PDB entry 1doj

[Scop | Full Entry | Seq (local cached copy) | More Options ]


HEADER    HYDROLASE/HYDROLASE INHIBITOR           21-DEC-99   1DOJ              
TITLE     CRYSTAL STRUCTURE OF HUMAN ALPHA-THROMBIN*RWJ-51438 COMPLEX           
TITLE    2 AT 1.7 A                                                             
COMPND    MOL_ID: 1;                                                            
COMPND   2 MOLECULE: ALPHA-THROMBIN;                                            
COMPND   3 CHAIN: A;                                                            
COMPND   4 EC: 3.4.21.5;                                                        
COMPND   5 MOL_ID: 2;                                                           
COMPND   6 MOLECULE: HIRUGEN;                                                   
COMPND   7 CHAIN: B;                                                            
COMPND   8 FRAGMENT: FRAGMENT OF HIRUDIN;                                       
COMPND   9 ENGINEERED: YES;                                                     
COMPND  10 MOL_ID: 3;                                                           
COMPND  11 MOLECULE: PROTEIN (RWJ-51438);                                       
COMPND  12 CHAIN: I;                                                            
COMPND  13 ENGINEERED: YES;                                                     
COMPND  14 OTHER_DETAILS: RWJ-51438 IS JOINED COVALENTLY TO SER195 OF           
COMPND  15 HUMAN ALPHA-THROMBIN                                                 
SOURCE    MOL_ID: 1;                                                            
SOURCE   2 ORGANISM_SCIENTIFIC: HOMO SAPIENS;                                   
SOURCE   3 ORGANISM_COMMON: HUMAN;                                              
SOURCE   4 CELLULAR_LOCATION: PLASMA;                                           
SOURCE   5 MOL_ID: 2;                                                           
SOURCE   6 SYNTHETIC: YES;                                                      
SOURCE   7 OTHER_DETAILS: HIRUGEN, COMES FROM HIRUDIN;                          
SOURCE   8 MOL_ID: 3;                                                           
SOURCE   9 SYNTHETIC: YES;                                                      
SOURCE  10 OTHER_DETAILS: RWJ-51438 WAS CHEMICALLY SYNTHESIZED                  
KEYWDS    THROMBIN, SERINE PROTEASE, ENZYME INHIBITION                          
EXPDTA    X-RAY DIFFRACTION                                                     
AUTHOR    R.RECACHA,M.J.COSTANZO,B.E.MARYANOFF,M.CARSON,L.DELUCAS,              
AUTHOR   2 D.CHATTOPADHYAY                                                      
REVDAT   2   01-APR-03 1DOJ    1       JRNL                                     
REVDAT   1   03-NOV-00 1DOJ    0                                                
JRNL        AUTH   R.RECACHA,M.J.COSTANZO,B.E.MARYANOFF,M.CARSON,               
JRNL        AUTH 2 L.DELUCAS,D.CHATTOPADHYAY                                    
JRNL        TITL   STRUCTURE OF HUMAN ALPHA-THROMBIN COMPLEXED WITH             
JRNL        TITL 2 RWJ-51438 AT 1.7 A: UNUSUAL PERTURBATION OF THE              
JRNL        TITL 3 60A-60I INSERTION LOOP.                                      
