PDB entry 1c1e

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HEADER    IMMUNE SYSTEM                           22-JUL-99   1C1E              
TITLE     CRYSTAL STRUCTURE OF A DIELS-ALDERASE CATALYTIC ANTIBODY              
TITLE    2 1E9 IN COMPLEX WITH ITS HAPTEN                                       
COMPND    MOL_ID: 1;                                                            
COMPND   2 MOLECULE: CATALYTIC ANTIBODY 1E9 (LIGHT CHAIN);                      
COMPND   3 CHAIN: L;                                                            
COMPND   4 FRAGMENT: FAB FRAGMENT;                                              
COMPND   5 MOL_ID: 2;                                                           
COMPND   6 MOLECULE: CATALYTIC ANTIBODY 1E9 (HEAVY CHAIN);                      
COMPND   7 CHAIN: H;                                                            
COMPND   8 FRAGMENT: FAB FRAGMENT                                               
SOURCE    MOL_ID: 1;                                                            
SOURCE   2 ORGANISM_SCIENTIFIC: MUS MUSCULUS;                                   
SOURCE   3 ORGANISM_COMMON: MOUSE;                                              
SOURCE   4 MOL_ID: 2;                                                           
SOURCE   5 ORGANISM_SCIENTIFIC: MUS MUSCULUS;                                   
SOURCE   6 ORGANISM_COMMON: MOUSE                                               
KEYWDS    CATALYTIC ANTIBODY, DIELS-ALDER, IMMUNOGLOBULIN                       
EXPDTA    X-RAY DIFFRACTION                                                     
AUTHOR    J.XU,I.A.WILSON                                                       
REVDAT   2   25-OCT-00 1C1E    3       ATOM   SHEET  HELIX  REMARK              
REVDAT   2 2                   3       SEQRES                                   
REVDAT   1   01-MAR-00 1C1E    0                                                
JRNL        AUTH   J.XU,Q.DENG,J.CHEN,K.N.HOUK,J.BARTEK,D.HILVERT,              
JRNL        AUTH 2 I.A.WILSON                                                   
JRNL        TITL   EVOLUTION OF SHAPE COMPLEMENTARITY AND CATALYTIC             
JRNL        TITL 2 EFFICIENCY FROM A PRIMORDIAL ANTIBODY TEMPLATE.              
JRNL        REF    SCIENCE                       V. 286  2345 1999              
JRNL        REFN   ASTM SCIEAS  US ISSN 0036-8075                               
REMARK   1                                                                      
REMARK   2                                                                      
REMARK   2 RESOLUTION. 1.90 ANGSTROMS.                                          
REMARK   3                                                                      
REMARK   3 REFINEMENT.                                                          
REMARK   3   PROGRAM     : X-PLOR 98.0                                          
REMARK   3   AUTHORS     : BRUNGER                                              
REMARK   3                                                                      
REMARK   3  DATA USED IN REFINEMENT.                                            
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.90                           
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 20.00                          
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 2.000                          
REMARK   3   DATA CUTOFF HIGH         (ABS(F)) : NULL                           
REMARK   3   DATA CUTOFF LOW          (ABS(F)) : NULL                           
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : 86.2                           
REMARK   3   NUMBER OF REFLECTIONS             : 30764                          
REMARK   3                                                                      
REMARK   3  FIT TO DATA USED IN REFINEMENT.                                     
REMARK   3   CROSS-VALIDATION METHOD          : NULL                            
REMARK   3   FREE R VALUE TEST SET SELECTION  : 5% RANDOM                       
REMARK   3   R VALUE            (WORKING SET) : 0.239                           
REMARK   3   FREE R VALUE                     : 0.294                           
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : NULL                            
REMARK   3   FREE R VALUE TEST SET COUNT      : 1521                            
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : NULL                            
REMARK   3                                                                      
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.                                  
