   61  5                                   5    
HELIX   41  41 VAL C   63  GLY C   72  1                                  10    
HELIX   42  42 GLY C   72  ILE C   79  1                                   8    
HELIX   43  43 ASP C   85  GLY C   98  1                                  14    
HELIX   44  44 GLY C  106  LYS C  125  1                                  20    
HELIX   45  45 PRO C  128  ASN C  137  1                                  10    
HELIX   46  46 GLY C  156  ALA C  159  5                                   4    
HELIX   47  47 THR C  177  ALA C  198  1                                  22    
HELIX   48  48 SER C  217  GLY C  233  1                                  17    
HELIX   49  49 THR C  268  TYR C  283  1                                  16    
HELIX   50  50 ASP C  296  LEU C  307  1                                  12    
HELIX   51  51 ASN C  322  LYS C  332  1                                  11    
HELIX   52  52 LYS C  341  GLY C  346  1                                   6    
HELIX   53  53 SER C  347  ALA C  361  1                                  15    
HELIX   54  54 ALA C  377  ALA C  387  1                                  11    
HELIX   55  55 ARG C  398  GLY C  416  1                                  19    
HELIX   56  56 GLU C  417  ALA C  419  5                                   3    
HELIX   57  57 ASN C  422  ILE C  427  5                                   6    
HELIX   58  58 ARG D   57  LYS D   61  5                                   5    
HELIX   59  59 VAL D   63  GLY D   72  1                                  10    
HELIX   60  60 GLY D   72  ILE D   79  1                                   8    
HELIX   61  61 ASP D   85  GLY D   98  1                                  14    
HELIX   62  62 GLY D  106  LYS D  125  1                                  20    
HELIX   63  63 PRO D  128  ASN D  137  1                                  10    
HELIX   64  64 GLY D  156  ALA D  159  5                                   4    
HELIX   65  65 THR D  177  ALA D  198  1                                  22    
HELIX   66  66 ALA D  219  GLY D  233  1                                  15    
HELIX   67  67 ALA D  247  GLU D  250  5                                   4    
HELIX   68  68 THR D  268  TYR D  283  1                                  16    
HELIX   69  69 ASP D  296  GLY D  308  1                                  13    
HELIX   70  70 ASN D  322  LYS D  332  1                                  11    
HELIX   71  71 LYS D  341  GLY D  346  1                                   6    
HELIX   72  72 SER D  347  ALA D  361  1                                  15    
HELIX   73  73 ALA D  377  THR D  386  1                                  10    
HELIX   74  74 ARG D  398  GLY D  416  1                                  19    
HELIX   75  75 GLU D  417  ALA D  419  5                                   3    
HELIX   76  76 ASN D  422  ILE D  427  5                                   6    
SHEET    1   A 3 ILE A   3  ILE A   6  0                                        
SHEET    2   A 3 PRO A  18  LEU A  26 -1  O  HIS A  25   N  VAL A   4           
SHEET    3   A 3 ARG A   9  ILE A  12 -1  O  ARG A   9   N  GLU A  21           
SHEET    1  A1 3 ILE A   3  ILE A   6  0                                        
SHEET    2  A1 3 PRO A  18  LEU A  26 -1  O  HIS A  25   N  VAL A   4           
SHEET    3  A1 3 VAL A  31  ALA A  35 -1  O  GLY A  32   N  VAL A  24           
SHEET    1   B 9 VAL A 147  PRO A 148  0                                        
SHEET    2   B 9 GLY A 389  LYS A 392  1  O  GLY A 389   N  VAL A 147           
SHEET    3   B 9 THR A 364  SER A 368  1  O  ALA A 365   N  GLN A 390           
SHEET    4   B 9 SER A 337  ILE A 340  1  O  ILE A 338   N  VAL A 366           