JRNL        REF    ACTA CRYSTALLOGR.,SECT.D      V.  56  1395 2000              
JRNL        REFN   ASTM ABCRE6  DK ISSN 0907-4449                               
REMARK   1                                                                      
REMARK   1 REFERENCE 1                                                          
REMARK   1  AUTH   X.QIU,K.P.PADMANABHAN,V.E.CARPEROS,A.TULINSKY,               
REMARK   1  AUTH 2 T.KLINE,J.M.MARAGANORE,J.W.FENTON II                         
REMARK   1  TITL   STRUCTURE OF THE HIRULOG 3-THROMBIN COMPLEX AND              
REMARK   1  TITL 2 NATURE OF THE S' SUBSITES OF SUBSTRATES AND                  
REMARK   1  TITL 3 INHIBITORS                                                   
REMARK   1  REF    BIOCHEMISTRY                  V.  31 11689 1992              
REMARK   1  REFN   ASTM BICHAW  US ISSN 0006-2960                               
REMARK   1 REFERENCE 2                                                          
REMARK   1  AUTH   J.H.MATTHEWS,R.KRISHNAN,M.J.COSTANZO,B.E.MARYANOFF,          
REMARK   1  AUTH 2 A.TULINSKY                                                   
REMARK   1  TITL   CRYSTAL STRUCTURE OF THROMBIN WITH                           
REMARK   1  TITL 2 THIAZOLE-CONTAINING INHIBITORS: PROBES OF THE S1'            
REMARK   1  TITL 3 BINDING SITE                                                 
REMARK   1  REF    BIOPHYS.J.                    V.  71  2830 1996              
REMARK   1  REFN   ASTM BIOJAU  US ISSN 0006-3495                               
REMARK   1 REFERENCE 3                                                          
REMARK   1  AUTH   W.BODE,D.TURK,A.KARSHIKOV                                    
REMARK   1  TITL   THE REFINED 1.9-A X-RAY CRYSTAL STRUCTURE OF                 
REMARK   1  TITL 2 D-PRO-PHE-ARG CHLOROMETHYLKETONE-INHIBITED HUMAN             
REMARK   1  TITL 3 ALPHA-THROMBIN: STRUCTURE ANALYSIS, OVERALL                  
REMARK   1  TITL 4 STRUCTURE, ELECTROSTATIC PROPERTIES, DETAILED                
REMARK   1  TITL 5 ACTIVE-SITE GEOMETRY, AND STRUCTURE-FUNCTION                 
REMARK   1  TITL 6 RELATIONSHIPS                                                
REMARK   1  REF    PROTEIN SCI.                  V.   1   426 1992              
REMARK   1  REFN   ASTM PRCIEI  US ISSN 0961-8368                               
REMARK   2                                                                      
REMARK   2 RESOLUTION. 1.70 ANGSTROMS.                                          
REMARK   3                                                                      
REMARK   3 REFINEMENT.                                                          
REMARK   3   PROGRAM     : X-PLOR 3.851                                         
REMARK   3   AUTHORS     : BRUNGER                                              
REMARK   3                                                                      
REMARK   3  DATA USED IN REFINEMENT.                                            
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.70                           
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 8.00                           
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 3.000                          
REMARK   3   DATA CUTOFF HIGH         (ABS(F)) : 1520225.350                    
REMARK   3   DATA CUTOFF LOW          (ABS(F)) : 0.0000                         
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : 84.2                           
REMARK   3   NUMBER OF REFLECTIONS             : 33357                          
REMARK   3                                                                      
REMARK   3  FIT TO DATA USED IN REFINEMENT.                                     
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT                      
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM                          
REMARK   3   R VALUE            (WORKING SET) : 0.174                           
REMARK   3   FREE R VALUE                     : 0.217                           
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 10.000                          
REMARK   3   FREE R VALUE TEST SET COUNT      : 3322                            
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : 0.004                           
REMARK   3                                                                      
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.                                  
REMARK   3   TOTAL NUMBER OF BINS USED           : 6                            
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 1.70                         
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 1.81                         
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 56.40                        
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : 3311                         
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2760                       
REMARK   3   BIN FREE R VALUE                    : 0.2980                       
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : 10.00                        
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : 3670                         
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : 0.016                        
REMARK   3                                                                      
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.                    