REMARK   3   TOTAL NUMBER OF BINS USED           : NULL                         
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : NULL                         
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : NULL                         
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : NULL                         
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : NULL                         
REMARK   3   BIN R VALUE           (WORKING SET) : NULL                         
REMARK   3   BIN FREE R VALUE                    : NULL                         
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : NULL                         
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : NULL                         
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : NULL                         
REMARK   3                                                                      
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.                    
REMARK   3   PROTEIN ATOMS            : 3338                                    
REMARK   3   NUCLEIC ACID ATOMS       : 0                                       
REMARK   3   HETEROGEN ATOMS          : 28                                      
REMARK   3   SOLVENT ATOMS            : 195                                     
REMARK   3                                                                      
REMARK   3  B VALUES.                                                           
REMARK   3   FROM WILSON PLOT           (A**2) : 15.17                          
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : NULL                           
REMARK   3   OVERALL ANISOTROPIC B VALUE.                                       
REMARK   3    B11 (A**2) : NULL                                                 
REMARK   3    B22 (A**2) : NULL                                                 
REMARK   3    B33 (A**2) : NULL                                                 
REMARK   3    B12 (A**2) : NULL                                                 
REMARK   3    B13 (A**2) : NULL                                                 
REMARK   3    B23 (A**2) : NULL                                                 
REMARK   3                                                                      
REMARK   3  ESTIMATED COORDINATE ERROR.                                         
REMARK   3   ESD FROM LUZZATI PLOT        (A) : NULL                            
REMARK   3   ESD FROM SIGMAA              (A) : NULL                            
REMARK   3   LOW RESOLUTION CUTOFF        (A) : NULL                            
REMARK   3                                                                      
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.                         
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : NULL                            
REMARK   3   ESD FROM C-V SIGMAA          (A) : NULL                            
REMARK   3                                                                      
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.                                   
REMARK   3   BOND LENGTHS                 (A) : 0.012                           
REMARK   3   BOND ANGLES            (DEGREES) : 1.70                            
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : NULL                            
REMARK   3   IMPROPER ANGLES        (DEGREES) : NULL                            
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL MODEL : NULL                                      
REMARK   3                                                                      
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA                
REMARK   3   MAIN-CHAIN BOND              (A**2) : NULL  ; NULL                 
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : NULL  ; NULL                 
REMARK   3   SIDE-CHAIN BOND              (A**2) : NULL  ; NULL                 
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : NULL  ; NULL                 
REMARK   3                                                                      
REMARK   3  NCS MODEL : NULL                                                    
REMARK   3                                                                      
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT          
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL                 
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL                 
REMARK   3                                                                      
REMARK   3  PARAMETER FILE  1  : NULL                                           
REMARK   3  TOPOLOGY FILE  1   : NULL                                           
REMARK   3                                                                      
REMARK   3  OTHER REFINEMENT REMARKS: MAXIMUM LIKELIHOOD TARGET USED            
REMARK   4                                                                      
REMARK   4 1C1E COMPLIES WITH FORMAT V. 3.0, 1-DEC-2006                         
REMARK   4                                                                      
REMARK   4 THIS IS THE REMEDIATED VERSION OF THIS PDB ENTRY.                    
REMARK   4 REMEDIATED DATA FILE REVISION 3.100 (2007-03-16)                     
REMARK 100                                                                      
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB .                              
REMARK 100 THE RCSB ID CODE IS RCSB009372.                                      