SHEET    5   B 9 GLN A 312  GLY A 315  1  O  GLY A 315   N  LEU A 339           
SHEET    6   B 9 ILE A 285  GLU A 289  1  O  VAL A 286   N  GLN A 312           
SHEET    7   B 9 THR A 241  ASP A 245  1  O  LEU A 242   N  VAL A 286           
SHEET    8   B 9 GLU A 167  GLN A 171 -1  O  GLU A 167   N  ASP A 245           
SHEET    9   B 9 MET A 150  ASN A 154 -1  O  MET A 150   N  ILE A 170           
SHEET    1   C 3 TYR A 252  LYS A 253  0                                        
SHEET    2   C 3 LYS A 256  LEU A 259 -1  O  LYS A 256   N  LYS A 253           
SHEET    3   C 3 LYS A 265  PHE A 267 -1  O  LYS A 265   N  LEU A 259           
SHEET    1   D 3 ILE B   3  ILE B  12  0                                        
SHEET    2   D 3 PRO B  18  LEU B  26 -1  O  THR B  19   N  ILE B  11           
SHEET    3   D 3 PHE B  30  ALA B  35 -1  O  PHE B  30   N  LEU B  26           
SHEET    1   E 9 VAL B 147  PRO B 148  0                                        
SHEET    2   E 9 GLN B 390  LYS B 392  1  N  ILE B 391   O  VAL B 147           
SHEET    3   E 9 THR B 364  SER B 368  1  O  ILE B 367   N  LYS B 392           
SHEET    4   E 9 SER B 337  ILE B 340  1  O  ILE B 338   N  VAL B 366           
SHEET    5   E 9 GLN B 312  GLY B 315  1  O  LEU B 313   N  SER B 337           
SHEET    6   E 9 ILE B 285  GLU B 289  1  O  VAL B 286   N  GLN B 312           
SHEET    7   E 9 THR B 241  ASP B 245  1  O  LEU B 242   N  VAL B 286           
SHEET    8   E 9 GLU B 167  GLN B 171 -1  O  GLU B 167   N  ASP B 245           
SHEET    9   E 9 MET B 150  ASN B 154 -1  O  MET B 150   N  ILE B 170           
SHEET    1   F 3 TYR B 252  LYS B 253  0                                        
SHEET    2   F 3 LYS B 256  LEU B 259 -1  O  LYS B 256   N  LYS B 253           
SHEET    3   F 3 LYS B 265  PHE B 267 -1  O  LYS B 265   N  LEU B 259           
SHEET    1   G 3 ILE C   3  ILE C  12  0                                        
SHEET    2   G 3 PRO C  18  LEU C  26 -1  O  THR C  19   N  ILE C  11           
SHEET    3   G 3 VAL C  31  ALA C  35 -1  O  GLY C  32   N  VAL C  24           
SHEET    1   H 9 VAL C 147  PRO C 148  0                                        
SHEET    2   H 9 GLY C 389  LYS C 392  1  O  GLY C 389   N  VAL C 147           
SHEET    3   H 9 THR C 364  SER C 368  1  O  ALA C 365   N  GLN C 390           
SHEET    4   H 9 SER C 337  ILE C 340  1  O  ILE C 338   N  VAL C 366           
SHEET    5   H 9 GLN C 312  GLY C 315  1  O  GLY C 315   N  LEU C 339           
SHEET    6   H 9 ILE C 285  GLU C 289  1  O  VAL C 286   N  GLN C 312           
SHEET    7   H 9 THR C 241  ASP C 245  1  O  LEU C 242   N  VAL C 286           
SHEET    8   H 9 GLU C 167  GLN C 171 -1  O  GLU C 167   N  ASP C 245           
SHEET    9   H 9 MET C 150  ASN C 154 -1  O  MET C 150   N  ILE C 170           
SHEET    1   I 2 TYR C 257  VAL C 258  0                                        
SHEET    2   I 2 ALA C 266  PHE C 267 -1  O  PHE C 267   N  TYR C 257           
SHEET    1   J 3 ILE D   3  ILE D  12  0                                        
SHEET    2   J 3 PRO D  18  LEU D  26 -1  O  THR D  19   N  ILE D  11           
SHEET    3   J 3 VAL D  31  ALA D  35 -1  O  GLY D  32   N  VAL D  24           
SHEET    1   K 9 VAL D 147  PRO D 148  0                                        
SHEET    2   K 9 GLY D 389  LYS D 392  1  O  GLY D 389   N  VAL D 147           
SHEET    3   K 9 THR D 364  SER D 368  1  O  ALA D 365   N  GLN D 390           
SHEET    4   K 9 SER D 337  ILE D 340  1  O  ILE D 338   N  VAL D 366           
SHEET    5   K 9 GLN D 312  GLY D 315  1  O  GLY D 315   N  LEU D 339           
SHEET    6   K 9 ILE D 285  GLU D 289  1  O  VAL D 286   N  GLN D 312           