REMARK   3   PROTEIN ATOMS            : 2837                                    
REMARK   3   NUCLEIC ACID ATOMS       : 0                                       
REMARK   3   HETEROGEN ATOMS          : 0                                       
REMARK   3   SOLVENT ATOMS            : 0                                       
REMARK   3                                                                      
REMARK   3  B VALUES.                                                           
REMARK   3   FROM WILSON PLOT           (A**2) : 16.20                          
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 19.50                          
REMARK   3   OVERALL ANISOTROPIC B VALUE.                                       
REMARK   3    B11 (A**2) : 1.02000                                              
REMARK   3    B22 (A**2) : -1.45000                                             
REMARK   3    B33 (A**2) : 0.43000                                              
REMARK   3    B12 (A**2) : 0.00000                                              
REMARK   3    B13 (A**2) : 0.00000                                              
REMARK   3    B23 (A**2) : 0.00000                                              
REMARK   3                                                                      
REMARK   3  ESTIMATED COORDINATE ERROR.                                         
REMARK   3   ESD FROM LUZZATI PLOT        (A) : 0.18                            
REMARK   3   ESD FROM SIGMAA              (A) : 0.13                            
REMARK   3   LOW RESOLUTION CUTOFF        (A) : 5.00                            
REMARK   3                                                                      
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.                         
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : 0.21                            
REMARK   3   ESD FROM C-V SIGMAA          (A) : 0.16                            
REMARK   3                                                                      
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.                                   
REMARK   3   BOND LENGTHS                 (A) : 0.010                           
REMARK   3   BOND ANGLES            (DEGREES) : 1.40                            
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : 26.30                           
REMARK   3   IMPROPER ANGLES        (DEGREES) : 0.61                            
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL MODEL : RESTRAINED                                
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA                
REMARK   3   MAIN-CHAIN BOND              (A**2) : 1.400 ; 1.500                
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : 2.060 ; 2.000                
REMARK   3   SIDE-CHAIN BOND              (A**2) : 2.420 ; 2.000                
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : 3.480 ; 2.500                
REMARK   3                                                                      
REMARK   3  NCS MODEL : NULL                                                    
REMARK   3                                                                      
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT          
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL                 
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL                 
REMARK   3                                                                      
REMARK   3  PARAMETER FILE  1  : TOPPAR:PROTEIN_REP.PARAM                       
REMARK   3  PARAMETER FILE  2  : INH3.PARAM                                     
REMARK   3  PARAMETER FILE  3  : ION.PARAM                                      
REMARK   3  PARAMETER FILE  4  : NAG.PARAM                                      
REMARK   3  PARAMETER FILE  5  : NULL                                           
REMARK   3  PARAMETER FILE  6  : NULL                                           
REMARK   3  TOPOLOGY FILE  1   : TOPPAR:TOPHCSDX.PRO                            
REMARK   3  TOPOLOGY FILE  2   : TOPPAR:TOPH19.SOL                              
REMARK   3  TOPOLOGY FILE  3   : INH2.TOP                                       
REMARK   3  TOPOLOGY FILE  4   : NAG.TOP                                        
REMARK   3  TOPOLOGY FILE  5   : SULF.TOP                                       
REMARK   3  TOPOLOGY FILE  6   : NULL                                           
REMARK   3                                                                      
REMARK   3  OTHER REFINEMENT REMARKS: NULL                                      
REMARK   4                                                                      
REMARK   4 1DOJ COMPLIES WITH FORMAT V. 3.0, 1-DEC-2006                         
REMARK   4                                                                      
REMARK   4 THIS IS THE REMEDIATED VERSION OF THIS PDB ENTRY.                    
REMARK   4 REMEDIATED DATA FILE REVISION 3.100 (2007-03-16)                     
REMARK 100                                                                      
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB .                              
REMARK 100 THE RCSB ID CODE IS RCSB010249.                                      