REMARK 200                                                                      
REMARK 200 EXPERIMENTAL DETAILS                                                 
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION                  
REMARK 200  DATE OF DATA COLLECTION        : 14-JUL-1997                        
REMARK 200  TEMPERATURE           (KELVIN) : 97.2                               
REMARK 200  PH                             : 5.50                               
REMARK 200  NUMBER OF CRYSTALS USED        : 1                                  
REMARK 200                                                                      
REMARK 200  SYNCHROTRON              (Y/N) : Y                                  
REMARK 200  RADIATION SOURCE               : SSRL                               
REMARK 200  BEAMLINE                       : BL7-1                              
REMARK 200  X-RAY GENERATOR MODEL          : NULL                               
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M                                  
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.98                               
REMARK 200  MONOCHROMATOR                  : NULL                               
REMARK 200  OPTICS                         : NULL                               
REMARK 200                                                                      
REMARK 200  DETECTOR TYPE                  : AREA DETECTOR                      
REMARK 200  DETECTOR MANUFACTURER          : MARRESEARCH                        
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : DENZO                              
REMARK 200  DATA SCALING SOFTWARE          : SCALEPACK                          
REMARK 200                                                                      
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 39213                              
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.900                              
REMARK 200  RESOLUTION RANGE LOW       (A) : 19.960                             
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 0.000                              
REMARK 200                                                                      
REMARK 200 OVERALL.                                                             
REMARK 200  COMPLETENESS FOR RANGE     (%) : 98.5                               
REMARK 200  DATA REDUNDANCY                : 3.200                              
REMARK 200  R MERGE                    (I) : 0.07000                            
REMARK 200  R SYM                      (I) : NULL                               
REMARK 200   FOR THE DATA SET  : 14.3000                            
REMARK 200                                                                      
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.                                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.90                     
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.97                     
REMARK 200  COMPLETENESS FOR SHELL     (%) : 99.4                               
REMARK 200  DATA REDUNDANCY IN SHELL       : 3.00                               
REMARK 200  R MERGE FOR SHELL          (I) : 0.83600                            
REMARK 200  R SYM FOR SHELL            (I) : NULL                               
REMARK 200   FOR SHELL         : 1.200                              
REMARK 200                                                                      
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH                              
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: NULL                         
REMARK 200 SOFTWARE USED: AMORE                                                 
REMARK 200 STARTING MODEL: NULL                                                 
REMARK 200                                                                      
REMARK 200 REMARK: NULL                                                         
REMARK 280                                                                      
REMARK 280 CRYSTAL                                                              
REMARK 280 SOLVENT CONTENT, VS   (%): 52.88                                     
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.61                     
REMARK 280                                                                      
REMARK 280 CRYSTALLIZATION CONDITIONS: PEG 10K, IMIDAZOLIUM MALATE, PH          
REMARK 280  5.5, VAPOR DIFFUSION, HANGING DROP, TEMPERATURE 277.15K             
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY                                            
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: C 2 2 21                         
REMARK 290                                                                      
REMARK 290      SYMOP   SYMMETRY                                                
REMARK 290     NNNMMM   OPERATOR                                                
REMARK 290       1555   X,Y,Z                                                   
REMARK 290       2555   -X,-Y,1/2+Z                                             
REMARK 290       3555   -X,Y,1/2-Z                                              
REMARK 290       4555   X,-Y,-Z                                                 
REMARK 290       5555   1/2+X,1/2+Y,Z                                           
REMARK 290       6555   1/2-X,1/2-Y,1/2+Z                                       
REMARK 290       7555   1/2-X,1/2+Y,1/2-Z                                       
REMARK 290       8555   1/2+X,1/2-Y,-Z                                          
REMARK 290                                                                      
REMARK 290     WHERE NNN -> OPERATOR NUMBER                                     
REMARK 290           MMM -> TRANSLATION VECTOR                                  
REMARK 290                                                                      
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS                            
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM             
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY                
REMARK 290 RELATED MOLECULES.                                                   
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       83.75000            
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000       83.75000            
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000        0.00000            
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000        0.00000            
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000            
REMARK 290   SMTRY1   5  1.000000  0.000000  0.000000       22.36500            
REMARK 290   SMTRY2   5  0.000000  1.000000  0.000000       66.22000            
REMARK 290   SMTRY3   5  0.000000  0.000000  1.000000        0.00000            
REMARK 290   SMTRY1   6 -1.000000  0.000000  0.000000       22.36500            
REMARK 290   SMTRY2   6  0.000000 -1.000000  0.000000       66.22000            
REMARK 290   SMTRY3   6  0.000000  0.000000  1.000000       83.75000            
REMARK 290   SMTRY1   7 -1.000000  0.000000  0.000000       22.36500            
REMARK 290   SMTRY2   7  0.000000  1.000000  0.000000       66.22000            
REMARK 290   SMTRY3   7  0.000000  0.000000 -1.000000       83.75000            
REMARK 290   SMTRY1   8  1.000000  0.000000  0.000000       22.36500            
REMARK 290   SMTRY2   8  0.000000 -1.000000  0.000000       66.22000            
REMARK 290   SMTRY3   8  0.000000  0.000000 -1.000000        0.00000            
REMARK 290                                                                      
REMARK 290 REMARK: NULL                                                         
REMARK 300                                                                      
REMARK 300 BIOMOLECULE: 1                                                       
REMARK 300 THIS ENTRY CONTAINS THE CRYSTALLOGRAPHIC ASYMMETRIC UNIT             
REMARK 300 WHICH CONSISTS OF 2 CHAIN(S). SEE REMARK 350 FOR                     
REMARK 300 INFORMATION ON GENERATING THE BIOLOGICAL MOLECULE(S).                