SHEET    7   K 9 THR D 241  ASP D 245  1  O  LEU D 242   N  VAL D 286           
SHEET    8   K 9 GLU D 167  GLN D 171 -1  O  GLU D 167   N  ASP D 245           
SHEET    9   K 9 MET D 150  ASN D 154 -1  O  MET D 150   N  ILE D 170           
SHEET    1   L 3 TYR D 252  LYS D 253  0                                        
SHEET    2   L 3 LYS D 256  VAL D 258  0                                        
SHEET    3   L 3 ALA D 266  PHE D 267  0                                        
LINK        MG    MG A1431                 OD2 ASP A 316                        
LINK        MG    MG A1431                 O   HOH Z  56                        
LINK        MG    MG A1431                 OD2 ASP A 245                        
LINK        MG    MG A1431                 OE2 GLU A 289                        
LINK        MG    MG A1431                 O   HOH Z  84                        
LINK        MG    MG A1431                 O   HOH Z  99                        
LINK        MG    MG B1431                 OD2 ASP B 245                        
LINK        MG    MG B1431                 OE2 GLU B 289                        
LINK        MG    MG B1431                 OD2 ASP B 316                        
LINK        MG    MG B1431                 O   HOH Y  86                        
LINK        MG    MG B1431                 O   HOH Y  87                        
LINK        MG    MG C1429                 O   HOH X  99                        
LINK        MG    MG C1429                 OE2 GLU C 289                        
LINK        MG    MG C1429                 O   HOH Z 142                        
LINK        MG    MG C1429                 O   HOH X  81                        
LINK        MG    MG D1432                 OE2 GLU D 289                        
LINK        MG    MG D1432                 OD2 ASP D 316                        
LINK        MG    MG D1432                 O   HOH W  82                        
LINK        MG    MG D1432                 O   HOH Z 147                        
LINK        MG    MG D1432                 OD2 ASP D 245                        
LINK        MG    MG D1433                 O   HOH W 109                        
LINK        MG    MG D1433                 O2  SO4 D1434                        
LINK        MG    MG D1433                 O   HOH W 108                        
LINK        MG    MG D1433                 O   SER D  41                        
LINK        MG    MG D1433                 OG  SER D  41                        
CRYST1  107.883  150.006  127.411  90.00 109.18  90.00 C 1 2 1       4          
ORIGX1      1.000000  0.000000  0.000000        0.00000                         
ORIGX2      0.000000  1.000000  0.000000        0.00000                         
ORIGX3      0.000000  0.000000  1.000000        0.00000                         
SCALE1      0.009269  0.000000  0.003224        0.00000                         
SCALE2      0.000000  0.006666  0.000000        0.00000                         
SCALE3      0.000000  0.000000  0.008310        0.00000                         
MTRIX1   1 -0.959910 -0.067360 -0.272100       51.40480    1                    
MTRIX2   1 -0.068800 -0.884390  0.461660       -5.53899    1                    
MTRIX3   1 -0.271740  0.461870  0.844300        8.97248    1                    
MTRIX1   2  0.993810  0.103560  0.040220      -11.81620    1                    
MTRIX2   2 -0.099570  0.669630  0.735990       42.34035    1                    
MTRIX3   2  0.049290 -0.735440  0.675790       29.33316    1                    
MTRIX1   3 -0.971780 -0.152240 -0.180200       38.67458    1                    
MTRIX2   3 -0.149470 -0.193590  0.969630       40.00534    1                    
MTRIX3   3 -0.182500  0.969200  0.165370       43.63807    1