REMARK 200                                                                      
REMARK 200 EXPERIMENTAL DETAILS                                                 
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION                  
REMARK 200  DATE OF DATA COLLECTION        : 04-MAR-1998                        
REMARK 200  TEMPERATURE           (KELVIN) : 100.0                              
REMARK 200  PH                             : 6.50                               
REMARK 200  NUMBER OF CRYSTALS USED        : 1                                  
REMARK 200                                                                      
REMARK 200  SYNCHROTRON              (Y/N) : N                                  
REMARK 200  RADIATION SOURCE               : ROTATING ANODE                     
REMARK 200  BEAMLINE                       : NULL                               
REMARK 200  X-RAY GENERATOR MODEL          : RIGAKU                             
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M                                  
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.5418                             
REMARK 200  MONOCHROMATOR                  : NULL                               
REMARK 200  OPTICS                         : NULL                               
REMARK 200                                                                      
REMARK 200  DETECTOR TYPE                  : IMAGE PLATE                        
REMARK 200  DETECTOR MANUFACTURER          : RIGAKU                             
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : DENZO                              
REMARK 200  DATA SCALING SOFTWARE          : SCALEPACK                          
REMARK 200                                                                      
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 126549                             
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.660                              
REMARK 200  RESOLUTION RANGE LOW       (A) : 99.000                             
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 0.000                              
REMARK 200                                                                      
REMARK 200 OVERALL.                                                             
REMARK 200  COMPLETENESS FOR RANGE     (%) : 87.1                               
REMARK 200  DATA REDUNDANCY                : 5.000                              
REMARK 200  R MERGE                    (I) : 0.04000                            
REMARK 200  R SYM                      (I) : NULL                               
REMARK 200   FOR THE DATA SET  : 16.3000                            
REMARK 200                                                                      
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.                                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.66                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 99.00                    
REMARK 200  COMPLETENESS FOR SHELL     (%) : 87.1                               
REMARK 200  DATA REDUNDANCY IN SHELL       : 3.00                               
REMARK 200  R MERGE FOR SHELL          (I) : 0.20000                            
REMARK 200  R SYM FOR SHELL            (I) : NULL                               
REMARK 200   FOR SHELL         : NULL                               
REMARK 200                                                                      
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH                              
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: NULL                         
REMARK 200 SOFTWARE USED: AMORE                                                 
REMARK 200 STARTING MODEL: NULL                                                 
REMARK 200                                                                      
REMARK 200 REMARK: NULL                                                         
REMARK 280                                                                      
REMARK 280 CRYSTAL                                                              
REMARK 280 SOLVENT CONTENT, VS   (%): 50.21                                     
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.47                     
REMARK 280                                                                      
REMARK 280 CRYSTALLIZATION CONDITIONS: 0.75 SODIUM ACETATE, 0.01% (W/V),        
REMARK 280  20% POLYETHYLENE GLYCOL 4000 (W/V), PH 6.5, VAPOR DIFFUSION,        
REMARK 280  HANGING DROP, TEMPERATURE 295.0K                                    
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY                                            
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 2                        
REMARK 290                                                                      
REMARK 290      SYMOP   SYMMETRY                                                
REMARK 290     NNNMMM   OPERATOR                                                
REMARK 290       1555   X,Y,Z                                                   
REMARK 290       2555   -X,-Y,Z                                                 
REMARK 290       3555   1/2-X,1/2+Y,-Z                                          
REMARK 290       4555   1/2+X,1/2-Y,-Z                                          
REMARK 290                                                                      
REMARK 290     WHERE NNN -> OPERATOR NUMBER                                     
REMARK 290           MMM -> TRANSLATION VECTOR                                  
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS                            
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM             
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY                
REMARK 290 RELATED MOLECULES.                                                   
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000       31.49100            
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       58.76050            
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000        0.00000            
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       31.49100            
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       58.76050            
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000            
REMARK 290                                                                      
REMARK 290 REMARK: NULL                                                         
REMARK 300                                                                      
REMARK 300 BIOMOLECULE: 1                                                       
REMARK 300 THIS ENTRY CONTAINS THE CRYSTALLOGRAPHIC ASYMMETRIC UNIT             
REMARK 300 WHICH CONSISTS OF 3 CHAIN(S). SEE REMARK 350 FOR                     
REMARK 300 INFORMATION ON GENERATING THE BIOLOGICAL MOLECULE(S).                
REMARK 350                                                                      
REMARK 350 GENERATING THE BIOMOLECULE                                           
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN           
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE                
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS          
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND                          
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.                               