REMARK 350                                                                      
REMARK 350 GENERATING THE BIOMOLECULE                                           
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN           
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE                
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS          
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND                          
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.                               
REMARK 350                                                                      
REMARK 350 BIOMOLECULE: 1                                                       
REMARK 350 APPLY THE FOLLOWING TO CHAINS: L, H                                  
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000            
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000            
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000            
REMARK 375                                                                      
REMARK 375 SPECIAL POSITION                                                     
REMARK 375 THE FOLLOWING ATOMS ARE FOUND TO BE WITHIN 0.15 ANGSTROMS            
REMARK 375 OF A SYMMETRY RELATED ATOM AND ARE ASSUMED TO BE ON SPECIAL          
REMARK 375 POSITIONS.                                                           
REMARK 375                                                                      
REMARK 375 ATOM RES CSSEQI                                                      
REMARK 375      HOH     3   LIES ON A SPECIAL POSITION.                         
REMARK 375      HOH   263   LIES ON A SPECIAL POSITION.                         
REMARK 375      HOH   289   LIES ON A SPECIAL POSITION.                         
REMARK 375      HOH   294   LIES ON A SPECIAL POSITION.                         
REMARK 375      HOH   332   LIES ON A SPECIAL POSITION.                         
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND LENGTHS                                      
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,2(A3,1X,A1,I4,A1,1X,A4,3X),F6.3)                  
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   RES CSSEQI ATM2   DEVIATION                     
REMARK 500    MET H 100B  SD    MET H 100B  CE    -0.087                        
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES                                       
REMARK 500                                                                      
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES              
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE               
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN               
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                 
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)              
REMARK 500                                                                      
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991                                
REMARK 500                                                                      
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3                                     
REMARK 500    LEU L   2   CA  -  CB  -  CG  ANGL. DEV. =-15.2 DEGREES           
REMARK 500    VAL L  51   N   -  CA  -  C   ANGL. DEV. = 11.4 DEGREES           
REMARK 500    PHE L  95   N   -  CA  -  CB  ANGL. DEV. = 12.7 DEGREES           
REMARK 500    PHE L  94   CA  -  C   -  N   ANGL. DEV. =-12.5 DEGREES           
REMARK 500    PHE L  95   C   -  N   -  CA  ANGL. DEV. = 15.6 DEGREES           
REMARK 500    LEU L 136   N   -  CA  -  C   ANGL. DEV. =-11.6 DEGREES           
REMARK 500    GLN H 133   N   -  CA  -  C   ANGL. DEV. = 11.0 DEGREES           
REMARK 500    ASN H 135   N   -  CA  -  C   ANGL. DEV. =-10.6 DEGREES           
REMARK 500                                                                      
REMARK 500 GEOMETRY AND STEREOCHEMISTRY                                         
REMARK 500 SUBTOPIC: TORSION ANGLES                                             
REMARK 500                                                                      
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:            
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;               
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).                             