REMARK 350                                                                      
REMARK 350 BIOMOLECULE: 1                                                       
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B, I                               
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 465                                                                      
REMARK 465 MISSING RESIDUES                                                     
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE                       
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)                
REMARK 465                                                                      
REMARK 465   M RES C SSSEQI                                                     
REMARK 465     GLY A    14L                                                     
REMARK 465     ARG A    15                                                      
REMARK 470                                                                      
REMARK 470 MISSING ATOM                                                         
REMARK 470 THE FOLLOWING RESIDUES HAVE MISSING ATOMS(M=MODEL NUMBER;            
REMARK 470 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;          
REMARK 470 I=INSERTION CODE):                                                   
REMARK 470   M RES CSSEQI  ATOMS                                                
REMARK 470     THR A   1H   OG1   CG2                                           
REMARK 470     PHE A   1G   CG    CD1   CD2   CE1   CE2   CZ                    
REMARK 470     ASP A  14M   CB    CG    OD1   OD2                               
REMARK 470     GLU A 247    CG    CD    OE1   OE2                               
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND LENGTHS                                      
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,2(A3,1X,A1,I4,A1,1X,A4,3X),F6.3)                  
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   RES CSSEQI ATM2   DEVIATION                     
REMARK 500    LYS A 169   CB    LYS A 169   CG     0.087                        
REMARK 500    LYS A 169   CB    LYS A 169   CG     0.101                        
REMARK 500    ASP A 243   CB    ASP A 243   CG     0.061                        
REMARK 500    ASP A 243   CB    ASP A 243   CG     0.056                        
REMARK 500    CH3 I 400   C     DPN I 401   N      0.164                        
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES                                       
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)              
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3                                     
REMARK 500    LYS A  14A  N   -  CA  -  C   ANGL. DEV. = 10.0 DEGREES           
REMARK 500    LEU A  33   N   -  CA  -  C   ANGL. DEV. = -8.6 DEGREES           
REMARK 500    GLY A  43   N   -  CA  -  C   ANGL. DEV. = -8.1 DEGREES           
REMARK 500    THR A  60I  N   -  CA  -  C   ANGL. DEV. = -7.8 DEGREES           
REMARK 500    THR A  74   N   -  CA  -  C   ANGL. DEV. =  8.2 DEGREES           
REMARK 500    TYR A  76   N   -  CA  -  C   ANGL. DEV. =-10.3 DEGREES           
REMARK 500    GLU A  97A  N   -  CA  -  C   ANGL. DEV. =  7.9 DEGREES           
REMARK 500    LEU A 123   N   -  CA  -  C   ANGL. DEV. = -9.4 DEGREES           
REMARK 500    ASP A 125   N   -  CA  -  C   ANGL. DEV. = -8.7 DEGREES           
REMARK 500    LEU A 130C  N   -  CA  -  C   ANGL. DEV. =  7.9 DEGREES           
REMARK 500    LYS A 169   CB  -  CA  -  C   ANGL. DEV. = 11.6 DEGREES           
REMARK 500    LYS A 169   CB  -  CA  -  C   ANGL. DEV. =-11.3 DEGREES           
REMARK 500    ALA A 190   N   -  CA  -  C   ANGL. DEV. = -9.8 DEGREES           
REMARK 500    PHE A 199   N   -  CA  -  C   ANGL. DEV. =-13.4 DEGREES           
REMARK 500    ASP A 243   CB  -  CA  -  C   ANGL. DEV. =-10.2 DEGREES           
REMARK 500    ASP A 243   CB  -  CA  -  C   ANGL. DEV. = 10.7 DEGREES           
REMARK 500    DPN I 401   C   -  N   -  CA  ANGL. DEV. =-12.1 DEGREES           
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: TORSION ANGLES                                             
REMARK 500                                                                      
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:            
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;               
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                             
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)                    
REMARK 500                                                                      
REMARK 500  M RES CSSEQI        PSI       PHI                                   
REMARK 500    DPN I 401     -130.86    179.34                                   
REMARK 900                                                                      
REMARK 900 RELATED ENTRIES                                                      
REMARK 900 RELATED ID: 1ABJ   RELATED DB: PDB                                   
REMARK 900 CONTAINS THE SAME PROTEIN COMPLEXED WITH D-PHE-PRO-ARG               
REMARK 900 CHLOROMETHYLKETONE (PPACK) WITH CHLOROMETHYLKETONE REPLACED          
REMARK 900 BY A METHYLENE GROUP                                                 
DBREF  1DOJ A    1H  247  UNP    P00734   THRB_HUMAN     328    622             
SEQRES   1 A  295  THR PHE GLY SER GLY GLU ALA ASP CYS GLY LEU ARG PRO          
SEQRES   2 A  295  LEU PHE GLU LYS LYS SER LEU GLU ASP LYS THR GLU ARG          
SEQRES   3 A  295  GLU LEU LEU GLU SER TYR ILE ASP GLY ARG ILE VAL GLU          
SEQRES   4 A  295  GLY SER ASP ALA GLU ILE GLY MET SER PRO TRP GLN VAL          
SEQRES   5 A  295  MET LEU PHE ARG LYS SER PRO GLN GLU LEU LEU CYS GLY          
SEQRES   6 A  295  ALA SER LEU ILE SER ASP ARG TRP VAL LEU THR ALA ALA          
SEQRES   7 A  295  HIS CYS LEU LEU TYR PRO PRO TRP ASP LYS ASN PHE THR          
SEQRES   8 A  295  GLU ASN ASP LEU LEU VAL ARG ILE GLY LYS HIS SER ARG          
SEQRES   9 A  295  THR ARG TYR GLU ARG ASN ILE GLU LYS ILE SER MET LEU          
SEQRES  10 A  295  GLU LYS ILE TYR ILE HIS PRO ARG TYR ASN TRP ARG GLU          
SEQRES  11 A  295  ASN LEU ASP ARG ASP ILE ALA LEU MET LYS LEU LYS LYS          
SEQRES  12 A  295  PRO VAL ALA PHE SER ASP TYR ILE HIS PRO VAL CYS LEU          
SEQRES  13 A  295  PRO ASP ARG GLU THR ALA ALA SER LEU LEU GLN ALA GLY          
SEQRES  14 A  295  TYR LYS GLY ARG VAL THR GLY TRP GLY ASN LEU LYS GLU          
SEQRES  15 A  295  THR TRP THR ALA ASN VAL GLY LYS GLY GLN PRO SER VAL          
SEQRES  16 A  295  LEU GLN VAL VAL ASN LEU PRO ILE VAL GLU ARG PRO VAL          
SEQRES  17 A  295  CYS LYS ASP SER THR ARG ILE ARG ILE THR ASP ASN MET          
SEQRES  18 A  295  PHE CYS ALA GLY TYR LYS PRO ASP GLU GLY LYS ARG GLY          
SEQRES  19 A  295  ASP ALA CYS GLU GLY ASP SER GLY GLY PRO PHE VAL MET          
SEQRES  20 A  295  LYS SER PRO PHE ASN ASN ARG TRP TYR GLN MET GLY ILE          
SEQRES  21 A  295  VAL SER TRP GLY GLU GLY CYS ASP ARG ASP GLY LYS TYR          
SEQRES  22 A  295  GLY PHE TYR THR HIS VAL PHE ARG LEU LYS LYS TRP ILE          
SEQRES  23 A  295  GLN LYS VAL ILE ASP GLN PHE GLY GLU                          
SEQRES   1 B   11  ASP PHE GLU GLU ILE PRO GLU GLU TYS LEU GLN                  
SEQRES   1 I    4  CH3 DPN PRO AR2                                              
MODRES 1DOJ AR2 I  403  ARG  ARGINYL-BENZOTHIAZOLE-6-CARBOXYLIC ACID            
MODRES 1DOJ TYS B  363  TYR  SULFONATED TYROSINE                                
MODRES 1DOJ CH3 I  400       METHYL GROUP                                       
MODRES 1DOJ DPN I  401  PHE  D-PHENYLALANINE                                    
MODRES 1DOJ ASN A   60G ASN  GLYCOSYLATION SITE                                 
HET    TYS  B 363      16                                                       
HET    CH3  I 400       1                                                       
HET    DPN  I 401      11                                                       
HET    AR2  I 403      23                                                       