REMARK 500                                                                      
REMARK 500 STANDARD TABLE:                                                      
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)                    
REMARK 500                                                                      
REMARK 500  M RES CSSEQI        PSI       PHI                                   
REMARK 500    VAL L  51      -46.98     70.94                                   
REMARK 500    GLN H 133     -104.14    139.43                                   
SEQRES   1 L  216  GLU LEU VAL MET THR GLN THR PRO LEU SER LEU PRO VAL          
SEQRES   2 L  216  SER LEU GLY ASP GLN ALA SER ILE SER CYS ARG SER SER          
SEQRES   3 L  216  GLN SER LEU VAL HIS SER ASN GLY ASN THR TYR LEU HIS          
SEQRES   4 L  216  TRP TYR LEU GLN LYS PRO GLY GLN SER PRO LYS PHE LEU          
SEQRES   5 L  216  ILE TYR LYS VAL SER ASN ARG PHE SER GLY VAL PRO ASP          
SEQRES   6 L  216  ARG PHE GLY GLY SER GLY SER GLY THR ASP PHE ILE LEU          
SEQRES   7 L  216  LYS ILE SER ARG VAL GLU ALA GLU ASP LEU GLY VAL TYR          
SEQRES   8 L  216  PHE CYS PHE GLN SER THR HIS PHE PHE PRO THR PHE GLY          
SEQRES   9 L  216  GLY GLY THR LYS LEU GLU ILE LYS SER ALA ASP ALA ALA          
SEQRES  10 L  216  PRO THR VAL SER ILE PHE PRO PRO SER SER GLU GLN LEU          
SEQRES  11 L  216  THR SER GLY GLY ALA SER VAL VAL CYS PHE LEU ASN ASN          
SEQRES  12 L  216  PHE TYR PRO LYS ASP ILE ASN VAL LYS TRP LYS ILE ASP          
SEQRES  13 L  216  GLY SER GLU ARG GLN ASN GLY VAL LEU ASN SER TRP THR          
SEQRES  14 L  216  ASP GLN ASP SER LYS ASP SER THR TYR SER MET SER SER          
SEQRES  15 L  216  THR LEU THR LEU THR LYS ASP GLU TYR GLU ARG HIS ASN          
SEQRES  16 L  216  SER TYR THR CYS GLU ALA THR HIS LYS THR SER THR SER          
SEQRES  17 L  216  PRO ILE VAL LYS SER PHE ASN ARG                              
SEQRES   1 H  219  GLN ILE GLN LEU VAL GLN SER GLY PRO GLU LEU LYS LYS          
SEQRES   2 H  219  PRO GLY GLU THR VAL LYS ILE SER CYS LYS ALA SER GLY          
SEQRES   3 H  219  TYR MET PHE THR ASN TYR GLY MET ASN TRP VAL LYS GLN          
SEQRES   4 H  219  ALA PRO GLY LYS ALA LEU LYS LEU MET GLY TRP ILE ASN          
SEQRES   5 H  219  PRO TYR THR GLY GLU SER THR PHE ALA ASP ASP PHE LYS          
SEQRES   6 H  219  GLY ARG PHE ALA PHE PHE LEU GLU THR SER ALA THR THR          
SEQRES   7 H  219  ALA TYR LEU GLN ILE ASN ASN LEU LYS ASN GLU ASP MET          
SEQRES   8 H  219  ALA THR TYR PHE CYS ALA ARG GLY THR THR ILE VAL ARG          
SEQRES   9 H  219  ALA MET ASP TYR TRP GLY GLN GLY THR SER LEU THR VAL          
SEQRES  10 H  219  SER SER ALA LYS THR THR PRO PRO SER VAL TYR PRO LEU          
SEQRES  11 H  219  ALA PRO GLY SER ALA ALA GLN THR ASN SER MET VAL THR          
SEQRES  12 H  219  LEU GLY CYS LEU VAL LYS GLY TYR PHE PRO GLU PRO VAL          
SEQRES  13 H  219  THR VAL THR TRP ASN SER GLY SER LEU SER SER GLY VAL          
SEQRES  14 H  219  HIS THR PHE PRO ALA VAL LEU GLN SER ASP LEU TYR THR          
SEQRES  15 H  219  LEU SER SER SER VAL THR VAL PRO SER SER PRO ARG PRO          
SEQRES  16 H  219  SER GLU THR VAL THR CYS ASN VAL ALA HIS PRO ALA SER          
SEQRES  17 H  219  SER THR LYS VAL ASP LYS LYS ILE VAL PRO ARG                  