HET    NAG  A 250      14                                                       
HET     NA    300       1                                                       
HET     NA    301       1                                                       
HETNAM     TYS SULFONATED TYROSINE                                              
HETNAM     CH3 METHYL GROUP                                                     
HETNAM     DPN D-PHENYLALANINE                                                  
HETNAM     AR2 ARGINYL-BENZOTHIAZOLE-6-CARBOXYLIC ACID                          
HETNAM     NAG N-ACETYL-D-GLUCOSAMINE                                           
HETNAM      NA SODIUM ION                                                       
HETSYN     NAG NAG                                                              
FORMUL   2  TYS    C9 H11 N O6 S                                                
FORMUL   3  CH3    C H4                                                         
FORMUL   3  DPN    C9 H11 N O2                                                  
FORMUL   3  AR2    C14 H17 N5 O3 S                                              
FORMUL   4  NAG    C8 H15 N O6                                                  
FORMUL   5   NA    2(NA 1+)                                                     
FORMUL   7  HOH   *349(H2 O)                                                    
HELIX    1   1 PHE A    7  SER A   11  5                                   5    
HELIX    2   3 ALA A   55  LEU A   59  1                                   5    
HELIX    3   5 ASP A  125  LEU A  130  1                                   9    
HELIX    4   6 GLU A  164  SER A  171  1                                   8    
HELIX    5   8 VAL A  231  PHE A  245  1                                  15    
SHEET    1   A 7 SER A  20  ASP A  21  0                                        
SHEET    2   A 7 GLN A 156  PRO A 161 -1  N  VAL A 157   O  SER A  20           
SHEET    3   A 7 LYS A 135  GLY A 140 -1  N  GLY A 136   O  LEU A 160           
SHEET    4   A 7 PRO A 198  LYS A 202 -1  O  PRO A 198   N  THR A 139           
SHEET    5   A 7 TRP A 207  TRP A 215 -1  N  TYR A 208   O  MET A 201           
SHEET    6   A 7 GLY A 226  HIS A 230 -1  N  PHE A 227   O  TRP A 215           
SHEET    7   A 7 MET A 180  ALA A 183 -1  O  PHE A 181   N  TYR A 228           
SHEET    1   B 7 GLN A  30  ARG A  35  0                                        
SHEET    2   B 7 GLU A  39  LEU A  46 -1  O  GLU A  39   N  ARG A  35           
SHEET    3   B 7 TRP A  51  THR A  54 -1  N  LEU A  53   O  SER A  45           
SHEET    4   B 7 ALA A 104  LEU A 108 -1  N  ALA A 104   O  THR A  54           
SHEET    5   B 7 LYS A  81  ILE A  90 -1  N  GLU A  86   O  LYS A 107           
SHEET    6   B 7 LEU A  64  ILE A  68 -1  O  LEU A  64   N  LEU A  85           
SHEET    7   B 7 GLN A  30  ARG A  35 -1  O  MET A  32   N  ARG A  67           
SSBOND   1 CYS A    1    CYS A  122                                             
SSBOND   2 CYS A   42    CYS A   58                                             
SSBOND   3 CYS A  168    CYS A  182                                             
SSBOND   4 CYS A  191    CYS A  220                                             
LINK         OG  SER A 195                 C   AR2 I 403                        
LINK         ND2 ASN A  60G                C1  NAG A 250                        
CISPEP   1 SER A   36A   PRO A   37          0        -0.07                     
CRYST1   62.982  117.521   47.989  90.00  90.00  90.00 P 21 21 2     4          
ORIGX1      1.000000  0.000000  0.000000        0.00000                         
ORIGX2      0.000000  1.000000  0.000000        0.00000                         
ORIGX3      0.000000  0.000000  1.000000        0.00000                         
SCALE1      0.015878  0.000000  0.000000        0.00000                         
SCALE2      0.000000  0.008509  0.000000        0.00000                         
SCALE3      0.000000  0.000000  0.020838        0.00000