HET    ENH    703      19                                                       
HET    DMR    702       9                                                       
HETNAM     ENH 1,7,8,9,10,10-HEXACHLORO-4-METHYL-4-AZA-                         
HETNAM   2 ENH  TRICYCLO[5.2.1.0(2,6)]DEC-8-ENE-3,5-DIONE                       
HETNAM     DMR 2-HYDROXY-SUCCINIC ACID                                          
FORMUL   3  ENH    C10 H5 CL6 N O2                                              
FORMUL   4  DMR    C4 H6 O5                                                     
FORMUL   5  HOH   *195(H2 O)                                                    
HELIX    1   1 GLU L   79  LEU L   83  5                                   5    
HELIX    2   2 SER L  121  SER L  127  1                                   7    
HELIX    3   3 LYS L  183  GLU L  187  1                                   5    
HELIX    4   4 MET H   28  TYR H   32  5                                   5    
HELIX    5   5 ASP H   61  LYS H   64  5                                   4    
HELIX    6   6 THR H   73  ALA H   75  5                                   3    
HELIX    7   7 LYS H   83  MET H   87  5                                   5    
HELIX    8   8 SER H  163  SER H  165  5                                   3    
HELIX    9   9 PRO H  213  SER H  216  5                                   4    
SHEET    1   A 4 MET L   4  THR L   7  0                                        
SHEET    2   A 4 ALA L  19  SER L  25 -1  N  SER L  22   O  THR L   7           
SHEET    3   A 4 ASP L  70  ILE L  75 -1  N  PHE L  71   O  CYS L  23           
SHEET    4   A 4 PHE L  62  SER L  67 -1  O  GLY L  63   N  LYS L  74           
SHEET    1   B 5 ASN L  53  ARG L  54  0                                        
SHEET    2   B 5 LYS L  45  TYR L  49 -1  N  TYR L  49   O  ASN L  53           
SHEET    3   B 5 LEU L  33  GLN L  38 -1  N  TRP L  35   O  ILE L  48           
SHEET    4   B 5 GLY L  84  GLN L  90 -1  N  VAL L  85   O  GLN L  38           
SHEET    5   B 5 THR L  97  PHE L  98 -1  O  THR L  97   N  GLN L  90           
SHEET    1   C 6 ASN L  53  ARG L  54  0                                        
SHEET    2   C 6 LYS L  45  TYR L  49 -1  N  TYR L  49   O  ASN L  53           
SHEET    3   C 6 LEU L  33  GLN L  38 -1  N  TRP L  35   O  ILE L  48           
SHEET    4   C 6 GLY L  84  GLN L  90 -1  N  VAL L  85   O  GLN L  38           
SHEET    5   C 6 THR L 102  ILE L 106 -1  O  THR L 102   N  TYR L  86           
SHEET    6   C 6 SER L  10  VAL L  13  1  N  LEU L  11   O  LYS L 103           
SHEET    1   D 4 THR L 114  PHE L 118  0                                        
SHEET    2   D 4 GLY L 129  PHE L 139 -1  O  VAL L 133   N  PHE L 118           
SHEET    3   D 4 TYR L 173  THR L 182 -1  O  TYR L 173   N  PHE L 139           
SHEET    4   D 4 VAL L 159  TRP L 163 -1  O  LEU L 160   N  THR L 178           
SHEET    1   E 4 GLU L 154  ARG L 155  0                                        
SHEET    2   E 4 ASN L 145  ILE L 150 -1  O  TRP L 148   N  ARG L 155           
SHEET    3   E 4 SER L 191  THR L 197 -1  N  THR L 193   O  LYS L 149           
SHEET    4   E 4 ILE L 205  ASN L 210 -1  N  ILE L 205   O  ALA L 196           
SHEET    1   F 4 GLN H   3  GLN H   6  0                                        
SHEET    2   F 4 VAL H  18  SER H  25 -1  O  LYS H  23   N  VAL H   5           
SHEET    3   F 4 THR H  77  ILE H  82 -1  N  ALA H  78   O  CYS H  22           
SHEET    4   F 4 PHE H  67  GLU H  72 -1  O  ALA H  68   N  GLN H  81           
SHEET    1   G 5 SER H  57  PHE H  59  0                                        
SHEET    2   G 5 LEU H  45  ILE H  51 -1  O  TRP H  50   N  THR H  58           
SHEET    3   G 5 GLY H  33  GLN H  39 -1  O  MET H  34   N  ILE H  51           
SHEET    4   G 5 ALA H  88  GLY H  95 -1  N  THR H  89   O  GLN H  39           
SHEET    1   H 6 SER H  57  PHE H  59  0                                        
SHEET    2   H 6 LEU H  45  ILE H  51 -1  O  TRP H  50   N  THR H  58           
SHEET    3   H 6 GLY H  33  GLN H  39 -1  O  MET H  34   N  ILE H  51           
SHEET    4   H 6 ALA H  88  GLY H  95 -1  N  THR H  89   O  GLN H  39           
SHEET    5   H 6 THR H 107  VAL H 111 -1  O  THR H 107   N  TYR H  90           
SHEET    6   H 6 GLU H  10  LYS H  12  1  N  GLU H  10   O  SER H 108           
SHEET    1   I 4 SER H 120  LEU H 124  0                                        
SHEET    2   I 4 MET H 137  TYR H 147 -1  O  GLY H 141   N  LEU H 124           
SHEET    3   I 4 TYR H 185  PRO H 194 -1  N  TYR H 185   O  TYR H 147           
SHEET    4   I 4 VAL H 171  THR H 173 -1  O  HIS H 172   N  SER H 190           
SHEET    1   J 4 SER H 120  LEU H 124  0                                        
SHEET    2   J 4 MET H 137  TYR H 147 -1  O  GLY H 141   N  LEU H 124           
SHEET    3   J 4 TYR H 185  PRO H 194 -1  N  TYR H 185   O  TYR H 147           
SHEET    4   J 4 VAL H 177  LEU H 178 -1  N  VAL H 177   O  THR H 186           
SHEET    1   K 3 THR H 153  TRP H 157  0                                        
SHEET    2   K 3 THR H 207  HIS H 212 -1  N  ASN H 209   O  THR H 156           
SHEET    3   K 3 THR H 217  LYS H 222 -1  O  THR H 217   N  HIS H 212           
SSBOND   1 CYS L   23    CYS L   88                                             
SSBOND   2 CYS L  134    CYS L  194                                             
SSBOND   3 CYS H   22    CYS H   92                                             
SSBOND   4 CYS H  142    CYS H  208                                             
CISPEP   1 THR L    7    PRO L    8          0         0.04                     
CISPEP   2 TYR L  140    PRO L  141          0         0.03                     
CISPEP   3 PHE H  148    PRO H  149          0         0.16                     
CISPEP   4 GLU H  150    PRO H  151          0        -0.24                     
CISPEP   5 ARG H  200    PRO H  202          0         0.21                     
CRYST1   44.730  132.440  167.500  90.00  90.00  90.00 C 2 2 21      8          
ORIGX1      1.000000  0.000000  0.000000        0.00000                         
ORIGX2      0.000000  1.000000  0.000000        0.00000                         
ORIGX3      0.000000  0.000000  1.000000        0.00000                         
SCALE1      0.022356  0.000000  0.000000        0.00000                         
SCALE2      0.000000  0.007551  0.000000        0.00000                         
SCALE3      0.000000  0.000000  0.005970        0.00000